特大冰雪灾害干扰下大明山常绿阔叶林木质残体的贮量特征

以2008年初遭受特大冰雪灾害干扰的大明山国家级自然保护区常绿阔叶林为研究对象,采用样方收获法测定林地上的木质残体贮量及其腐解等级特征。结果表明:大明山常绿阔叶林木质残体的总贮量为37.249t·hm-2,其中粗木质残体和细木质残体的贮量分别为33.143和4.106t·hm-2,依次占林地木质残体总贮量的88.98%和11.02%;倒木和落枝是大明山常绿阔叶林木质残体贮量的主要贡献者,分别占木质残体总贮量的61.6%和34.6%;木质残体各腐解等级的贮量百分比序列为2级(69.22%)>4级(13.62%)>3级(12.98%)>5级(3.96%)>1级(0.22%);2008年初的特大冰雪灾害干扰对大明山常绿阔叶林木质残体输入量的有显著影响,导致腐解2级的贮量高达25.784t·hm-2。     

森林粗木质物残体（CWD）的研究进展

森林粗木质物残体（CWD）在保持森林生态系统的完整性方面发挥着重要的生态功能,影响着系统内外相关的生物和非生物过程,不仅为微生物、节肢动物、鸟类以及哺乳动物提供栖息环境,也是生态系统中重要的碳库和养分库,而且在减少土壤侵蚀和森林水文生态方面具重要的作用．文章总结了CWD的研究历史及状况,系统地阐述了CWD的形成和来源、贮量、分解、碳元素和营养元素的贮藏功能以及CWD对森林的更新作用、对生物多样性的维持和水文生态功能等．对今后CWD研究的重点及方向提出了建议．     

A quantitative framework for evaluating the mass balance of in-stream organic debris

A quantitative framework is developed for analyzing the mass budget of in-stream woody debris. Wood budgets are necessary for defining the relative importance of different recruitment processes over short and long periods, for designing spatially explicit simulation models, and for estimating the range of variability. The framework is used to analyze century-long patterns of large woody debris in streams that are governed by episodic forest death (fire and wind), forest growth and chronic mortality, bank erosion, mass wasting, decay, and stream transport. Simplified mathematical expressions are used to represent climatic, hydrologic, geomorphic, and biotic processes. Results are expressed in terms of time series and probability distributions. Predictions include that in areas of longer fire rotation (500 years) toppling of fire-killed trees comprises only 15% of the long-term wood budget yet chronic stand mortality that affect the large standing forest biomass ensures continuous large volumes of wood in streams. In contrast, toppling of fire-killed trees in forest environments with shorter fire rotations (150 years) comprise about 50% of the wood budget and indicates that field observers have a significantly higher chance of encountering low wood volumes in streams. Wood recruitment by bank erosion should increase irregularly downstream and bank erosion recruitment should exceed mortality recruitment at a bank erosion rate of approximately 5 cm per year. Recruitment from debris flows represents the single largest point source of woody debris to streams. The rarity of debris flows, in conjunction with a 3% per year annual decay rate, limits the contribution of wood from debris flows to about 12% of the long-term wood budget. Fluvial transport of wood promotes an increase in both inter-jam spacing and jam volume downstream. The proportion of woody debris transported into a reach in comparison to lateral recruitment approaches an asymptotic maximum of 50% when tree height approaches channel width. The relationships among process rates, their spatial variance across landscapes, and the resulting probability distributions of long-term patterns of wood abundance are proposed as a set of general theoretical principles. New data on wood supply and storage at the network scale are needed to fully test the predictions made in this analysis.     

森林生态系统粗木质残体生态功能研究

粗木质残体(CWD)是森林生态系统中重要的结构性与功能性单元。本文在对国内外CWD 研究历史进行简要回顾的基础上,对其生态功能及其分解过程进行了系统分析。指出了目前CWD 研究的热点及存在的问题,并建议加强对CWD生态服务功能的定量化研究,为其在维持生物多样性、生产力和其它生态过程方面的经营管理提供科学指导。参57。     

Loss of carbon sequestration potential after several decades of shifting cultivation in the Southern Yucatán

Cumulative losses from shifting cultivation in the tropics can affect the local to regional to global balance of carbon and nutrient cycles. We determined whether shifting cultivation in the Southern Yucatán causes feedbacks that limit future forest productivity and carbon sequestration potential. Specifically, we tested how the recovery of carbon stocks changes with each additional cultivation-fallow cycle. Live aboveground biomass, coarse woody debris, fine woody debris, forest floor litter and soil were sampled in 53 sites (39 secondary forests 2–25 years old, with one to four cultivation-fallow cycles, and 14 mature forests) along a precipitation gradient in Campeche and Quintana Roo, Mexico. From the first to the third or fourth cultivation-fallow cycle, mean carbon stocks in live aboveground biomass debris declined 64%. From the first to the third cycle, coarse woody debris declined by 85%. Despite declining inputs to soil with each cultivation-fallow cycle, soil carbon stocks did not further decline after the initial conversion from mature to secondary forest. The combined aboveground and soil carbon stock declined almost 36% after conversion from mature forest, however two additional cultivation cycles did not promote further significant decline, largely because of the stability of the soil carbon pool. Although age was the dominant factor in predicting total carbon stocks of secondary forests under shifting cultivation, the number of cultivation-fallow cycles should not be neglected. Understanding change beyond the first cycle of deforestation will enhance forest management at a local scale by improving predictions of secondary forest productivity and related agricultural productivity. A multi-cycle approach to deforestation is critical for regional and national evaluation of forest-based carbon sequestration. Finally, models of the global carbon cycle can be better constrained with more accurate quantification of carbon fluxes from land-use change.     

Biofuel harvests, coarse woody debris, and biodiversity - A meta-analysis

Forest harvest operations often produce large amounts of harvest residue which typically becomes fine (foliage, small limbs and trees) and coarse woody debris (snags and downed logs). If removed at harvest, residual biomass has potential to be a local energy source and to produce marketable biofuel feedstock. But, CWD in particular serves critical life-history functions (e.g., breeding, foraging, basking) for a variety of organisms. Unfortunately, little is known about how forest biodiversity would respond to large scale removal of harvest residues. We calculated 745 biodiversity effect sizes from 26 studies involving manipulations of CWD (i.e., removed or added downed woody debris and/or snags). Diversity and abundance of both cavity- and open-nesting birds were substantially and consistently lower in treatments with lower amounts of downed CWD and/or standing snags, as was biomass of invertebrates. However, cumulative effect sizes for other taxa were not as large, were based on fewer studies, and varied among manipulation types. Little is currently known about biodiversity response to harvest of fine woody debris. Predicting the effects of biomass harvests on forest biodiversity is uncertain at best until more is known about how operational harvests actually change fine and coarse woody debris levels over long time periods. Pilot biomass harvests report post-harvest changes in CWD levels much smaller than the experimental changes involved in the studies we analyzed. Thus, operational biomass harvests may not change CWD levels enough to appreciably influence forest biodiversity, especially when following biomass harvest guidelines that require leaving a portion of harvest residues. Multi-scale studies can help reduce this uncertainty by investigating how biodiversity responses scale from the small scale of manipulative experiments (i.e., 10-ha plots) to operational forest management and how biodiversity response to CWD levels might vary at different spatial and temporal scales and in different landscape contexts.     

Effect of improvement cutting on the community structure of aphyllophoraceous fungi on Okinawa Island

Improvement cutting, which decreases tree density by cutting and thinning while leaving logging residues, has been practiced on Okinawa Island since the 1970s and changes not only the vegetation composition but also the amount and composition of woody debris on the forest floor. To quantitatively compare the effect of improvement cutting with the effects of woody debris and vegetation on the community structure of aphyllophoraceous fungi, we conducted a field survey of vegetation composition, the amount and composition of woody debris, and the aphyllophoraceous fungi flora on Okinawa Island in December 2006. We established 3 plots in forest stands without logging and 13 plots in a forest that had undergone improvement cutting from 1 to 22 years before our field survey. The number and volume of woody debris tended to be highest soon after improvement cutting. We recorded 1,172 fungal samples belonging to 81 species in four orders. Fungal species richness was correlated with the composition of the woody debris. Fungal species composition in forest plots 1 year after improvement cutting differed from most of the other logged forests. Variation partitioning revealed that the effects of improvement cutting explained 14.1 % of the total variation in fungal species composition, and that differences in woody debris and vegetation not related to improvement cutting explained 40.4 %. Changes in the composition of woody debris caused by improvement cutting explained at most 2.7 % of the variation. This suggests that improvement cutting does not have a strong effect on fungal community structure, but woody debris does.     

Effects of management on coarse woody debris volume and composition in boreal forests in northern Sweden

Forest management practices have led to a reduction in the volume and a change in the composition of coarse woody debris (CWD) in many forest types. This study compared CWD volume and composition in reserves and two types of managed forest in the central boreal zone of Sweden. Ten areas were surveyed, each containing clear-cut, mature managed and old-growth stands, to determine the volume of standing and lying CWD in terms of species composition, decay class and size class. Volumes of CWD on clear-cuts and in mature managed forests were high compared with previous studies. Old-growth forests (72.6 m³ ha^?1) contained a greater volume of CWD than mature managed forests (23.3 m³ ha^?1) and clear-cuts (13.6 m³ ha^?1). Differences were greatest for the larger size classes and intermediate decay stages. Despite stand ages being up to 144 years, CWD volume and composition in managed forests was more similar to clear-cuts than to old-growth forests.     

森林粗木质残体研究综述

森林粗死木残体（coarse woody debris,CWD）是指所有地上和地下小头直径≥2．5cm的枯立木（snag）、倒木（Log）、树桩（Stump）、大枯枝（Large Branch）、小枯枝（Twig）、连接在活树上的枯死小树枝（Suspended Twig）等的总称。该文综述了近几十年CWD的研究成果,总结了CWD的概念及其在碳氮循环、生物多样性维护等方面的生态功能,CWD的贮量和养分动态。大量的研究证明,CWD对森林生态系统具有重要的生态功能,但它的蓄积量、空间分布以及它的生态意义因所研究的森林生态系统植被林型而异。另外,CWD还受诸多生物学和非生物学因子的影响,研究各因子与CWD的关系对于保持CWD的生态功能也是十分必要的。     

Decomposition of Scots pine fine woody debris in boreal conditions: Implications for estimating carbon pools and fluxes

Litter quality and environmental effects on Scots pine (Pinus sylvestris L.) fine woody debris (FWD) decomposition were examined in three forestry-drained peatlands representing different site types along a climatic gradient from the north boreal (Northern Finland) to south (Southern Finland) and hemiboreal (Central Estonia) conditions. Decomposition (percent mass loss) of FWD with diameter ≤10 mm (twigs) and FWD with diameter >10 mm (branches) was measured using the litter bag method over 1–4-year periods. Overall, decomposition rates increased from north to south, the rate constants (k values) varying from 0.128 to 0.188 year⁻¹ and from 0.066 to 0.127 year⁻¹ for twigs and branches, respectively. On average, twigs had lost 34%, 19% and 19%, and branches 25%, 17% and 11% of their initial mass after 2 years of decomposition at the hemiboreal, south boreal and north boreal sites, respectively. After 4 years at the south boreal site the values were 48% for twigs and 42% for branches. Based on earlier studies, we suggest that the decomposition rates that we determined may be used for estimating Scots pine FWD decomposition in the boreal zone, also in upland forests. Explanatory models accounted for 50.4% and 71.2% of the total variation in FWD decomposition rates when the first two and all years were considered, respectively. The variables most related to FWD decomposition included the initial ash, water extractives and Klason lignin content of litter, and cumulative site precipitation minus potential evapotranspiration. Simulations of inputs and decomposition of Scots pine FWD and needle litter in south boreal conditions over a 60-year period showed that 72 g m⁻² of organic matter from FWD vs. 365 g m⁻² from needles accumulated in the forest floor. The annual inputs varied from 5.7 to 15.6 g m⁻² and from 92 to 152 g m⁻² for FWD and needles, respectively. Each thinning caused an increase in FWD inputs, up to 510 g m⁻², while the needle inputs did not change dramatically. Because the annual FWD inputs were lowered following the thinnings, the overall effect of thinnings on C accumulation from FWD was slightly negative. The contribution of FWD to soil C accumulation, relative to needle litter, seems to be rather minor in boreal Scots pine forests.     

Family forest owner preferences for biomass harvesting in Massachusetts

U.S. forests, including family-owned forests, are a potential source of biomass for renewable energy. Family forest owners constitute a significant portion of the overall forestland in the U.S., yet little is known about family forest owners' preferences for supplying wood-based biomass. The goal of this study is to understand how Massachusetts family forest owners feel about harvesting residual woody biomass from their property. The study estimates the probability that Massachusetts landowners will harvest biomass as part of a timber harvest using data from a survey of 932 Massachusetts family forest owners. Logistic regression results suggest that the likelihood of harvesting for biomass is quite low, and that the supply of participation in biomass harvesting is inelastic with respect to price. These low probabilities may be due to the method used to account for preference uncertainty, as well as the unique nature of Massachusetts forests, forest markets, and landowner attitudes in comparison to other states (e.g., Minnesota). The study suggests that it would be more effective to target renewable energy policy toward different regions and/or markets rather than develop a uniform national policy.     

昆明西山国家森林公园主要林型地表可燃物的特征

通过外业调查和实验室测定,利用绝对含水率和载量2个关键指标,研究了持续干旱背景下昆明西山国家森林公园的华山松、云南油杉、滇青冈、地盘松、麻栎和旱冬瓜等6种主要林型的可燃物特征。结果表明:地表可燃物、粗死木质残体的含水率最高的都为云南油杉林,分别为55.36%,40.95%,最低都为滇青冈林,分别为10.26%,10.58%,差异显著,最高与最低分别相差约5.4倍、3.9倍;载量都达到较高水平,地表可燃物载量最大为华山松林(4.53±1.11)kg/m2,最低为旱冬瓜林(2.08±0.19)kg/m2,相差约2.2倍,粗死木质残体载量最高为地盘松林为(0.94±0.16)kg/m2,最低为旱冬瓜为(0.07±0.03)kg/m2,相差约13.4倍。     

Effects of preparation of clear‐cut forest sites on the soil mycobiota with consequences for Scots pine growth and health

Fungal abundance and diversity were assessed in two differently managed forest soils. In soil 1, chipped woody debris had been spread on the top soil and mixed with the soil by standard deep ploughing to 50 cm. In soil 2, woody debris was removed and there was no ground preparation. Ten years later, the different methods of management resulted in unexpectedly distinctive habitat heterogeneity with a high variation in structure of the mycobiota communities. Incorporation of woody debris into soil resulted in lower abundance of fungi but greater fungal diversity (which may enhance ecosystem stability) and was beneficial for the growth of Scots pines. Removal of post‐harvest woody debris from the surface of the clear‐cut site with no subsequent ground preparation, compared with incorporating chipped debris and deep ploughing, resulted in the soil of the subsequent 10‐year‐old Scots pine plantation having: (a) 159‐fold greater abundance of fungi, (b) lower fungal diversity, (c) different fungal community composition, (d) higher proportion of mycorrhizal fungi, (e) no root rot pathogens, (f) less tree biomass. The study showed that removal of post‐harvest woody debris and no ground preparation is potentially beneficial for the health of young coniferous trees.     

Estimating the effects of adjacency and green-up constraints on landowners of different sizes and spatial arrangements located in the southeastern U.S.

The maximum clearcut size and green-up period is important for land managers adhering to voluntary and regulatory guidelines. Therefore the impact of actual and hypothetical clearcut size restrictions is a concern for forest landowners who manage land and intend to practice forestry for profit. In this research, the effect of a 97.1 ha (240 ac) clearcut size constraint with a green-up period of 2-yrs is assessed for forest landowners with different forest land sizes, ownership patterns, and age class distributions. A meta heuristic which consists of threshold accepting, 1-opt tabu search, and 2-opt tabu search is used to develop spatially-constrained forest plans for 27 hypothetical forest landowners. These results are compared to a relaxed solution produced with linear programming, and statistical analyses are used to determine significant differences. The analysis provided enough evidence to suggest that two factors (size of ownership pattern and initial age class distribution), and one interaction factor (ownership size × initial age class distribution) are significant in explaining the differences in the percent reduction in forest plan value among the forests managed by the hypothetical forest landowners. From an absolute value reduction perspective, small-sized older forests were most affected. From a percent value point of view, one can conclude from this analysis that landowners with small-sized forests and young initial age class distributions will be significantly more affected by potential adjacency and green-up restrictions in the southeastern U.S. than other types of landowners.     

Ten-year responses of ponderosa pine growth,vigor, and recruitment to restoration treatments in the Bitterroot Mountains,Montana, USA

Little is known about ponderosa pine forest ecosystem responses to restoration practices in the Northern Rocky Mountains, USA. In this study, restoration treatments aimed at approximating historical forest structure and disturbances included modified single-tree selection cutting, with and without prescribed burning. We compared the effectiveness of restoration treatments on growth, vigor, and composition of recruitment responses with untreated controls. We used a randomized block design to detect treatment differences in mean individual tree basal area increment (BAInc10), growth efficiency (GE), and recruitment abundance between two restoration treatments (Cut-only and Cut-burn) and a Control. We further examined treatment effects by tree age-class (Young, Mature, Presettlement) using a spatial ANOVA model that incorporates the spatial autocorrelation among trees within experimental units. Ten years after implementing restoration treatments, mean individual tree BAInc10 and GE were significantly higher for treated units relative to Control units; all three age-classes benefited similarly from restoration treatments relative to the Control, with the greatest response in the Cut-only and moderate response in the Cut-burn. When treated units were compared, Cut-burn negatively affected BAInc10 and GE relative to Cut-only. Presettlement trees responded positively to treatment relative to the Control, particularly for BAInc10, demonstrating the potential of these old trees to respond to reduced competition. The Cut-burn treatment, in contrast, negatively affected the BAInc10 and GE response of postsettlement trees when compared to Cut-only. Restoration treatments did not reduce the amount of Douglas-fir recruits. In addition, the recruitment of both ponderosa pine and Douglas-fir species was associated with the proximate cover of woody debris in Cut-only and Control treatments. Finally, special consideration needs to be taken for spring Cut-burn treatments, which appeared to dampen growth and vigor, relative to Cut-only, particularly for Young and Mature trees, and increased recruitment of ponderosa pine and particularly Douglas-fir.     

Mapping dead wood distribution in a temperate hardwood forest using high resolution airborne imagery

Dead wood, in the form of coarse woody debris and standing dead wood, or snags, is an essential structural component of forest ecosystems. It plays a key role in nutrient cycling, ecosystem functions and provision of habitat for a wide variety of species. In order to manage dead wood in a temperate hardwood forest, an understanding of its availability and spatial distribution is important. This research evaluates airborne digital camera remote sensing for mapping temperate forest dead wood across an area within Gatineau Park, Canada. Two approaches were evaluated: (1) direct detection and mapping of canopy dead wood (dead branches and tall snags) through the combination of three techniques in a hybrid classification: ISODATA clustering, object-based classification, and spectral unmixing, and (2) indirect modelling of coarse woody debris and snags using spectral and spatial predictor variables extracted from the imagery. Indirect modelling did not provide useful results while direct detection was successful with field validation showing 94% accuracy for detected canopy level dead wood objects (i.e. 94% of validation sites with canopy dead wood were detected correctly) and 90% accuracy for control sites (i.e. 90% of validation sites with no canopy level dead wood were identified correctly). The procedures presented in this paper are repeatable and could be used to monitor dead wood over time, potentially contributing to applications in forest carbon budget estimation, biodiversity management, and forest inventory.     

California's regulatory forest carbon market: Viability for northeast landowners

Carbon markets have the potential to reward landowners for improved forest management and forest conservation. To date, the Over the Counter (OTC) voluntary market represents the greatest opportunity for forest landowners to participate in carbon transactions. However, lack of a consistent carbon price signal and sporadic demand coupled by high transaction costs has prevented widespread participation from family forest landowners. Adoption of a U.S. based cap-and-trade program reduces price risk and may provide incentives for sustainable forest management across large areas. Yet few studies have examined the supply side of carbon offsets and factors affecting project financial viability. To address this gap, we assessed how (1) property characteristics (i.e. stocking level, forest type, size etc.); (2) silvicultural treatments; and (3) protocol and legislative requirements affect the financial viability of compliance forest offset projects, focusing on California's Air Resource Board (ARB) program due to its significance as the world's second largest carbon market. We used forest inventory data from 25 properties in the northeastern United States to examine the viability of the sites as ARB offset projects. We utilized the U.S. Forest Service Forest Vegetation Simulator for our growth and yield simulations. To examine the factors that influence project viability, we used a classification and regression tree analysis performed in S-Plus software. Results indicate C stocking and property size are the most important property characteristics driving return on investment. However, protocol requirements and legislative assumptions impacting long-term monitoring costs are also important factors. While reduced price risk in a compliance carbon market has the potential to improve forest management in North America; high initial project development costs, long-term monitoring obligations, and legislative uncertainty are significant barriers that will limit family forest landowner market participation. The model developed here can be used by U.S. landowners to assess the financial viability of their property as a compliance offset project and can be utilized by policymakers to develop cost-effective climate change policy.     

Riparian trees,shrubs, and forest regeneration in the coastal mountains of Oregon

Riparian trees and shrubs are important providers of shade, bank stability, and woody debris needed for optimal stream quality and fish habitat in the coastal mountains of Oregon, but more data are needed to relate this woody vegetation to environmental variables. Trees, shrubs, and forest regeneration were studied in 22 riparian environments to provide those data. Conifer basal area increased with elevation, stream gradient, time since disturbance, and distance from the stream; it decreased with stream width. Salmonberry (Rubus spectabilis Pursh) cover and stink currant (Ribes bracteosum Dougl. ex Hook.) cover were highest near the streams. Dwarf Oregon grape (Berberis nervosa Pursh) cover and salal (Gaultheria shallon Pursh) cover were lowest near the streams. Although forest regeneration was poor everywhere, it decreased with total shrub cover and increased with stream gradient. Existing riparian conifer stands should be maintained wherever a continuing supply of coarse woody debris is required.