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1.
Denitrification rates in soils of six subalpine plant communities in an eastern Sierra Nevada watershed were determined by the acetylene inhibition method. Soil atmosphere samples were collected monthly from June 1986 through May 1987 in a riparian, wet meadow, dry meadow, north-facing forest, south-facing forest and barren site and analyzed for N2O content using gas chromatography. Soil temperature, moisture, organic matter, C,N,C:N ratio, NO3N and pH were examined to assess their effects on denitrification rates. Mean denitrification rates for the year varied from 103.3 μg m−2h−1 in the north-facing forest to 120.2 μg m−2h−1 at the barren site, but did not differ significantly among any of the six plant communities. However, comparisons among months within individual communities revealed that the denitrification rates in each community varied significantly over the year, and in three of the six sites significant correlations between denitrification rates and other soil parameters were detected. Soil acidity was positively correlated with denitrification rate in the riparian and wet meadow communities, and in the dry meadow, soil moisture was positively correlated while soil temperature and organic matter were negatively correlated with denitrification rate. Comparisons among sites within individual months revealed significant differences in denitrification rates in June, September, October and January, but no single site consistently exhibited the highest or lowest rate in all 4 months, and only in October, when denitrification rates were positively correlated with soil temperature and moisture, was variation in denitrification rates among sites explained by other soil parameters. For the six plant communities overall, soil denitrification rates were highly variable from June to October, increased sharply from October to December, and then declined from December to May.  相似文献   

2.
The capacity of prescribed fire to restore forest conditions is often judged by changes in forest structure within a few years following burning. However, prescribed fire might have longer-term effects on forest structure, potentially changing treatment assessments. We examined annual changes in forest structure in five 1 ha old-growth plots immediately before prescribed fire and up to eight years after fire at Sequoia National Park, California. Fire-induced declines in stem density (67% average decrease at eight years post-fire) were nonlinear, taking up to eight years to reach a presumed asymptote. Declines in live stem biomass were also nonlinear, but smaller in magnitude (32% average decrease at eight years post-fire) as most large trees survived the fires. The preferential survival of large trees following fire resulted in significant shifts in stem diameter distributions. Mortality rates remained significantly above background rates up to six years after the fires. Prescribed fire did not have a large influence on the representation of dominant species. Fire-caused mortality appeared to be spatially random, and therefore did not generally alter heterogeneous tree spatial patterns. Our results suggest that prescribed fire can bring about substantial changes to forest structure in old-growth mixed conifer forests in the Sierra Nevada, but that long-term observations are needed to fully describe some measures of fire effects.  相似文献   

3.
Typically, after large stand-replacing fires in mid-elevation Sierra Nevada forests, dense shrub fields occupy sites formerly occupied by mature conifers, until eventually conifers overtop and shade out shrubs. Attempting to reduce fuel loads and expedite forest regeneration in these areas, the USDA Forest Service often disrupts this cycle by the logging of fire-killed trees, replanting of conifers and killing of shrubs. We measured the effects of these treatments on live and dead fuel loads and alien species and modeled potential fire behavior and fire effects on regenerating forests. Sampling occurred in untreated, logged and herbicide-treated stands throughout the Sierra Nevada in four large fire areas 4–21 years after stand-replacing fires. Logging fire-killed trees significantly increased total available dead fuel loads in the short term but did not affect shrub cover, grass and forb cover, alien species cover or alien species richness. Despite the greater available dead fuel loads, fire behavior was not modeled to be different between logged and untreated stands, due to abundant shrub fuels in both logged and untreated stands. In contrast, the herbicide treatment directed at shrubs resulted in extremely low shrub cover, significantly greater alien species richness and significantly greater alien grass and forb cover. Grass and forb cover was strongly correlated with solar radiation on the ground, which may be the primary reason that grass and forb cover was higher in herbicide treated stands with low shrub and tree cover. Repeat burning exacerbated the alien grass problem in some stands. Although modeled surface fire flame lengths and rates of spread were found to be greater in stands dominated by shrubs, compared to low shrub cover conifer plantations, surface fire would still be intense enough to kill most trees, given their small size and low crown heights in the first two decades after planting.  相似文献   

4.
Fire plays an important role in shaping many Sierran coniferous forests, but longer fire return intervals and reductions in area burned have altered forest conditions. Productive, mesic riparian forests can accumulate high stem densities and fuel loads, making them susceptible to high-severity fire. Fuels treatments applied to upland forests, however, are often excluded from riparian areas due to concerns about degrading streamside and aquatic habitat and water quality. Objectives of this study were to compare stand structure, fuel loads, and potential fire behavior between adjacent riparian and upland forests under current and reconstructed active-fire regime conditions. Current fuel loads, tree diameters, heights, and height to live crown were measured in 36 paired riparian and upland plots. Historic estimates of these metrics were reconstructed using equations derived from fuel accumulation rates, current tree data, and increment cores. Fire behavior variables were modeled using Forest Vegetation Simulator Fire/Fuels Extension.Riparian forests were significantly more fire prone under current than reconstructed conditions, with greater basal area (BA) (means are 87 vs. 29 m2/ha), stand density (635 vs. 208 stems/ha), snag volume (37 vs. 2 m3/ha), duff loads (69 vs. 3 Mg/ha), total fuel loads (93 vs. 28 Mg/ha), canopy bulk density (CBD) (0.12 vs. 0.04 kg/m3), surface flame length (0.6 vs. 0.4 m), crown flame length (0.9 vs. 0.4 m), probability of torching (0.45 vs. 0.03), predicted mortality (31% vs. 17% BA), and lower torching (20 vs. 176 km/h) and crowning indices (28 vs. 62 km/h). Upland forests were also significantly more fire prone under current than reconstructed conditions, yet changes in fuels and potential fire behavior were not as large. Under current conditions, riparian forests were significantly more fire prone than upland forests, with greater stand density (635 vs. 401 stems/ha), probability of torching (0.45 vs. 0.22), predicted mortality (31% vs. 16% BA), and lower quadratic mean diameter (46 vs. 55 cm), canopy base height (6.7 vs. 9.4 m), and frequency of fire tolerant species (13% vs. 36% BA). Reconstructed riparian and upland forests were not significantly different. Our reconstruction results suggest that historic fuels and forest structure may not have differed significantly between many riparian and upland forests, consistent with earlier research suggesting similar historic fire return intervals. Under current conditions, however, modeled severity is much greater in riparian forests, suggesting forest habitat and ecosystem function may be more severely impacted by wildfire than in upland forests.  相似文献   

5.
The fisher (Martes pennanti) is a forest mustelid endemic to North America that has experienced range reductions in Pacific states that have led to their listing under the Endangered Species Act as warranted but precluded by higher priorities. The viability of the southern Sierra Nevada fisher population is of particular concern due to its reduced historical range, isolated nature, and low genetic variability. We located resting structures of radio-collared fishers in the southern Sierra Nevada and compared resting and available habitat to examine selection for specific features of resting sites. Resting structures provide protection from predators and unfavorable weather and are believed to be the most limiting habitat element across fisher home ranges. Resting structures were found primarily in live trees (76%) and snags (15%). Trees used by fishers for resting were among the largest available and frequently had mistletoe infestations. Ponderosa pines (Pinus ponderosa) were used more often than expected and incense cedars (Calocedrus decurrens) less than expected. Snags were also large and in fairly advanced stages of decay. Habitat at fisher resting sites had higher canopy cover, greater basal area of snags and hardwoods, and smaller and more variable tree sizes compared to random sites. Resting sites were also found on steeper slopes and closer to streams. Canopy cover was consistently the most important variable distinguishing rest and random sites. In western North America, fishers are generally associated with late-successional forests, but changes in these forests due to logging and fire suppression have resulted in a transition to forest stands characterized by fewer large trees and more small stems. These conditions are consistent with our finding that the large rest structures were surrounded by smaller than average trees. Management practices that support the growth and retention of greater numbers of large trees and snags, while maintaining a minimum of 61% (based on moosehorn) or 56% (generated via Forest Vegetation Simulator) canopy cover and a complex horizontal and vertical forest structure, can improve and provide for future fisher habitat.  相似文献   

6.
We review some of the unique features of biogeochemical cycling in forests of the eastern Sierra Nevada Mountains, USA. As is the case for most arid and semi-arid ecosystems, spatial and temporal variability in nutrient contents and fluxes are quite high. “Islands of fertility” are common in these forests, a result of spatial variations in both litterfall and decomposition rates. Dry summer conditions greatly inhibit biological activity in the O horizon, and thus most annual litter decomposition takes place beneath the snowpack when moisture is available. Snowmelt duration is shortened near tree boles because of local warming, resulting in earlier drying of the O horizon, significantly lower decomposition rates, and increased O horizon mass. Water and nutrient fluxes vary spatially because of snowdrift in winter and surface runoff over hydrophobic soils in summer and fall. Moisture variability in the vertical as well as the horizontal dimension has significant consequences for nutrient fluxes. Because of the very dry summers, rooting in the O horizons is absent in these forests, and thus competition between microbes and trees for nutrients in that horizon is non-existent. Nutrients mineralized from the O horizon and not taken up by plants enrich runoff through the O horizons over hydrophobic mineral soils, resulting in very high concentrations of inorganic N and P in runoff waters. Substantial temporal variations in water and nutrient fluxes occur on a seasonal (with snowmelt being the dominant hydrologic event of the year), annual, and decadal basis. The most significant temporal variation is due to periodic fire, which we estimate causes annualized N losses that are two orders of magnitude greater than those associated with leaching and runoff. We hypothesize that fire suppression during the 20th century may have contributed to the deterioration of nearby Lake Tahoe by allowing buildups of N and P in O horizons which could subsequently leach from the terrestrial ecosystem to the Lake in runoff. In general, we conclude that biogeochemical cycling in these forests is characterized by greater spatial and temporal variability than in more mesic forest ecosystems.  相似文献   

7.
Urban development typically has extensive and intensive effects on native ecosystems, including vegetation communities and their associated biota. Increasingly, urban planning strives to retain elements of native ecosystems to meet multiple social and ecological objectives. The ecological integrity of native forests in an urbanizing landscape is challenged by a myriad of impacts, such as forest management and invasive species. Environmental protection efforts in the Lake Tahoe basin, spanning the California/Nevada border in the Sierra Nevada mountains, over the past half century have resulted in the retention of thousands of parcels of remnant native forest located throughout the urbanizing landscape. The basin landscape provides an opportunity to evaluate the effects of land development on the composition and structure of remnant native forests along a gradient of urbanization. We sampled 118 sites located in remnant forests in the lower montane zone surrounded by 0–70% development. We also sampled forest structure in the landscape surrounding 75 of these sites to evaluate the contribution of remnant forests to the retention of native forest elements in the larger landscape. We characterized plant species composition and cover, vertical structure, and the density of trees, snags, and logs, as well as levels of ground disturbance and human activity. We found that remnant native forests retained much of their compositional and structural character along the development gradient, including large tree density, total canopy cover, and plant species richness. Notable exceptions were reductions in the density and decay stage of snags and logs, and the density of understory trees. We also observed increases in the richness and cover of herb and grass species and increases in the number of exotic plant species. In contrast, structural complexity was reduced in the landscape surrounding forest remnants in all measures except large tree density. We conclude that remnant native forests contribute significantly to maintaining native species in an urbanizing landscape, and that land conservation practices have an important role to play in protecting native forest ecosystems.  相似文献   

8.
Xu M  DeBiase TA  Qi Y  Goldstein A  Liu Z 《Tree physiology》2001,21(5):309-318
We estimated total ecosystem respiration from a ponderosa pine (Pinus ponderosa Dougl. ex Laws.) plantation in the Sierra Nevada Mountains near Georgetown, California, from June to October, 1998. We apportioned ecosystem respiration among heterotrophic, root, stem and foliage based on relationships for each component that considered microclimate and vegetation characteristics. We measured each respiration component at selected sampling points, and scaled the measurements up to the ecosystem based on modeled relationships. Over the study period, total mean ecosystem respiration was 5.7 +/- 1.3 mumol m-2 s-1 (based on daily mean), comprising about 67% from soil-surface CO2 efflux, 10% from stem and branch respiration and 23% from foliage respiration. Shrub leaves contributed about 24% to total foliage respiration, and current-year needles (1998 age class) accounted for 40% of total tree needle respiration. Root respiration accounted for 47% of soil-surface CO2 efflux. We conclude that ecosystem respiration can be estimated based on daily mean air and soil temperatures through exponential relationships with r2 values of 0.85 and 0.87, respectively. When based on both air and soil temperatures, about 91% of the variation in total ecosystem respiration could be explained by a linear regression.  相似文献   

9.
More than a century of fire exclusion and past timber management practices in many Sierra Nevada mixed-conifer forests have led to increased stand densities and fuel accumulation, with a corresponding risk of large, high severity wildfires. To reduce hazardous fuel accumulations and restore the health and natural processes of forest ecosystems, fuel management programs often employ thinning and prescribed fire treatments, both alone and in combination. We evaluated forest floor and mineral soil chemical and physical characteristics following these treatments in a managed Sierra Nevada mixed-conifer forest using a fully replicated study design with four separate treatments: THIN, BURN, THIN + BURN, and an untreated CONTROL. Compared to the CONTROL, the BURN and THIN + BURN treatments consumed a large amount of the forest floor, reducing the mass and depth by more than 80%. These treatments reduced the forest floor C and N pools by more than 85%, resulting in reductions of 25 Mg C ha−1 and more than 700 kg N ha−1 from the forest floor. Despite these large losses from the organic horizons, no significant differences in mineral soil total C and N pools were detected among treatments. Compared with the CONTROL and THIN treatments, the BURN and THIN + BURN significantly increased the mineral soil NO3-N concentration, pool of inorganic N, pH, and exposed bare soil. The THIN + BURN treatment significantly increased the concentrations of NH4-N and exchangeable Ca relative to the CONTROL. No significant differences in the net rates of nitrification, N mineralization, or bulk density were detected among the four treatments. The BURN treatment reduced mineral soil C concentration and CEC, while the THIN + BURN treatment had the greatest increase in inorganic N. Fire effects on soil pH and inorganic N were moderated in skid trails due to reduced fuel continuity and consumption. In light of the current management emphasis on hazardous fuels reduction, we recommend that researchers investigating fire effects in harvested stands include skid trail influences in their study design.  相似文献   

10.
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12.
Due to increases in tree density and hazardous fuel loading in Sierra Nevadan forests, land management is focusing on fuel reduction treatments to moderate the risk of catastrophic fires. Fuel treatments involving mechanical and prescribed fire methods can reduce surface as well as canopy fuel loads. Mastication is a mechanical method which shreds smaller trees and brush onto the surface fuel layer. Little data exist quantifying masticated fuel beds. Despite the paucity of data on masticated fuels, land managers desire fuel loading, potential fire behavior and fire effects such as tree mortality information for masticated areas. In this study we measured fuel characteristics before and after mastication and mastication plus prescribed burn treatments in a 25-year old ponderosa pine (Pinus ponderosa C. Lawson) plantation. In addition to surface fuel characteristics and tree data collection, bulk density samples were gathered for masticated material. Regressions were created predicting masticated fuel loading from masticated fuel bed depth. Total masticated fuel load prior to fire treatment ranged from 25.9 to 42.9 Mg ha−1, and the bulk density of masticated fuel was 125 kg m−3. Mastication treatment alone showed increases in most surface fuel loadings and decreases in canopy fuel loads. Masticated treatment in conjunction with prescribed burning reduced both surface and canopy fuel loads. Detailed information on fuel structure in masticated areas will allow for better predictions of fire behavior and fire effects for fire in masticated fuel types. Understanding potential fire behavior and fire effects associated with masticated fuels will allow managers to make decisions on the possibility of mastication to create fuel breaks or enhance forest health.  相似文献   

13.
Understanding seasonal variations of photosynthetic parameters is critical for accurate modeling of carbon dioxide (CO2) uptake by ecosystems. Maximum carboxylation velocity (Vcmax), maximum rate of electron transport (Jmax), leaf respiration in the light (R(day)), light-saturated assimilation (Amax) and maximum quantum yield (Phi) were calculated from leaf gas exchange measurements made monthly throughout the year on leaves of three co-occuring evergreen species in a Pinus ponderosa Dougl. ex P. Laws. & C. Laws. forest with shrubs in the understory (Arctostaphylos manzanita Parry and Ceanothus cordulatus Kellogg.). The seasonality and relationships of the photosynthetic parameters with environmental and physiological variables differed among the species. The nitrogen-fixing species, C. cordulatus had the highest values of the parameters and the largest seasonal variation, whereas A. manzanita exhibited the lowest seasonality and weaker correlations with environmental variables. In general, variations in Vcmax were highly correlated with light, leaf mass per area and leaf nitrogen content on an area basis. Temporal scaling of the parameters with each other seemed possible for C. cordulatus and P. ponderosa. However, lags between these variables and Vcmax likely reflect the influences of other factors. The acclimation relationships found along vertical light gradients within canopies in other studies cannot be applied to seasonal variations. The Jmax to Vcmax ratio varied seasonally for P. ponderosa and A. manzanita, being lower at high light, high air temperature and low soil water content.  相似文献   

14.
We summarized the status of wolves (Canis lupus), elk (Cervis elaphus), and woody browse conditions during the 20th century for the upper Gallatin elk winter range in southwestern Montana, USA. During this period, wolves were present until about the mid-1920s, absent for seven decades, and then returned to the basin in 1996. A chronosequence of photographs, historical reports, and studies indicated willows (Salix spp.) along streams became heavily browsed and eventually suppressed following the removal of wolves, apparently due to unimpeded browsing by elk. However, after wolf establishment in 1996, browsing intensity on willows lessened in some areas and we hypothesized that, at both a landscape and fine scale, browsing pressure reflects terrain configurations influencing predation risk (nonlethal effects), in conjunction with lower elk densities (lethal effects). We measured browsing intensity and heights of Booth willow (S. boothii) along 3000 m reaches of the Gallatin River and a tributary to examine the potential influence of wolf/elk interactions upon willow growth. Where the Gallatin Valley is relatively narrow (high predation risk), willows began releasing in 1999 and by 2002 were relatively tall (150–250 cm). In contrast, willow heights along a wider portion of the Gallatin Valley, along the open landscape of the tributary, and an upland site (all low predation risk) generally remained low (<80 cm). We identified terrain and other features that may contribute to the perceived risk of wolf predation, by elk for a given site. Although alternative mechanisms are discussed, changes in willow communities over time following wolf removal and their subsequent reintroduction were consistent with a top-down trophic cascade model involving nonlethal and possibly lethal effects. If similar top-down effects upon vegetation hold true in other regions of North America and other parts of the world where wolves have been extirpated, wolf recovery may represent a management option for helping to restore riparian plant communities and conserve biodiversity.  相似文献   

15.
The northern and central Appalachian forests are subject to high levels of atmospheric acid deposition (AD), which has been shown in some forests to negatively impact forest growth as well as predispose the forest system to damage from secondary stresses. The purpose of this study was to evaluate the possible contribution of AD to changes in composition and productivity of the Monongahela National Forest, and to evaluate soil-based indicators of acidification that might be useful for detecting AD-related forest changes. Soils adjacent to 30 Forest Inventory and Analysis (FIA) sites were sampled and analyzed for a suite of acidity indicators. These indicators were correlated with the periodic mean annual volume increment (PMAVI) of the forest stands on FIA plots for the 10-yr period 1989–2000. PMAVI ranged from −9.5 to 11.8 m3 ha−1 yr−1, with lower-than-expected growth (<3 m3 ha−1 yr−1) on two-thirds of the sites. In the surface horizon, effective base saturation, Ca2+ concentration, base saturation, K+ concentration, Ca/Al molar ratio, and Mg/Al molar ratio, were positively correlated with PMAVI and Fe concentration was negatively correlated with PMAVI (p ≤ 0.1). In the subsurface horizon pH(w) and effective base saturation were positively correlated and Al3− concentration and K+ concentration were negatively correlated with PMAVI. We hypothesized that NO3-N/NH4-N ratio would also be correlated with PMAVI, but it was not. Correlations between soil chemical indicators and PMAVI suggest that AD may contribute, in part, to the lower-than-expected forest growth on the Monongahela National Forest.  相似文献   

16.
Fire is an important ecological process in many western U.S. coniferous forests, yet high fuel loads, rural home construction and other factors have encouraged the suppression of most wildfires. Using mechanical thinning and prescribed burning, land managers often try to reduce fuels in strategic areas with the highest fuel loads. Riparian forests, however, are often designated as areas where only limited management action can take place within a fixed-width zone. These highly productive forests have developed heavy fuel loads capable of supporting stand-replacing crown fires that can alter wildlife habitat and ecosystem function, and contribute to stream channel erosion. Objectives of this study were to determine whether adjacent coniferous riparian and upland forests burned historically with different frequencies and seasonalities, and whether these relationships varied by forest, site, and stream characteristics. We measured dendrochronological fire records in adjacent riparian and upland areas across a variety of forest, site and stream conditions at 36 sites in three sampling areas in the northern Sierra Nevada.  相似文献   

17.
通过连续2年的外业调查和实验室分析,研究了持续干旱背景下西山国家森林公园的华山松、云南油杉、地盘松、麻栎和旱冬瓜等5个主要林型的可燃物特征,结果表明:与2012年的载量相比较,华山松林的载量增加了2.68kg · m^ -2,增加59.16%,地盘松林的载量增加了4.06kg · m^ -2,增加1.28倍,增长很明显;云南油杉的载量增加了0.01kg · m^ -2,麻栎林的载量增加了0.05kg · m^ -2,增加不明显;而旱冬瓜林的载量减少了0.12kg· m -2,减少了5.77%。  相似文献   

18.
19.
Mori A  Takeda H 《Tree physiology》2004,24(6):661-670
Light-related plasticity of crown morphology and within-crown characteristics were investigated in understory sun and shade saplings of three codominant subalpine conifers, Abies mariesii M.T. Mast., Abies veitchii Lindl. and Picea jezoensis var. hondoensis (Mayr) Rehd. Compared with those of sun saplings, current-year shoots of shade saplings allocated less biomass to needles, resulting in less dense needle packing and hence less mutual needle shading. The proportion of lateral branch biomass in foliage was either similar in sun and shade saplings or greater in shade saplings, depending on the species, suggesting that, over the lifetime of a branch, greater needle longevity in shade compensates for reduced biomass investment in needles of current-year shoots of shade saplings. Saplings with slower-growing branches tended to have greater needle life spans, suggesting that plasticity of branch growth rate and plasticity of needle life span are interdependent. Both Abies species showed greater light-related plasticity of needle life span and branch growth than P. jezoensis. The greater shade tolerance of the Abies species derives from their broad flattened crowns with slow-growing branches. This type of crown development incurs substantial support costs, but the long needle life span of shade saplings of the Abies species compensates, at least in part, for their low annual investment in foliage, especially in the case of A. mariesii, which has a longer needle life span and slower-growing and stouter branches than A. veitchii. Compared with the Abies species, P. jezoensis had a less plastic crown morphology, and less variability of needle life span and branch growth in response to light, resulting in lower shade tolerance. However, compared with the flattened crown of Abies shade saplings, the conical crown of P. jezoensis saplings imposes a smaller support cost, making this species better adapted to rapid height growth than to survival in shade.  相似文献   

20.
Fuel treatment of wildland vegetation is the primary approach advocated for mitigating fire risk at the wildland-urban interface (WUI), but little systematic research has been conducted to understand what role fuel treatments play in controlling large fires, which factors influence this role, or how the role of fuel treatments may vary over space and time. We assembled a spatial database of fuel breaks and fires from the last 30 years in four southern California national forests to better understand which factors are consistently important for fuel breaks in the control of large fires. We also explored which landscape features influence where fires and fuel breaks are most likely to intersect. The relative importance of significant factors explaining fuel break outcome and number of fire and fuel break intersections varied among the forests, which reflects high levels of regional landscape diversity. Nevertheless, several factors were consistently important across all the forests. In general, fuel breaks played an important role in controlling large fires only when they facilitated fire management, primarily by providing access for firefighting activities. Fire weather and fuel break maintenance were also consistently important. Models and maps predicting where fuel breaks and fires are most likely to intersect performed well in the regions where the models were developed, but these models did not extend well to other regions, reflecting how the environmental controls of fire regimes vary even within a single ecoregion. Nevertheless, similar mapping methods could be adopted in different landscapes to help with strategic location of fuel breaks. Strategic location of fuel breaks should also account for access points near communities, where fire protection is most important.  相似文献   

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