A methodology for multi-objective cropping system design based on simulations. Application to weed management

Weeds are harmful for crop production but important for biodiversity. In order to design cropping systems that reconcile crop production and biodiversity, we need tools and methods to help farmers to deal with this issue. Here, we developed a novel method for multi-objective cropping system design aimed at scientists and technical institutes, combining a cropping system database, decision trees, the “virtual field” model FlorSys and indicators translating simulated weed floras into scores in terms of weed harmfulness (e.g. crop yield loss, weed-borne parasite risk, field infestation), weed-mediated biodiversity (e.g. food offer for bees) and herbicide use intensity. 255 existing cropping systems were simulated with FlorSys, individual indicator values were aggregated into a multi-performance score, and decision trees were built to identify combinations of management practices and probabilities for reaching performance goals. These trees are used to identify the characteristics of existing cropping systems that must be changed to achieve the chosen performance goals, depending on the user's risk strategy. Alternative systems are built and simulated with FlorSys to evaluate their multi-criteria performance. The method was applied to an existing oilseed rape/wheat/barley rotation with yearly mouldboard ploughing from Burgundy which was improved to reconcile weed harmfulness control, reduced herbicide use and biodiversity promotion, based on a risk-minimizing strategy. The best alternative replaced a herbicide entering plants via shoot tips (during emergence) and roots after barley sowing by a spring herbicide entering via leaves, introduced crop residue shredding before cereals and rolled the soil at sowing, which reduced the risk of unacceptable performance from 90% to 40%. When attempting to reconcile harmfulness control and reduced herbicide use, the best alternative changed the rotation to oilseed rape/wheat/spring pea/wheat, replaced one herbicide in oilseed rape by mechanical weeding, delayed tillage before rape and applied the PRE herbicide before oilseed rape closer to sowing. This option reduced the risk of unacceptable performance to 30%. None of the initial or alternative cropping systems succeeded in optimal performance, indicating that more diverse cropping systems with innovative management techniques and innovative combinations of techniques are needed to build the decision trees. This approach can be used in workshops with extension services and farmers in order to design cropping systems. Compared to expert-based design, it has the advantage to go beyond well-known options (e.g. plough before risky crops) to identify unconventional options, with a particular focus on interactions between cultural techniques.     

Multi-model uncertainty analysis in predicting grain N for crop rotations in Europe

Realistic estimation of grain nitrogen (N; N in grain yield) is crucial for assessing N management in crop rotations, but there is little information on the performance of commonly used crop models for simulating grain N. Therefore, the objectives of the study were to (1) test if continuous simulation (multi-year) performs better than single year simulation, (2) assess if calibration improves model performance at different calibration levels, and (3) investigate if a multi-model ensemble can substantially reduce uncertainty in reproducing grain N. For this purpose, 12 models were applied simulating different treatments (catch crops, CO₂ concentrations, irrigation, N application, residues and tillage) in four multi-year rotation experiments in Europe to assess modelling accuracy. Seven grain and seed crops in four rotation systems in Europe were included in the study, namely winter wheat, winter barley, spring barley, spring oat, winter rye, pea and winter oilseed rape. Our results indicate that the higher level of calibration significantly increased the quality of the simulation for grain N. In addition, models performed better in predicting grain N of winter wheat, winter barley and spring barley compared to spring oat, winter rye, pea and winter oilseed rape. For each crop, the use of the ensemble mean significantly reduced the mean absolute percentage error (MAPE) between simulations and observations to less than 15%, thus a multi–model ensemble can more precisely predict grain N than a random single model. Models correctly simulated the effects of enhanced N input on grain N of winter wheat and winter barley, whereas effects of tillage and irrigation were less well estimated. However, the use of continuous simulation did not improve the simulations as compared to single year simulation based on the multi-year performance, which suggests needs for further model improvements of crop rotation effects.     

Ertragsbildung von getreidereichen Fruchtfolgen und Getreide-monokulturen in einem extensiven und intensiven Anbausystem

Yield formation in cereal-rich crop rotations and monocultures in an extensive and intensive crop-management system
In a long duration trial, conducted from 1979/80 to 1992 at TU-Munich's research station in Roggenstein, the performance of monocultures of winter wheat, winter barley and winter rye, as well as numerous cereal-crop rotations were compared in an extensive and intensive crop-management system. The results obtained can be summarized as follows.
Over the course of 13 years, the influence of the immediately preceding crop on the yield of the main crops was of much greater significance than the rotation as a whole. With winter wheat, no yield differences could be observed between monoculture and cereal crop rotation (if the rotation did not include oats). Oats, rape, field bean, pea, potato and maize as preceding crops, however, in crop management systems, led to, on average, an increase in yield of 13 dt/ha from the following wheat. Winter barley yields were not significantly different in monoculture, cereal crop rotations and crop rotations containing 66% cereals. Furthermore, winter rye yields were the same in monocultures and cereal crop rotations. With all cereals, intensification of fertilizing and chemical plant protection led to a considerable increase in yield, but did not diminish the effects of the preceding crop. Hence, even with the use of modern agronomical techniques it is impossible to compensate for yield losses due to crop rotation.     

Reducing potential gene escape in time by appropriate post-harvest tillage—Evidence from field experiments with oilseed rape at 10 sites in Europe

Persistence of oilseed rapeseed in soil can result in weed problems but also reduce oil quality of following rape crops or result in unwanted gene escape which is particularly relevant in the context of genetically modified oilseed rape. In this paper data from 13 field experiments at sites in England, Austria and Germany are presented where tillage operations were tested that potentially reduce the build-up of a seed bank. In the majority of experiments seed losses were artificially simulated by broadcasting ca. 10,000 freshly ripened rapeseed m⁻² onto cereal stubbles. Oilseed rapeseedlings in autumn, the seed bank in winter–spring and yields of the following crop winter wheat were assessed as a function of tillage regime. During summer and autumn 19–70% of the seeds germinated and emerged. This part of the population was killed by following tillage operations or herbicide applications. However, 0–29%, in moist years 0–5%, of the initially broadcasted seeds developed dormancy and remained ungerminated in the soil until the following winter–spring.

Delaying incorporation of the seeds by leaving the stubble untouched for up to 4 weeks resulted in a reduced seed bank in almost every case. Also, repeated stubble tillage compared to an early single stubble tillage operation resulted in a smaller seed bank. The type of primary tillage (ploughing versus non-inversion cultivation) had no clear effect. No relation was found between the number of seedlings in autumn and the size of the seed bank the following winter–spring. Grain yield of the following crop winter wheat was not adversely affected by delayed stubble tillage.

The results indicate that stubble tillage aiming at a reduced seed bank of oilseed rape should focus on conditions avoiding induction of secondary dormancy rather than improving germination conditions. This means that, under the climatic conditions of central and western Europe, the stubble should be left untouched for several weeks after harvest before starting the usual tillage sequence with stubble tillage and ploughing or a non-inversion tillage sequence.     

Sunflower allelopathy for weed control in agriculture systems

Recent developments in weed science and allied aspects have involved several interdisciplinary approaches. In this context, indiscriminate use of herbicides for weed control has become a questionable subject, which besides controlling the weeds, the chemical herbicides are harmful in many ways to soil, crops, other plants and the environment as a whole. Taking into consideration ecologically sound weed management, in modern days the reliance on chemical herbicides has decreased and a shift towards naturally occurring biological herbicides has received great attention throughout the world. Sunflower is an annual dicotyledonous plant, herbaceous, erect, and a native of North America. It is thermo and photo-insensitive, hence it can be grown year round in sub-tropical and tropical countries. Only two spp. Helianthus annuus L. and Helianthus tuberosum are cultivated for food, the remaining spp., are ornamentals weeds and wild plants. However, H. annuus is allelopathic and inhibits the growth and development of other plants thus reducing their productivity. Sunflower is a major oil-yielding crop in India and its cultivation in northwest India started 25 to 30 years ago in areas located in the plains. In this region, rice-wheat rotation became very popular owing to its high yields; however, these crops are highly infested by weeds, thus farmers use herbicides for their control. Hence, this rotation consumes a maximum quantity of herbicides in this region, which has resulted in several problems viz., environmental pollution, human health hazards, and development of herbicide resistance in weeds. Thus, serious ecological questions about the reliance on herbicides for weed control in this rotation have been raised. One of the alternatives to overcome these problems is with the use of allelopathic strategies, including the use of weed-smothering crops for weed management and for the sustainability of agriculture. The field, pot culture, and laboratory studies have shown that inclusion of sunflower crops in rotation and intercropping considerably reduced the weed population in the current and succeeding crops. Rhizosphere soil of sunflower drastically smothered the weed germination, population, and biomass. The residual suppression effect of sunflower also persisted in the next crop up to 75 days. Thus, it is conceptualized that the inclusion of such oilseed crops before the rice crop in the rice-wheat rotation may provide satisfactory weed control in the succeeding rice crops and may minimize the use of herbicides. Likewise, the replacement of sorghum by summer sunflower oilseed crops may also help in the control of summer as well as winter weeds. More studies in this direction may provide avenues for satisfactory weed management in agro-ecosystems and may help to minimize the use of herbicides and thereby pave the way to develop sustainable agricultural practices for biodiversity conservation and enhancing biological integrity.     

Assessing non-chemical weeding strategies through mechanistic modelling of blackgrass (Alopecurus myosuroides Huds.) dynamics

Because of environmental and health safety issues, it is necessary to develop strategies that do not rely on herbicides to manage weeds. Introducing temporary grassland into annual crop rotations and mechanical weeding are the two main features that are frequently used in integrated and organic cropping systems for this purpose. To evaluate the contribution of these two factors in interaction with other cropping system components and environmental conditions, the present study updated an existing biophysical model (i.e. AlomySys) that quantifies the effects of cropping system on weed dynamics. Based on previous experiments, new sub-models were built to describe the effects on plant survival and growth reduction of mechanical weeding resulting from weed seedling uprooting and covering by soil, and those of grassland mowing resulting from tiller destruction. Additional modifications described the effect of the multi-year crop canopy of grassland on weed survival, growth, development and seed return to the soil. The improved model was used to evaluate the weed dynamics over 27 years in the conventional herbicide-based cropping system most frequently observed in farm surveys (i.e. oilseed rape/winter wheat/winter barley rotation with superficial tillage) and then to test prospective non-chemical scenarios. Preliminary simulations tested a large range of mechanical weeding and mowing strategies, varying operation frequencies, dates and, in the case of mechanical weeding, characteristics (i.e. tool, working depth, tractor speed). For mechanical weeding soon after sowing, harrowing was better than hoeing for controlling weed seed production. The later the operation, the more efficient the hoeing and the less efficient the harrowing. Tractor speed had little influence. Increasing tilling depth increased plant mortality but increased weed seed production because of additional seed germination triggering by the weeding tool. Decreasing the interrow width for hoeing was nefarious for weed control. The best combinations were triple hoeing in oilseed rape and sextuple harrowing in cereals. The best mowing strategy was mowing thrice, every 4–6 weeks, starting in mid-May. The best individual options were combined, simulated over 27 years and compared to the herbicide-based reference system. If herbicide applications were replaced solely by mechanical weeding, blackgrass infestation could not be satisfactorily controlled. If a three-year lucerne was introduced into the rotation, weed infestations were divided by ten. Replacing chisel by mouldboard ploughing before winter wheat reduced weed infestations at short, medium and long term to a level comparable to the herbicide-based reference system.     

Vorfruchtwirkungen verschiedener Blattfrüchte auf Winterweizen und Einfluß von Anbaumaßnahmen

Effects of preceding "break crops" on winter wheat and influence of cultural practices
Between 1979—83 there was tested the effect of 5 "break crops" (biennial alfalfa, clover-grass, faba bean, winter-rape, silomaize) on succeeding winter wheat and the interaction with changed cultural practices such as N-fertilization and conventional or reduced primary tillage (plough ↔ rotary tiller).
After the break crops wheat yield decreased within a range of 4 dt/ha as follows: alfalfa → beans → rape/ clover-grass/maize. Raising N-fertilization hardly allowed to improve the value of the break crops; most likely that of maize and with rotavating. Adaption of N-dressings promised greater influence.
The manner of primary tillage exerted stronger influence on the yield than the level of N-fertilization. Concerning long-term yield the rotary tiller equaled the plough after 4 break crops; after faba bean it was superior. Increasing N-fertilization tendentially favoured the rotary tiller after bean, rape and maize. Using the rotary tiller, at beginning of growth there may be calculated on 15 % higher NO₃-values and/or higher N-mineralization or N-transformation.
Eyespot disease could not be prevented sufficiently and purposefully by an other break crop, N-fertilization or primary tillage. Recurrent rotavating suppressed weed less than ploughing.     

Indentification of strong and weak yield determining components of winter oilseed rape compared with winter wheat

Yield formation of winter oilseed rape was compared with that of winter wheat in terms of absorption of PAR, radiation use efficiency and harvest index by means of crop growth simulation. Cumulative incoming PAR from April 15 until maturity, cumulative absorption of PAR, radiation use efficiency and harvest index were lower for winter oilseed rape compared with winter wheat. To improve potential seed yield of winter oilseed rape, a crop type with increased cumulative absorption of PAR, radiation use efficiency and harvest index is desirable.     

Effect of Different Preceding Crops and Crop Rotations on Yield of Winter Oil-seed Rape (Brassica napus L.)

To determine the effect of different preceding crops and crop rotations on the grain yield of oil-seed rape, a long-term rotation experiment was conducted at the Hohenschulen experimental station in Kiel, NW Germany. Additional factors included the nitrogen fertilization and the fungicide application. The results reported herein are based upon the harvest years 1988 to 1993. Averaged over the different rotations and husbandry treatments, the grain yields in the 6 experimental years varied between 2.71 t ha^?1 and 3.99 t ha^?1. In contrast, the effect of the different husbandry treatments was smaller and non significant. Averaged over 6 years, only the fungicide application caused small yield increase of 0.2 t ha^?1. The highest grain yields of 3.77 t ha^?1 or 3.65 t ha^?1 occurred when oil-seed rape was directly following peas. Low yields between 3.15 tha^?1 and 3.33 tha^?1 were obtained when oil-seed rape was grown after oilseed rape. The lowest grain yield of 3.13 t ha^?1 was produced with oil-seed rape grown in monoculture only. In rotations with oil-seed rape following a preceding cereal crop (wheat or barley), the grain yields averaged between 3.22 tha^?1 in a two course rotation and up to 3.44 tha^?1 in a four course rotation. In general, the yields of oil-seed rape increase with the length of the rotation and the length of the break between two oilseed rape crops. The yield component number of seeds per m² was affected by the previous cropping accordingly, whereas the thousand seed weight did not respond to the cropping history. Based upon disease assessments in the first years of this experiment, we argue that an increase in the incidence of fungal diseases has considerably contributed to the yield decrease of oil-seed rape in short rotations.     

Strahlungsnutzung durch Bestände ölreicher Körnerfruchtarten — Winterraps, Öllein und Sonnenblume im Vergleich

Radiation use by oil seed crops — a comparison of winter rape, linseed and sunflower For the production of grain crops rich in oil, winter rape, linseed and sunflower are similarly suitable at many arable locations. We wanted to compare the extent to which radiation (PAR) is intercepted and utilized by the individual species for dry matter and yield production. For this purpose, a 2‐year field experiment comprising the factors genotype, N fertilization and soil tillage was conducted. For five phases of crop development, growth rates (CGR, RGR and NAR) and PAR utilization were calculated. At full ripeness, total dry matter, grain, oil and energy yields, the crop‐ and year‐specific PAR supply, its interception and utilization for dry matter production and the resulting energy binding were determined. Due to the different individual vegetation periods, the PAR supply of the crops differed. The crop assimilation areas also differed, with values for winter rape and sunflower higher than those for linseed. The yield productivity of winter rape and sunflower was also higher than that of linseed. N fertilization increased yield to different extents for the different crops. On average, winter rape and sunflower produced the same amounts of dry matter and energy yield. Due to a higher harvest index, sunflower had the highest grain yield, and because the oil concentration in grain was comparatively high sunflower produced the highest oil yield, too. Under cool and wet climate conditions, however, the productivity of sunflower is offset by a relatively high yield risk because of uncertain ripening. The highest PAR utilization by linseed did not compensate for its very short vegetation period in combination with the lowest PAR interception.     

Competition and Control of Volunteer Jerusalem Artichoke in Various Crops

A major production constraint in Jerusalem artichoke ( Helianthus tuberosus L.) is caused by tubers which are not recovered at harvest. Such lost tubers raise a serious weed problem the following season. Winter wheat, oat, spring oilseed rape, sugarbeet, maize and ryegrass were grown in a field which had Jerusalem artichoke as the preceding crop in order to obtain information about their competitive ability and the efficacy of various control measurements against Jerusalem artichoke infestation. The Jerusalem artichoke treatments in these crops were: total control by regular hand weeding (TOC), mechanical/chemical control (MCC), and no control (NOC). Under the NOC treatment, Jerusalem artichoke infestation at harvest was variable among crops, with the number of shoots ranging from 9 to 25 m⁻² in oat and maize stands respectively. The number of Jerusalem artichoke shoots in the MCC plots was reduced by 50 to 99 % in oat and maize, respectively. The highest crop yields in each of the six species were realized under the TOC treatment. Insignificant yield reductions were observed in the NOC treatment of wheat, oat, rape and ryegrass. However, under this management yield reductions of 91 and 81 % occurred in sugarbeet and maize respectively. Depending on the preceding crop, 1–9 shoots m⁻² of Jerusalem artichoke were still recorded under the MCC plots in the following season. Consequently, for complete elimination of infestation, volunteers must be controlled in the second and probably in the third year following a Jerusalem artichoke crop.     

Novel Traits in Novel Crops： Fueling the Growth of Agrobiotechnology

Biotechnology has contributed greatly to improvements in agriculture during the past decades. Advances have been most spectacular in the fi eld of herbicide and insect resistance and have benefi ted primarily four major crops,     

Effects of cover crops on soil mineral nitrogen and on the yield and nitrogen content of maize

Current agricultural practice favours winter cover crops, which can not only optimize N management in field crop rotation; but also affect subsequent crops. Three field experiments were carried out in Eastern Slovenia to examine the effects of Italian ryegrass (Lolium multiflorum Lam.), winter rape (Brassica napus ssp.oleifera (Metzg.) Sinsk), subclover (Trifolium subterraneum L.), and crimson clover (Trifolium incarnatum L.) as winter cover crops on the mineral N (N_min) content of soil and on the yield and N content of subsequent maize (Zea mays L.), fertilized with 120 kg N ha⁻¹. Italian ryegrass and winter rape decreased soil N_min contents before winter and in spring more than both clovers. In contrast, clovers accumulated significantly higher amounts of N in organic matter and had lower C/N ratios than winter rape and especially Italian ryegrass. In comparison to the control (bare fallow without cover crop), clovers increased the whole above ground maize dry matter yield, maize grain yield and N contents in whole above ground plants and in grain. The yields and N contents of maize following winter rape were on the same level as the control, while yields and N contents of maize following Italian ryegrass were, in two of the experiments, at the same level as the control. The effects of Italian ryegrass on the maize as subsequent crop in the third experiment were markedly negative. Maize in the control treatment exploited N much more efficiently than in treatments with cover crops. Therefore, cover crop N management should be improved, especially with a view to optimizing the timing of net N mineralization in accordance with the N demands of the subsequent crop.     

不同作物秸秆腐解液对节节麦种子萌发和幼苗生长的影响

节节麦(Aegilops tauschii Coss.)作为麦田恶性杂草之一,给中国小麦生产带来了严重的经济损失。为探究农作物秸秆在节节麦生态防治方面的作用,本试验采用了滤纸培养皿法和小杯法研究了不同浓度的玉米、油菜、水稻秸秆及稻壳腐解液对节节麦种子萌发率、发芽指数及幼苗根长、苗高和干重5项生物指标的影响。结果表明,当浓度为6.25 mg/mL时,玉米、油菜秸秆腐解液对节节麦幼苗干重表现为促进作用,对其他测定指标均为抑制作用。稻壳腐解液除了对根长具有抑制作用外,对其他指标均具有促进作用。水稻秸秆腐解液对5项指标均具有促进作用。当浓度大于50 mg/mL时,4种秸秆腐解液对5项指标均具有抑制作用,且随着浓度的增加,抑制作用增强。总体来说,作物秸秆腐解液对节节麦的化感作用强弱依次为玉米秸秆、油菜秸秆、稻壳、水稻秸秆,且对根的化感作用尤为明显。     

The Exploitation of Crop Allelopathy in Sustainable Agricultural Production

Crop allelopathy may be useful to minimize serious problems in the present agricultural production such as environmental pollution, unsafe products, human health concerns, depletion of crop diversity, soil sickness and reduction of crop productivity. Several crops including alfalfa, buckwheat, maize, rice, rye, sorghum, sunflower, wheat, etc. are affected either by their own toxicity or phytotoxin exudates when their residues decompose in the soil, that show strong suppression on weed emergences. Allelopathic crops when used as cover crop, mulch, smother crops, green manures, or grown in rotational sequences are helpful in reducing noxious weeds and plant pathogen, improve soil quality and crop yield. Those crop plants, particularly the legumes, incorporated at 1–2 tons ha⁻¹ (alfalfa, buckwheat, rice by-products), which can give weed reduction and increase of rice yield by 70 and 20 %, respectively, are suggested for use as natural herbicides. Allelochemicals from allelopathic crops may aid in the development of biological herbicides and pesticides. Cultivating a system with allelopathic crops plays an important role in the establishment of sustainable agriculture. The introduction of allelopathic traits from accessions with strong allelopathic potential to the target crops will enhance the efficacy of crop allelopathy in future agricultural production.     

Population dynamics of volunteer oilseed rape (Brassica napus L.) affected by tillage

Volunteer plants of oilseed rape (Brassica napus L.) from persistent seeds in soil can affect subsequent crops. Apart from the agricultural disadvantages, the environment and the marketing of the seeds may also be affected, particularly if plants with special ingredients or genetically modified (gm) plants are grown. In order to investigate the influence of soil cultivation and genotype on seed persistence and gene flow via volunteers, a field experiment was set up testing four tillage treatments and two cultivars in a split-plot design. The cultivars tested were near-isogenic to two gm cultivars. To simulate harvesting losses, 10 000 seeds m⁻² were broadcast on a soil in July. The subsequent tillage treatments were combinations of immediate or delayed stubble tillage by a rotary tiller, primary tillage with plough or cultivator, or zero tillage. Over the following year, the fate of the seeds was determined. Immediate stubble tillage with following cultivator or plough resulted in 586 resp. 246 seeds m⁻² in the soil seed bank. After delayed stubble tillage with following plough, 76 seeds m⁻² were found, and no soil seed bank was built up in the zero tillage treatment. Nevertheless, in the zero tillage treatment, several robust volunteer plants survived the herbicide application before the direct drilling in autumn until following spring. In the zero tillage treatment and in the cultivator treatment, 0.19 volunteers m⁻² resp. 0.06 volunteers m⁻² flowered simultaneously to ordinarily sown oilseed rape in the following crop of winter wheat and produced 73 resp. 18 seeds m⁻². Delayed stubble tillage reduced the risk of gene escape via the soil seed bank, while zero tillage resulted in the highest risk of gene escape by pollen and by production of a new generation of seeds. In terms of a labelling threshold for gm food this number of seeds would be below the threshold of 0.9% of transgenic parts in conventially bred food or feed.     

Towards an optimal management regime to facilitate the coexistence of GM herbicide tolerant and non-GM oilseed rape

Although a national strategy for the coexistence of GM and conventional/organic crops in Ireland has been published (McGill et al., 2005), measures pertaining to the coexistence of GM and non-GM oilseed rape (OSR) crops was omitted pending the completion of regional specific research. As the evaluation of gene flow between coexisting OSR varieties cannot be solely studied through field experiments and to assist policy makers in the generation of coexistence guidelines for GM herbicide tolerant (HT) OSR, the present work utilized the gene flow model GeneSys to simulate the influence of management strategies on gene escape from GM OSR across the Irish agri-environment. The model predicted that the incorporation of GMHT OSR into an existing winter wheat-based rotation without a complimentary modification to the cropping regime would rapidly compromise coexistence, recorded as the % harvest impurities in non-GM OSR crops at the silo and the % of non-GM OSR fields whose harvest exceeded the European Union labelling threshold of 0.9% GM content. Enhanced herbicide efficacy, the application of spring weed control, the inclusion of a single spring crop (potato, barley, maize) post-OSR, reduced regional harvest impurities in non-GM OSR crops below the 0.9% threshold. The simulations showed that the establishment of GM zones by clustering of GMHT OSR fields presents for early adopters a viable mechanism to achieve coexistence with neighboring non-GM OSR sites. By incorporating a 50 m non-OSR (GM/non-GM) buffer zone to circumvent each GM cluster, the simulations concluded that regional harvest purity can be assured with up to 6 separate clusters accommodating up to 10% regional GM adoption across the landscape. Significantly, these simulations were completed using GMHT OSR as a break crop in the standard winter wheat rotation, which in previous simulations was deemed to be impractical for coexistence purposes. Organizing cultivation through GM clusters, presents a workable option to those early adopters who will seek to investigate the potential of the technology by (i) concentrating the necessity for intensive volunteer control in a specific, localised area (ii) increasing the potential for allocating machinery via GM/non-GM purposes and (iii) facilitating regulatory requirements in regards to farmer-to-farmer communication.     

Mechanical wounding under field conditions: A potential tool to increase the allelopathic inhibitory effect of cover crops on weeds?

To increase the inhibitory effect of soil-incorporated cover crop residues on germination and early growth of weeds, the allelochemical content of the cover crop at the time of soil incorporation should be maximal. We investigated whether mechanical damaging in spring induced the production of allelochemicals in late-summer sown lucerne, winter rye and winter oilseed rape. Allelopathic activity and biomass of intact cover crop plants were determined throughout spring, and mechanically damaged plants were monitored till 14 days after wounding. Allelopathic activity was determined directly by lettuce seedling bioassays (lucerne and winter rye) or indirectly by glucosinolate quantification (winter oilseed rape). During spring, the allelopathic activity per unit biomass of intact plants showed a gradual decline for winter rye and lucerne, and a steep decline at the onset of flowering for winter oilseed rape. All cover crop species attained the highest allelopathic activity per unit area at the end of the sampling period as the increase in biomass surpassed the decline in allelopathic activity per unit biomass. Although mechanical wounding enhanced the allelopathic activity per unit biomass of all three species, the effect was only minor and often just sufficient to compensate for the loss in biomass resulting from wounding. This study therefore indicates that the best option for maximizing the inhibitory effect is to incorporate residues of intact cover crops as late as possible.     

Effects of 15N Split-application on Soil and Fertiliser N Uptake of Barley, Oilseed Rape and Wheat in Different Cropping Systems

In intensive farming systems, farmers split up and apply the N fertilization to winter cereals and oilseed rape (OSR) at several dates to meet the need of the crop more precisely. Our objective was to determine how prior fertilizer N application as slurry and/or mineral N affects contributions of fertilizer‐ and soil‐derived N to N uptake of barley (1997), oilseed rape (OSR; 1998) and wheat (1999). In addition, residual fertilizer N effects were observed in the subsequent crop. Since autumn 1991, slurry (none, slurry in autumn, in spring, in autumn plus in spring) and mineral N fertilizer (0, 12 and 24 g N m⁻²) were applied annually. Each year, the treatments were located on the same plots. In 1997–1999, the splitting rates of the mineral N fertilization were labelled with ¹⁵N. Non‐fertilizer N uptake was estimated from the total N uptake and the fertilizer ¹⁵N uptake. All three crops utilized the splitting rates differently depending on the time of application. Uptake of N derived from the first N rate applied at the beginning of spring growth was poorer than that from the second splitting rate applied at stem elongation (cereals) or third splitting rate applied at ear emergence or bud formation (all three crops). In contrast, N applied later in the growing season was taken up more quickly, resulting in higher fertilizer N‐use efficiency. Mineral N fertilization of 24 g N m⁻² increased significantly non‐fertilizer N uptake of barley and OSR at most of the sampling dates during the growing season. In cereals, slurry changed the contribution of non‐fertilizer N to the grain N content only if applied in spring, while OSR utilized more autumn slurry N. In OSR and wheat, only small residual effects occurred. The results indicate that 7 years of varying N fertilization did not change the contribution of soil N to crop N uptake.     

Continuous cropping systems and weed selection

Summary Changes in the weed flora of agro-ecosystems can occur as long-term changes or temporary fluctuations in species composition. The rate at which weed population shifts occur depends on the selection pressure imposed, genetic variability among weed populations, plant characteristics and environmental factors. Agronomic practices associated with cropping systems including crop rotation, tillage, herbicide use, soil amendments, and mechanization of harvesting that impart a range of selection pressures on weed populations are discussed in this review. Widespread use of herbicides has had the greatest impact on weed selection in recent years. Evolution of herbicide resistant weeds presents an enormous challenge to farmers. Development of herbicide tolerant crops has provided another tool for farmers however the selection pressure on weeds and potential impact on weed population shifts will require judicious use of this technology. Simulation models provide an excellent opportunity to predict future weed population shifts in response to management practices. Further insight into future management changes on weed selection must proceed towards an investigation of the processes rather than the outcomes. In particular, this must involve an understanding of the ecological factors and processes that are likely to determine the weed responses to particular management regimes.