Territorial Behaviour in Certain Horned Ungulates,With Special Reference to the Examples of Thomson's and Grant's Gazelles

An attempt has been made to distinguish between action area, home range, and territory in the Bovidae. The establishing of subjective boundaries is considered to be the most important criterion of territoriality. The existence of such boundaries becomes evident from certain behavioural symptoms; “defence” or better, localized dominance which may lead to intolerance, is one of them.

Not all bovids are territorial. Within the territorial species, there seem to be at least two types: (a) The animals, usually in pairs, may, under favourable conditions, stay in their territories permanently; (b) Only the males are territorial and stay in temporary territories, usually for several weeks or months. This last type is obviously more common in horned ungulates than the first one. Within this second type (b), there are species-specific differences. For example, in Grant's gazelle (Gazella granti), under certain environmental conditions, this type of territoriality is combined with harem behaviour, but in the co-inhabiting Thomson's gazelle (Gazella thomsoni), the females roam through the territories of the males and stay together with the same buck only for a few hours per day.

Even within one and the same species, there can be variations, apparently linked with differences in environmental conditions. This is discussed, using the examples of the Uganda kob (Adenota kobj, the wildebeest (Connochaetes taurinus), and Grant's gazelle. Finally, there can be differences in the territorial behaviour of the same individual according to the phases of territoriality (beginning, peak, end) which is shown by the example of Thomson's gazelle.     

Comparative tick counts on game, cattle and sheep on a working game ranch in Kenya

The main objective of the study was to determine the possible influence of host species on the maintenance of ticks in the field by determining the relative contribution of game animals compared to domestic animals. The study was carried out on a game ranch 32 km south-east of Nairobi. Tick counts were carried out on 30 Zebu cattle (Bos indicus) aged 1 to 3 years and 20 red Maasai sheep (Ovis aries) aged 6 months to 1 year grazing with game animals in a common area for a period of 2 years and these counts were compared with those on eland and Thomson's gazelle. Half-body counts were carried out on the cattle and sheep once every week. To avoid excessive stress, the animals were dipped in amitraz whenever the half body counts exceeded 50 fully engorged female ticks of any species. Tick counts on two wild animal species (eland (Taurotragus oryx) and Thomson's gazelle (Gazella thomsonii)) were carried out during the weekly culling of the herbivores. The results revealed that there was no significant difference in the number ticks per square metre between the wild ungulates and the domestic animals.     

Assessing the impacts of domesticated versus wild ungulates on terrestrial small mammal assemblages at Telperion Nature Reserve,South Africa

Grazing by large mammals alters vegetation physiognomy, consequently changing habitat suitability for small mammal communities. We investigated the response of terrestrial small mammals to grazing by wild and domesticated ungulates at the boundary of a protected area (Telperion Nature Reserve) and surrounding cattle ranches in Mpumalanga, South Africa over two seasons. Fifteen paired grids were set on either side of the boundary fence at which small mammals were trapped in Sherman live traps placed flat on the ground. A total of 11 760 trap nights resulted in the capture of 187 animals belonging to 14 species (11 rodents, two shrews and one elephant shrew). The small mammal communities in grasslands grazed by domesticated or wild ungulates were similar in abundance, species richness, diversity and demographic parameters, likely due to the fact that vegetation structure of the two grazing systems was also similar. We used generalised linear models to show that rock and grass cover were plausible predictors of small mammal abundance in this system. Rock cover showed a positive relationship with small mammal abundance whilst grass cover showed a negative relationship. Our observations suggest that at the scale of our study and with the current stocking densities, wild and domesticated ungulates have similar impacts on the small mammal community.     

Ovicidal and larvicidal activity in vitro of Spigelia anthelmia Linn. extracts on Haemonchus contortus

The prevalence of antibodies to Neospora caninum was examined in six wild Artiodactyla species, and in five wild Carnivora species from Kenya. Blood sera (104 wild ungulates from Marula Estates (MEs), and 31 wild carnivores from Masai-Mara reserve and from other wildlife areas in northern and Southern Kenya), were screened using a Neospora agglutination test (NAT), with a twofold dilution (1:40-1:320 titres). Presence of NAT antibodies to N. caninun is reported here for the first time in zebra (Equus burchelli), eland (Taurotragus oryx), African buffalo (Syncerus caffer), Thompson gazelle (Gazella thompsoni), impala (Aepyceros melampus), warthog (Phacochoerus aethiopicus), spotted hyena (Crocuta crocuta) and in free-ranging cheetah (Acinonyx jubatus). At 1:80 dilution, prevalence was 61.5% in eland, 58.5% in zebra, 19.2% in Thompson gazelle, 33.3% in warthog, 50% in African buffalo, 30% in lion (Panthera leo), 20% in cheetah, and 33.3% in spotted hyena. Antibodies up to 1:320 titre were detected in eland (38.4%), zebra (19.5%), Thompson gazelle (3.8%) and lion (5%). Amongst herbivores, sero-prevalence was significantly (P<0.05) higher, at all dilutions, in "grazer/digger" species (e.g. eland and zebra) than in non-"grazer/digger" species (e.g. impala and Thompson gazelle). No antibodies to N. caninum were found in two leopards (Panthera pardus) and one serval (Felis serval). Our results indicates a steady presence of N. caninum in wild mammals from Kenya. The hypothesis of a sylvatic cycle of N. caninum could be suggested, but more data are needed to verify the hypothesis, as to evaluate the role of N. caninum infection on the dynamics of wild animals population in the study area.     

The ecological potential of the kudu for commercial production in savanna regions

Abstract

This paper considers the choice of a browsing ungulate to supplement cattle in savanna regions susceptible to problems of bush encroachment, and evaluates the kudu in particular. Relevant aspects of its ecology considered include (i) the vegetation components utilized at different seasons and in different areas, with regard to principal food species, reserve dry season species, rejected plant species and potential food limitations; (ii) social grouping patterns and home range extent; (iii) population densities, reproductive parameters, survival rates and population dynamics in relation to rainfall; (iv) impact on the vegetation of browsing and branch breakage. Features of the kudu are compared with those of other browsing ungulates, including giraffe, eland, impala and goats. It is concluded that the kudu is a prime candidate for inclusion alongside cattle in mixed species ranching enterprises in most regions of savanna vegetation.     

Oviparity in Two South African Skinks of the Genus Mabuya,With Notes on Hatching

This study was conducted as part of the interdisciplinary biodiversity project BIOTA and describes small mammal communities on two differently managed farmlands (cattle and game farm) in Namibia over the course of one year. Species diversity, density and activity are discussed and compared to data on vegetation parameters, and on grazing and trampling effects by large herbivores. Small mammal abundance was related to vegetation cover on both farm areas. Species composition differed between the cattle and the game farm. These differences and the dominance structure are discussed in the view of different grazing techniques of cattle and wild ungulates.     

Cattle Selection for Aspen and Meadow Vegetation: Implications for Restoration

There is concern over the decline of aspen and the lack of successful regeneration due to excessive browsing of aspen suckers by cattle and other wild and domestic ungulates. We conducted a 2-yr study on Lassen National Forest, California, to aid development of cattle grazing strategies to enhance aspen regeneration. We evaluated seasonal biomass, nutritional quality, and utilization by cattle of aspen suckers, aspen herbaceous understory vegetation, and meadow herbaceous vegetation within six aspen–meadow complexes. Aspen suckers had greater nutritional quality compared to aspen understory and meadow vegetation regardless of season or year. Nutritional quality declined with season in all three vegetation types. Early-growing season foraging by cattle focused on meadow and aspen understory vegetation. Mid-growing season decreases in meadow and aspen understory nutritional quality coincided with a marked increase in utilization of aspen suckers. By late-growing season, utilization on aspen suckers was significantly greater than aspen understory or meadow vegetation. Managers can use early-growing season grazing to reduce aspen consumption by cattle, set stocking rates so that adequate herbaceous vegetation is available throughout the growing season, provide nutritional supplements to reduce demand for nutritious aspen suckers, construct protective fencing, and implement grazing systems that insure years with mid- and late-growing season rest from heavy browsing.     

Relationships Between Cattle and Biodiversity in Multiuse Landscape Revealed by Kenya Long-Term Exclosure Experiment

On rangelands worldwide, cattle interact with many forms of biodiversity, most obviously with vegetation and other large herbivores. Since 1995, we have been manipulating the presence of cattle, mesoherbivores, and megaherbivores (elephants and giraffes) in a series of eighteen 4-ha (10-acre) plots at the Kenya Long-term Exclosure Experiment. We recently (2013) crossed these treatments with small-scale controlled burns. These replicated experimental treatments simulate different land management practices. We seek to disentangle the complex relationships between livestock and biodiversity in a biome where worldwide, uneasy coexistence is the norm. Here, we synthesize more than 20 yr of data to address three central questions about the potentially unique role of cattle in savanna ecology: 1) To what extent do cattle and wild herbivores compete with or facilitate each other? 2) Are the effects of cattle on vegetation similar to those of wildlife, or do cattle have unique effects? 3) What effects do cattle and commercial cattle management have on other savanna organisms? We found that 1) Cattle compete at least as strongly with browsers as grazers, and wildlife compete with cattle, although these negative effects are mitigated by cryptic herbivores (rodents), rainfall, fire, and elephants. 2) Cattle effects on herbaceous vegetation (composition, productivity) are similar to those of the rich mixture of ungulates they replace, differing mainly due to the greater densities of cattle. In contrast, cattle, wild mesoherbivores, and megaherbivores have strongly guild-specific effects on woody vegetation. 3) Both cattle and wild ungulates regulate cascades to other consumers, notably termites, rodents, and disease vectors (ticks and fleas) and pathogens. Overall, cattle management, at moderate stocking densities, can be compatible with the maintenance of considerable native biodiversity, although reducing livestock to these densities in African rangelands is a major challenge.     

Farm-level risk factors for the occurrence,new infection or persistence of tuberculosis in cattle herds from South-Central Spain

Eradication of tuberculosis (TB) in cattle is a priority in the EU. However, and despite the resources invested, TB eradication is still a goal yet to be accomplished. As a consequence, the identification of risk factors contributing to TB transmission and persistence is key to a better understand and, ultimately, more cost-effectively control and eradicate this complex zoonotic disease worldwide. This study aimed to identify the factors contributing to the TB occurrence, new infection, and persistence in one of the most TB-prevalent regions in the South-Central part of Spain (SCS), Ciudad Real (CR), and for which high quality and detailed information on cattle and wild ungulate demographics, management, and sanitary status was available for up to 5 years (2007–2011). Multilevel logistic modeling was used for this purpose. Results of this study revealed that the risk for TB (occurrence, new infection, and/or persistence) in cattle herds from CR is related to TB-persistence on farm in previous years, extensive production systems (beef and bullfighting being more risky that dairy) and large farm in terms of cattle number. Also, the presence and proximity of fenced hunting estates (which are usually intensively managed) significantly contributed to the risk of TB occurrence and persistence in CR. This association suggests that wild ungulates may play a role as TB-reservoirs and transmit TB to cattle in the CR, a region where TB is endemic and is associated with extensive farming of beef cattle. To the best of author's knowledge this is the first study to (i) evaluate the wild ungulate-domestic interface at a farm level using detailed information over an extensive region and period of time (2007–2011) and (ii) to identify a direct association between TB in cattle and the proximity and management practices of wild ungulates. Methods and results presented here may support policies to better prevent and control TB in the SCS and in other regions/countries with similar epidemiological conditions.     

Postfire Succession in Big Sagebrush Steppe With Livestock Grazing

Prescribed fire in rangeland ecosystems is applied for a variety of management objectives, including enhancing productivity of forage species for domestic livestock. In the big sagebrush (Artemisia tridentata Nutt.) steppe of the western United States, fire has been a natural and prescribed disturbance, temporarily shifting vegetation from shrub–grass codominance to grass dominance. There is limited information on the impacts of grazing to community dynamics following fire in big sagebrush steppe. This study evaluated cattle grazing impacts over four growing seasons after prescribed fire on Wyoming big sagebrush (Artemisia tridentata subsp. Wyomingensis [Beetle & Young] Welsh) steppe in eastern Oregon. Treatments included no grazing on burned and unburned sagebrush steppe, two summer-grazing applications after fire, and two spring-grazing applications after fire. Treatment plots were burned in fall 2002. Grazing trials were applied from 2003 to 2005. Vegetation dynamics in the treatments were evaluated by quantifying herbaceous canopy cover, density, annual yield, and perennial grass seed yield. Seed production was greater in the ungrazed burn treatments than in all burn–grazed treatments; however, these differences did not affect community recovery after fire. Other herbaceous response variables (cover, density, composition, and annual yield), bare ground, and soil surface litter did not differ among grazed and ungrazed burn treatments. All burn treatments (grazed and ungrazed) had greater herbaceous cover, herbaceous standing crop, herbaceous annual yield, and grass seed production than the unburned treatment by the second or third year after fire. The results demonstrated that properly applied livestock grazing after low-severity, prescribed fire will not hinder the recovery of herbaceous plant communities in Wyoming big sagebrush steppe.     

Livestock Riparian Guidelines May Not Promote Woody Species Recovery Where Wild Ungulate Populations Are High

Stubble height, streambank alteration, and woody species use are indicators used to monitor livestock impacts on riparian areas in the western United States. Effects of wild ungulates on riparian conditions are often not monitored and assumed to be represented by indicators developed for livestock. We tested this assumption by evaluating effects of elk (Cervus canadensis) and mule deer (Odocoileus hemionus) on grazing indicators along Meadow Creek, a salmonid-bearing stream in northeastern Oregon. Wild ungulates reduced stubble height by 20% to 30%. Mean streambank alteration was 1.1% (ranged from 0.3–8%). Woody species use was negatively related to stubble height and positively related to streambank alteration (P < 0.05). Despite maintenance of stubble height and streambank alteration within regulatory guidelines, wild ungulate use of preferred woody species was moderate to high (> 50%). Adherence to guidelines developed for livestock may not result in desired riparian conditions where wild ungulate populations are high.     

Behavioral and Ecological Implications of Bunched,Rotational Cattle Grazing in East African Savanna Ecosystem

Rangelands and the wildlife and livestock they support are critical to human livelihoods, but rangeland ecosystems increasingly suffer from overgrazing and degradation. Planned grazing, a strategy that commonly involves time-controlled rotations of high-density (bunched) groups of cattle across a pasture, is marketed as a method to enhance rangeland health and lessen livestock impacts. However, the behavioral mechanisms underlying any potential rangeland improvements resulting from rotational, high-density planned grazing have rarely been examined. To investigate these mechanisms, we compared planned grazing with conventional continuous grazing management in a savanna ecosystem in Kenya. We surveyed cattle grazing behavior, measured changes in vegetation characteristics through surveys conducted before and after cattle grazing, and measured native ungulate abundance following grazing using camera traps. Stocking rates were held constant across treatments, resulting in a commensurate decline in total foliar hits per pin (a proxy for vegetative biomass) across treatments. Planned grazing management altered cattle behavior and reduced grazing selectivity by restricting movements, causing cattle to walk more slowly while grazing and to take more bites per step. Vegetation survey results supported this finding: cattle in the planned grazing treatment ate significantly more Pennisetum grasses (typically avoided because of their unpalatability), creating the opportunity for regrowth of more palatable species after seasonal rains. We also documented significantly higher zebra presence in planned grazing plots after cattle grazing, likely due to increased relative abundance of more palatable grass species. This investigation of grazing behavior, and specifically decreased grazing selectivity as a mechanism underpinning the benefits of planned grazing, shows that when conducted at appropriate stocking densities, planned grazing has the potential to help mitigate rangeland degradation and improve rangeland sustainability for both livestock and wildlife in pastoral African savanna ecosystems.     

放牧牛羊对云贵地区禾草/白三叶草地群落稳定性的影响

禾草/白三叶(Trifolium repens)草地是中国南方喀斯特地区主要刈牧地之一,探究不同家畜放牧下禾草/白三叶草地植被构成和群落稳定性,可一定程度揭示其植被演替特征。本研究定量分析了多年放牧牛(grazing cattle pasture,GC)、放牧羊(grazing sheep pasture,GS)的禾草/白三叶草地群落特征、物种多样性、生物量构成及群落稳定性。结果显示:1)草地死物质量及植物物种数为GC>GS,植物物种Simpson指数为GS>GC;2)草地禾草[多年生黑麦草(Lolium perenne)、鸭茅(Dactylis glomerata)及原生禾草]、原生双子叶(马鞭草科、车前科、酢浆草科和十字花科)及适口性中等植物地上生物量组分为GC>GS,播种白三叶地上生物量组分为GS>GC;且GC草地地上生物量以禾草为主(77.20%),GS草地地上生物量以禾草(51.79%)和豆科(28.02%)为主;3)草地土壤K含量为GC>GS;4)GC草地植物种Raunkiaer频度直方图呈反J型,群落演替度(37.08)比GS草地(45.45)的低。研究认为,GC草地群落处于相对稳定状态,而GS草地群落处于更高演替阶段,牛放牧更利于禾草/白三叶草地群落稳定性和物种多样性的维持。因此,放牧时可实行牛、羊混牧制,或牛、羊轮牧制,以提高禾草/白三叶草地的稳定性和生产力。     

Effects of Wildlife on Cattle Diets in Laikipia Rangeland,Kenya

The impacts of wild herbivores on cattle diet selection were investigated in an East African rangeland during August 2001 and February 2002. The study compared cattle diets in plots exclusively accessible to cattle (C) and those accessible to megaherbivores (elephants and giraffes), non-megaherbivore wild herbivores > 15 kg (zebras, hartebeests, Grant's gazelles, oryx, elands, and buffaloes) and cattle (MWC); or non-megaherbivore wild herbivores and cattle (WC). There were no treatment differences in selection of most grass species in either sampling period (P > 0.05). However, selection of forbs differed among treatments during February when conditions were relatively dry and percent of bites taken by cattle on this forage class increased (P < 0.005) from 1.8% ± 0.3 to 7.7% ± 1.6 (mean ± SE). During this period, cattle took a lower percent of bites on forbs in MWC (4.3% ± 1.7; P = 0.01) and WC (5.9% ± 2.2; P = 0.03) than in C (12.9% ± 0.9). These patterns were generally driven by Commelina spp., which comprised 65% ± 9.4 of total bites on forbs. Notably, these differences were associated with differences in cover of forbs, which was positively correlated with percent of bites on forbs (r² = 0.86, P < 0.01). Because forbs may be critical components of cattle diets in such rangelands during relatively dry periods, these dietary changes may indicate potential seasonal costs of wildlife to cattle production. Looking for ways to offset such costs may be worthwhile for livestock properties that accommodate wildlife.     

Seasonal changes in biomass concentration in three coastal grassland communities in Transkei

The density of forage available to wild ungulates was measured in terms of biomass concentration. Data were collected in three grassland communities for several months. In two of the communities burning and grazing treatments were also applied. The biomass concentration was greatest in the shortest sward. Results indicate that after burning biomass concentrations were temporarily below the level required by a LSU to maintain body mass. This temporary shortage existed for 5–6 months after a fire. This may be partly responsible for the increased grazing frequently observed following a fire.     

Cattle Grazing Fails to Control Shrub Encroachment in Mediterranean Landscapes

The Common Agricultural Policy supports the use of free-ranging cattle herds to control woody encroachment and fire hazards in Europe. There is, however, little empirical evidence about the effectiveness of extensive grazing to preserve open landscapes in the Mediterranean Basin. In this work, we evaluated the effects of extensive beef cattle grazing on the vegetation structure in a Mediterranean ecosystem using a twofold framework: 1) analyzing temporal changes in the forest, shrub, and grassland cover in areas under different grazing pressures for 16 yr (1993 ? 2009) and 2) studying diet selection to assess the impact of cattle on the local Mediterranean vegetation. Our landscape structure analyses revealed a remarkable change in land cover over the study period. However, woody community dynamics seemed to be more related to natural vegetation succession than to cattle effects. Extensive grazing seemed to preserve grasslands but only at high stocking rates. On the other hand, the diet analyses supported the lack of a role for cattle in encroachment control. Beef cattle diets were based on herbaceous plants (59%) with lower contribution of woody ones (41%). Cattle only showed a significant preference (P < 0.05) for few woody species (Erica multiflora, Olea europaea, Quercus ilex, and Rosmarinus officinalis), mostly at high-density stocking rates. Hence, our results support the idea that extensive cattle grazing alone exerts a negligible effect on shrub encroachment and thus on the risk of fire in the studied Mediterranean area. We urge a redesign of current research to truly integrate extensive cattle grazing as High Nature Value farming in European policies to successfully meet its putative goals, such as shrub encroachment control and wildfire risk prevention.     

Vigilance in Cattle: The Influence of Predation,Social Interactions,and Environmental Factors

Vigilant behavior in wild ungulates is critical to guard against predation. However, few studies have examined vigilant behavior in domesticated ungulates. Considering the expansion of large predator populations, understanding vigilant behavior and factors that influence it will help with the management of livestock. We observed adult female cattle (Bos taurus L.) in open-range conditions where large predators (wolves [Canis lupus L.] and mountain lions [Puma concolor (L.).]) were common threats during summers of 2005 and 2006 in eastern Arizona. This study was designed to determine 1) to what extent cattle exhibit vigilant behavior compared to published data on wild ungulates, 2) whether predation events influence vigilance rates of cattle, and 3) whether social and environmental factors affect vigilance of cattle. Cattle exhibited vigilant behavior (3% ± 0.19%) during peak foraging periods, but at a lower rate than wild ungulates. Cows with calves were more than twice as vigilant (4.5% ± 0.46%) as those without calves (2.0% ± 0.27%). Single cattle and groups of two to five exhibited higher vigilance rates (4.2% ± 0.79%) than groups of six to 20 (2.5% ± 0.32%) and groups of > 20 (3.0% ± 0.41%). Cattle in groups of > 20 increased vigilance as visual obstruction increased. Mother cows whose calves were preyed upon (n = 5) exhibited a 3% to 48% increase in vigilance within 3 d after their calves were killed; this rate returned to near baseline levels after 10 d. Conversely, mother cows reduced foraging after their calves were killed from 88.5% ± 1.69% to 43.5% ± 11.4%; foraging rate also returned to near baseline levels after 10 d. Cattle exhibit vigilance at lower levels compared to wild ungulates, but this behavior appears to be at least partially an antipredatory behavior. Our findings provide support that predators can influence cattle behavior.     

Influence of Forest Management and Previous Herbivory on Cattle Diets

Grazing cattle and timber harvest are common practices associated with forested rangelands. Therefore, the objective was to document the effects of timber harvest and herbivory on nutritional quality and botanical composition of steer diets in grand fir (Abies grandis [Dougl. ex D. Don] Lindl.) and ponderosa pine (Pinus ponderosa P. & C. Lawson) forests. Three replicated grand fir sites were arranged as a split-plot design; timber harvest treatments—1) no harvest (CON), 2) thinning (TH), 3) clearcut (CL)—were whole plots, and herbivory treatments—1) large ungulate grazing (Graze), 2) wild ungulate grazing (CExc), and 3) exclusion of large ungulate grazing (TExc)—were the subplots. Three replicated ponderosa pine sites were arranged as a split-plot design; timber harvest treatments—1) CON and 2) TH—were whole plots, and herbivory treatments—1) GR, 2) BG, and 3) EX—were subplots. Diet samples were collected in June and August of 2001 and 2002. Crude protein, in vitro organic matter digestibility, acid detergent fiber, and neutral detergent fiber of the diets were only affected by season of use and were higher (P < 0.05) quality during the June grazing period. Botanical composition of diets was determined with the use of microhistological analysis of ruminal masticate. Within grand fir sites, graminoids were the major constituent in the diet (65%–91%), forbs intermediate (8%–31%), and shrubs least (0.2%–3.5%). Within ponderosa pine sites graminoids were the major constituent in the diet (83%–88%), forbs intermediate (10%–14%), and shrubs least (2%–3%). Season of use did not affect (P > 0.10) botanical composition in either grand fir or ponderosa pine sites. Timing of grazing had a greater influence on diet quality; however, previous herbivory and(or) timber harvest had a greater influence on composition of diets than did timing of grazing.     

Initial Effects of Brush Cutting and Shoot Removal on Willow Browse Quality

We examined the initial effects of brush cutting (removal of all aboveground biomass), as well as clipping (removal of current annual shoots) and ungulate browsing (collectively referred to as shoot removal) on the morphology and nutrient quality of Scouler's willow (Salix scouleriana J. Barratt ex Hook.) for ungulates on sites 2 and 4 years after brush cutting. We specifically assessed changes in the biomass, tannin content, digestible energy, and digestible protein of shoots from brush-cut willows relative to shoots of uncut willows to determine how browse plants respond to this form of vegetation management. In winter, the resprouted current annual shoots of willows that had been brush cut were larger in mass and lower in digestible protein than shoots of uncut willows for at least 4 years after brush cutting. Shoots of brush-cut willows were also lower in tannin and digestible energy than the shoots of uncut plants for two winters after brush cutting. In the second winter after brush cutting, shoot biomass decreased and tannin content increased with increasing shoot removal during the previous winter. In the fourth winter after brush cutting, shoot mass increased and digestible energy decreased in shoots with greater shoot removal. Nutrient quality was otherwise unaffected by the amount of shoot removal during the previous winter. Because of the occasional importance of site effects in this study, we recommend that long-term studies maximize the number of sampled sites. Because brush cutting alters the quality of regenerating browse and can affect how ungulates utilize such browse for several years after brush cutting, we further recommend that forest vegetation managers consider potential impacts of brush cutting on ungulate winter range.     

Interspace/Undercanopy Foraging Patterns of Beef Cattle in Sagebrush Habitats

Forage selection patterns of cattle in sagebrush (Artemisia L.) communities are influenced by a variety of environmental and plant-associated factors. The relative preference of cattle for interspace versus under-sagebrush canopy bunchgrasses has not been documented. Potential preferences may indirectly affect habitat for sage-grouse and other ground-nesting birds. Our objectives were to investigate grazing patterns of cattle with respect to undercanopy (shrub) and interspace tussocks, determine the influence of cattle grazing on screening cover, and relate shrub morphology to undercanopy grazing occurrence. Eighteen-day replicated trials were conducted in the summers of 2003 and 2004. Findings suggest cattle initially concentrate grazing on tussocks between shrubs, and begin foraging on tussocks beneath shrubs as interspace plants are depleted. Grazing of undercanopy grass tussocks was negligible at light-to-moderate utilization levels (< 40% by weight). Grass tussocks under spreading, umbrella-shaped shrub canopies were less likely (P < 0.001) to be grazed than those beneath erect, narrow canopies. Horizontal screening cover decreased (P < 0.001) with pasture utilization. At the trial’s end, removal of 75% of the herbaceous standing crop induced about a 5% decrease in screening cover in all strata from ground level to 1 m with no differences among strata (P = 0.531). This implied that shrubs constituted the majority of screening vegetation. Our data suggest that conservative forage use, approaching 40% by weight, will affect a majority (about 70%) of interspace tussocks and a lesser proportion (about 15%) of potential nest-screening tussocks beneath sagebrush. Probability of grazing of tussocks beneath shrubs, however, is also affected by shrub morphology. These findings will help managers design grazing programs in locales where habitat for ground nesting birds is a concern.