Juvenile–mature genetic correlations in <Emphasis Type="Italic">Pinus radiata D. Don</Emphasis>. under different nutrient × water regimes in Spain

Individual and family heritabilities and juvenile–mature genetic correlations were estimated for growth and biomass traits recorded in treatment with optimal and limiting water and/or nutrient availability to study how the different growth environments affected genetic parameter estimates. Thirty open-pollinated families, randomly selected among the 58 families used in field progeny tests in Galicia, were cultivated for 30 weeks in a climatic chamber under controlled conditions. Two water regimes (high and low water supply) combined with two nutrient regimes (high and low nutrient supply) were applied by subirrigation. Several growth, branching and dry mass traits were assessed 30 weeks after sowing and compared with field performance (height, diameter and volume) of 4-year-old progeny tests established at three different sites in Galicia (NW Spain). Both the irrigation and the fertilization treatments had a strong effect in all the assessed traits except irrigation for the number of branches. Heritabilities for growth and biomass traits were moderate to high (0.13–0.77) in individual treatments. However, when analyzing all treatments together, the impact of the family × treatment interactions led to a reduction (0.20–0.35) in the heritability estimates. The results indicated that the genotype × water and genotype × nutrient interactions may be important and could not be ignored in the Galician radiata pine breeding program. Climatic chamber–field correlations were different between different traits measured at climatic chamber experiments and the three field tests. The correlations were larger with the well-watered treatments, suggesting that further development of early testing methods for radiata pine in Galicia should include treatments with no limiting water availability.     

马尾松高产脂种子园家系选择

以马尾松（Pinus rnassoniana）高产脂种子园自由授粉家系子代测定林为对象,分析了4年生家系生长性状（树高、胸径、材积和活轮枝数）及抗虫性的遗传变异规律,估算了各性状的遗传力,并进行了相关性分析。结果表明,除抗虫性存在显著差异外,材积、胸径、树高、活轮枝数生长性状在家系问均存在极显著的遗传差异;生长性状受较强遗传力控制,而抗虫性受中强度遗传力控制;家系各生长性状问具有高度相关性,而产脂力、抗虫性和各生长性状间没有相关性。利用最佳线性预测的单性状（材积）及多性状联合（抗虫性和家系产脂力）估算家系育种值,选出了2个马尾松优良家系作为桂东南育种和推广材料。     

The effects of timing of pre-commercial thinning and stand density on stem form and branch characteristics of Pinus sylvestris

The effects of pre-commercial thinning on tree and branch characteristicswere analysed at two experimental sites in northern Sweden,where pre-thinning of Scots pine stands to 600, 1000 and 1800stems per hectares at heights of 1.5, 3, 5 and 7 m had beencombined in a factorial design. A total of 90 trees were cutand branches were measured in whorls sampled at 1-m intervals,starting with the whorl closest to 0.5 m above ground. Effectsof the treatments on tree and branch characteristics were analysedby a fixed-effect analysis of variance model, which was appliedseparately for each experimental site and whorl height. Diameterat breast height (d.b.h.) and living crown to height ratio decreased,while the height/d.b.h. ratio (stem form) increased with increasingstand density, and generally with increased height at thinning.No treatment effect was found on the number of branches perwhorl or branch angle, but branch diameter (BD) was significantlyinfluenced by both stand density after thinning and height atthinning. BD decreased with increases in stand density and decreasedwith increases in height at pre-commercial thinning. Relativebranch size (RBS), defined as the ratio between the sum of thediameters of branches in a whorl and stem perimeter under barkat the location of that whorl, was significantly influencedonly by stand height at the time of treatment. RBS decreasedby 4–6 per cent in whorls located below the current baseof the living crown with each step increase in stand heightat pre-commercial thinning. The results indicate that the numberof stems after pre-commercial thinning has a greater impacton tree size and shape than timing of pre-commercial thinning.Stand density after thinning also has a greater impact on branchsize, in the sense that it will affect branch size further upthe tree than the timing of pre-commercial thinning. However,by delaying the time for pre-commercial thinning it is possibleto reduce the RBS in the lower part of the stem. This mightbe an important measure to improve timber quality later in therotation period, which cannot be done solely by regulating standdensity.     

杉木第二代种子园子代遗传变异及优良家系选择研究

来自广东省龙山林场杉木第二代种子园的39个自由授粉家系参加了1998年多点子代测定和2001年单点子代测定试验,结果表明：11年生子代测定林的树高、胸径、单株材积在家系和地点间差异达到极显著水平（P〈0．01）,8年生子代测定林生长性状的差异较小;家系平均树高、胸径、材积遗传力分别为0．24、0．26、0．30,主要生长性状受中等偏下的遗传力控制;第二代种子园材积遗传增益在1．88％～5．85％．其遗传增益和现实增产效果高于1．5代种子园,也高于初级种子园。对11年生多点子代测定林家系进行了丰产稳定性分析,选择出广谱丰产稳定家系5、8、17、19、23、26、28号,平均单株材积比群体均值提高3．39％;丰产家系3、7、15、21号和家系20、22、25号,分别适合龙山林场和曲江林场及相似立地条件地区限制性推广利用,平均单株材积比群体均值分别提高9．91％和4．89％;8年生单点子代测定林,以单株材积高于群体均值作为入选标准,有1、2、4、6、7、8、10、11、12、14、15、19等12个家系入选,平均单株材积比群体均值提高6．76％。     

青海云杉半同胞子代苗期生长性状遗传分析

以青海云杉无性系种子园内30个青海云杉半同胞家系为研究对象,在张掖市肃南县塔尔沟苗圃开展苗期试验,测定5a生苗期苗高、地径、新梢长、侧枝数等生长性状。分析结果表明：（1）青海云杉半同胞子代苗家系间苗高、地径、新梢长存在极显著差异;遗传变异系数较高,均在19．76％以上,苗高、地径、新梢长、侧枝数指标的家系遗传力均较大,均在0．830以上;苗高与地径、新梢长间存在极显著正相关,地径与新梢长、侧枝数存在极显著正相关;（2）以苗高为首要选择指标,按照1倍标准差选择出158#、106#、160#、154#、109#、119#6个优良家系,各性状值高于30个家系的平均值。     

马尾松实生种子园自由授粉子代测定及亲本家系选择增益估算

以福建省白砂国有林场的马尾松实生种子园自由授粉子代测定林为材料,分析了5、6年生家系生长量(树高、胸径及材积)及形质指标(通直度、冠幅及侧枝粗度)的遗传变异规律,估算了各性状的遗传力。结果表明,材积、胸径、树高生长性状及冠幅、通直度、侧枝粗度等形质指标在家系间和家系内均存在显著的遗传差异,且家系内变异大于家系间。家系、单株遗传力分别在0.493 0.731、0.080 0.199之间。利用最佳线性预测的单性状(材积)及多性状联合(材积、冠幅、侧枝粗度及通直度)估算了6年生家系育种值。分析了材积单性状选择方式、多形状联合选择以及冠幅约束选择在不同入选率下亲本家系的预测及现实遗传增益(材积及通直度)的变化规律,结果表明,在等权重条件下,联合材积及通直度指标进行亲本家系选择是最佳选择方式,能够获得相对较高的遗传增益。     

Maturation in Douglas-fir: I. Changes in stem, branch and foliage characteristics associated with ontogenetic aging

Two experiments were conducted to characterize changes associated with ontogenetic aging in coastal Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and to identify possible maturation "markers" for this species. In the first experiment (Experiment 1), scions of ages 1, 4 and 9 years were collected from four seed zones in field progeny tests and grafted onto 1-year-old rootstocks. In Experiment 2, scions from five full-sib families of ages 1 and 10 years were collected from one progeny test and similarly grafted. Grafts for both experiments were planted in the field in a completely randomized design. The grafts were measured after two and six growing seasons. In Experiment 1, graft diameter, number of terminal bud flushes per year, number of branches, and branch length decreased with age. Plagiotropic angle and needle weight increased. All variables except needle length had a significant age x seed zone interaction. After 6 years, internodal stem diameters, numbers of nodal and internodal branches, and length and diameter of internodal branches decreased with age, and there were age x seed zone interactions with most variables. In Experiment 2, graft height and diameter, number of flushes, number and size of lateral branches, needle length and weight decreased with age. After 6 years, height and diameter, size and number of nodal and internodal branches, and leaf chlorophyll concentration (measured in April) decreased, but there were relatively few age x family interactions. An analysis based on traits that were significantly affected by age, but that did not interact with seed zone or family, indicated that main stem diameter, nodal branch length and nodal branch diameter were the most consistent and reliable maturation markers.     

Comparison of terrestrial laser scanning and X-ray scanning in measuring Scots pine (Pinus sylvestris L.) branch structure

While X-ray scanning is increasingly used to measure the interior quality of logs, terrestrial laser scanning (TLS) could be used to collect information on external tree characteristics. As branches are one key indicator of wood quality, we compared TLS and X-ray scanning data in deriving whorl locations and each whorl’s maximum branch and knot diameters for 162 Scots pine (Pinus sylvestris L.) log sections. The mean number of identified whorls per tree was 37.25 and 22.93 using X-ray and TLS data, respectively. The lowest TLS-derived whorl in each sample tree was an average 5.56?m higher than that of the X-ray data. Whorl-to-whorl mean distances and the means of the maximum branch and knot diameters in a whorl measured for each sample tree using TLS and X-ray data had mean differences of ?0.12?m and ?6.5?mm, respectively. One of the most utilized wood quality indicators, tree-specific maximum knot diameter measured by X-ray, had no statistically significant difference to the tree-specific maximum branch diameter measured from the TLS point cloud. It appears challenging to directly derive comparative branch structure information using TLS and X-ray. However, some features that are extractable from TLS point clouds are potential wood quality indicators.     

Modeling internode length and branch characteristics for Pinus radiata in New Zealand

It is recognized that estimation of internode length and maximum branch size is important for the prediction of clearwood in unpruned timber stands, as well as for evaluating the quality and value of logs in general. A review of existing branch models reveals a diversity of approaches as well as a tendency for results to be species specific. Here, a branch model is developed for Pinus radiata in New Zealand, capable of predicting successive internode lengths, the number of branches on each branch cluster, and the size of each branch up to the green crown (GC) at site index age 20. Inputs to the model include tree height and diameter at breast height (dbh) (both at age 20) and basal area per hectare of the top 100 stems. Further optional inputs are an ocular count of the number of branch clusters up to the green crown, and branch factor (BF)—the size of the biggest branch in the first cluster encountered above 6 m. The vertical distributions of internode length and maximum branch diameter are found to reach maxima around 0.3–0.4 of relative height. Internode length and the number of branches per whorl were found to be independent of tree size, site index and stand density. Stems per hectare is not required as an explicit predictor variable but it appears implicitly through tree dbh. Some model output is given and the results are discussed.     

Crown architecture of Abies balsamea from four canopy positions

Data collected from four distinct canopy positions from each of 39 Abies balsamea (L.) Miller trees were used to construct models to describe the cumulative leaf area distribution within the crown and to predict the needle mass of individual branches, the average branch angle, branch diameter, branch length, and crown radius per whorl, and the average number of living branches per whorl. We tested the hypotheses that regression models are equal among canopy positions and that a model to predict branch needle mass is valid at the northern and southern extremes of the central climatic zone of Maine. Canopy position had an effect on the models constructed to predict needle mass, branch angle, branch diameter, branch length, crown radius, and the number of living branches per whorl. However, compared with an expanded model that incorporated parameters calculated for each crown class, there was only a small loss in model precision when a general model constructed from data pooled from all crown classes was used to predict needle mass, branch angle, and branch diameter. Regression equations unique to each crown class were needed to predict crown shape and leaf area distribution in the crown satisfactorily. Our branch needle mass model, which was constructed from data collected at the southern extreme of the central climatic zone of Maine, consistently underestimated needle branch mass when applied to the northern extreme of the central climatic zone.     

Height growth recovery and crown development in top‐damaged Pinus sylvestris trees

Leader shoot damage was imposed on young Scots pines (3 m high) in four ways: by caging the uppermost whorl including the leader shoot with 10 pine shoot beetles; by caging the two uppermost whorls with 20 beetles; by topping the tree below the 1984 whorl; by topping the trees as above and removing all current shoots in whorl 1983. Six years later, top‐damaged trees were still significantly shorter than undamaged controls, although the annual height growth rate had recovered to the pre‐damage level within 3 years. Recovery occurred in two ways: lateral branches from the nearest intact whorl took dominance over the damaged leader or secondary branches developed from interfascicular buds on the damaged leader shoot. Approximately one and two years of height growth were lost in the former and latter case, respectively. Only 5 of the 34 top‐damaged trees escaped technical defects (crooks and/or vertical branches). Stem diameter and crown development were slightly affected.     

杉木优良家系选择的研究

应用杉木382个半同胞家系的子代测定数据,分析主要生长性状的变异原因和规律,估算重要数量性状的遗传参数,探讨抗逆性的表达和遗传差异。 选出可在川南和川东南杉木产区推广的速生型优良家系和川西杉木产区推广的抗逆速生型优良家系共119个,材积的平均增产率约43％、遗传增益约22％。 根据杉木生长性状间的相关,树高年度相关和亲—子树高、胸径、材积的相关研究,讨论了杉木遗传改良中的一些技术问题。     

Effect of pruning on branch production and water relations in widely spaced ponderosa pines

This study dealt with the effects of pruning on branch and leaf area (F _a) production of ponderosa pines growing in silvopastoral systems in Patagonia. We hypothesized that pruning positively influences the number of branches per whorl and their basal area growth rate, changing F _a production. In addition, we studied some water relations in order to explain potential differences in branch growth rates between treatments. Two mathematical models were developed to estimate branch and total F _a. The averaged diameter at the third year of pruning was, for high-pruned trees 3.1 and 3.6 cm at the bottom and middle of the crown, against 4 and 4.4 cm for low-pruned trees. Pruning did not produce changes in the number of branches per whorl (approximately 7.6 branches per whorl). Water stress may be responsible of this lower branch growth in pruned trees. Water potential, stomatal conductance and transpiration were lower in high- than in low-pruned trees.     

闵东桉树引种抗逆性的早期选择

比较了桉树7个品系在3个不同海拔的成活率、耐寒性、抗虫性、树干弯曲程度和径高比5个性状。结果表明:不同品系的耐寒性、抗虫性、树干弯曲程度、径高比存在显著差异,海拔高度对成活率、耐寒性、树干弯曲程度、径高比具有显著影响;多重比较结果表明:比较适宜闵东发展的桉树品系是尾赤桉201、其次是尾巨桉3229和尾圆桉184。     

火炬松纸浆材优良家系多性状选择

对６年生火炬松自由授粉子代测定林进行了生长、材性和干形等性状的遗传参数估算和相关关系分析。树高、木材基本密度在家系间存在极显著差异，且具有较高的遗传力，胸径、材积、干材干重和管胞长度具中度遗传力，树干通直度和分枝性状未发现显著的家系间差异。干材干重与生长性状呈高度正相关，与木材基本密度呈微弱的负相关，而与管胞长度呈负相关。以浆材为选择目的进行家系指数选择，认为包含产量（干材干重）和质量（木材基本密     

Patterns in aboveground carbon allocation and tree architecture that favor stem growth in young Scots pine from high latitudes

The production and allocation of aboveground biomass and the characteristics of tree architecture were examined in eight-year-old Scots pine (Pinus sylvestris L.). Considerable among-tree variation existed in tree architecture, total aboveground dry mass production, and dry mass partitioning among tree parts. A linear relationship existed between needle and branch mass. Stem mass was directly proportional to tree height, which in turn was directly proportional to the allocation ratio between stem mass and total needle + branch mass production. The architectural characteristics that were related to a high proportional allocation to stem and high stemwood production were a large mean shoot volume, large mean number of branches per whorl, long needle retention and a high crown length/crown width ratio. Individual trees were found that combined high stemwood production with both high harvest index and high stemwood specific gravity.     

Implication of genotype × site interaction on <Emphasis Type="Italic">Pinus radiata</Emphasis> breeding in Galicia

The magnitude and practical importance of family × site interactions for growth and form traits considered in radiata pine (Pinus radiata D.Don) breeding were analysed by four different methods: type B genetic correlation, regression coefficient, mean rank deviation and ecovalence. The material analysed consisted in growth, form and frost resistance assessments of 58 open pollinated half-sib families at 3 to 4 years old across three sites in Galicia (NW Spain). Analysis of variance revealed that family × site interactions were quantitatively important for all traits (diameter, volume, branch angle, straightness and frost resistance). The losses in potential genetic gain for a breeding population were important for all traits under individual selection but only for diameter and frost resistance under family selection. Omission of the ten most interactive families from the analysis considerably reduced the losses for all traits, although losses in potential genetic gain remained important for frost resistance under family selection. The results indicate that elimination of these families from breeding programmes would be an effective strategy for selecting for stability in performance across sites for all traits except frost resistance. In order to overcome this problem, selecting varieties for frost resistance specifically adapted to various regional contexts would be an option. However further studies under controlled conditions are required before making final decisions for radiata pine breeding programmes.     

洞庭湖区短轮伐期杨树人工林生物量调查

为实现杨树人工林的分类经营,提高杨树人工林的经济效益,采用标准木法对洞庭湖区杨树人工林中不同径阶林木生物量进行了测定。结果表明:7年生杨树人工林的径阶集中分布在14~20 cm。不同径阶杨树单株不同器官生物量均遵循树干树枝树根(含桩)树皮树叶的分配规律;随着径阶的增长,树干在单株、林分内所占的比例均呈增长趋势;小于5 cm的主干、树枝和根的生物量与地上部分的胸径和树高均成幂函数关系,相关系数分别为0.839 4、0.940 5和0.994 7。     

Crown and branch attributes of mid-aged Betula alnoides plantations in response to planting density

Both crown and branch characteristics greatly affect tree growth and timber quality, and their development is closely related to planting density (PD). Here crown and branch attributes of Betula alnoides were investigated in a 14-year-old experimental plantation in southern China with five planting densities ranging from 500 to 3333 stems per hectare (sph). The results showed that high PD significantly reduced crown and branch sizes. The ratios of crown diameter to stem diameter at breast height at the stand level and for dominant and co-dominant trees were almost constant regardless of PD. PD also had no significant influence on the number and angle of branches, and only planting with the highest density significantly increased branch density and mortality. Insertion angle (IA) of dead branches was greater than that of live branches and these differences were significant at the two low planting densities (833 and 500?sph). Linear mixed models revealed that branch diameter (BD) was negatively correlated with its relative height in the crown and IA. The binary-logistic regression model indicated that branch status (live or dead) was codetermined by BD and PD. These findings can help farmers cultivate high-quality wood of B. alnoides and obtain a higher economic return.     

马尾松实生种子园五年生的植株生长与分枝

本文研究了马尾松实生种子园建立第五年时植株的生长与分枝状况,研究的性状包括树高、胸径、枝条总数、年高生长、 年枝条生长量、年分枝数、不同分枝处树干直径等。对总的生长与分枝性状(含树高、胸径、枝条总数)分析表明:家系间只在胸径方面存在显著差异。对分年度的生长与分枝性状的研究表明:家系间在年高生长、年枝条生长量、年枝条直径方面无显著差异;年分枝数在刚建立的前两年家系间没有显著差异,后两年则差异显著;不同分枝处树干直径在连续的四年内,家系间均存在显著差异。不同年度的生长与分枝的趋势分析结果发现,大部分家系的年高生长与年枝条生长呈递增型,而大部分家系每年的分枝数却为波动型。这些研究结果表明,该马尾松实生种子园到建立的第五年时,家系间的差异大部分都不显著。究其原因,一方面可能是马尾松初期生长节律;另一方面可能是因为建园家系复杂的父本构成。因此,这个时期进行家系选择不太有效,建议在利用优树自由授粉子代建立实生种子园时,对他们先期进行苗期选择,而不是采用所有的子代。图1表4参20。