The effect of herbage surface water and the provision of supplementary forage on the intake and feeding behaviour of cattle

High rainfall and the corresponding low herbage DM content has been shown to adversely affect cattle intakes and production. This could arise from physical restrictions on intake or digestion rate, or other behavioural limitations such as reduced palatability. A knowledge of the reason for the intake reduction would assist in determining the optimum supplement to feed. Eight steers were offered four diets in a changeover design. The diets were: A, cut herbage; B, cut herbage with added surface water; C, as B but with a low dry matter (DM) forage supplement (silage); and D, as B but with a high DM forage supplement (hay). The addition of surface water to herbage reduced DM intake and feeding time but had no effect on biting rate or rate of intake. Neither hay nor silage was eaten in sufficient quantities to restore DM intake, even though the silage was of similar nutrient composition to the herbage.     

The effect of drainage on herbage growth and soil development

The effects of drainage on the yield of perennial ryegrass ( Lolium perenne ) swards grown for silage, and on soil development under grassland, were studied between 1982 and 1985.
Over the 4 years of the experiment, drainage resulted in a mean increase in herbage production of 1-5 t dry matter (DM) ha⁻¹, 16% above the yield recorded on the undrained site. Drainage also led to increased herbage nitrogen contents at every cut except one, so that total nitrogen recovery was increased by an average of 24-8 kg N ha⁻¹ (15%) when drainage was installed. These total differences were mainly due to differences that were present at the first cut taken in June each year; drainage led to an average 24% increase in yield, and an average 27% increase in nitrogen recovery at the first cut.
Detailed sward measurements taken in 1985 endeavoured to explain these differences; drainage increased depth to free water over winter, enabling increased root efficiency and nitrogen availability, greater tillering and more rapid leaf extension in early spring. There was some suggestion that drainage also led to an improved topsoil structure in terms of improved aggregation and aeration. This is important for long-term soil development as well as short-term herbage yield.     

The effect of herbage allowance upon the herbage intake and performance of suckler cows and calves

Twenty-four Hereford × Friesian cows and their South Devon cross calves were allocated to three herbage allowances allotted daily for three periods of 8 weeks in a Latin square design. The daily allowances were 17, 34 and 51 g dry matter per kg cow plus calf live weight. Milk production was depressed by 0·2 and 1·2 kg d^-1 at the medium and low allowances. The corresponding falls in liveweight gain were for cows 0·26 and 0·25, and for calves 0·27 and 0·24 kg d^-1. Residual sward height after grazing gave a better indication of the animals' reaction to sward conditions and the management imposed than actual herbage allowance. The quantity per unit area and the composition of material present were important factors influencing intake. Calves were unable to compete with their dams to maintain herbage intake at the lower allowances and therefore are likely to benefit from additional feeding or creep grazing when residual sward height falls below 6cm for periods in excess of 1–2 weeks.     

The effect of surface height of swards continuously stocked with cattle on herbage production and clover content over four years

The long-term influence of sward height from April to July (Phase 1) and from July to early October (Phase 2) on total herbage and white clover production was measured over four years (1988–1991) as herbage accumulation. A subsidiary experiment to determine the influence of leaf area index (LAI) on gross canopy photosynthesis was conducted to aid interpretation of growth from herbage accumulation data. Target sward heights in 0·5 ha plots in two blocks were 5,7 or 9 cm in Phase 1 and 7 or 9 cm in Phase 2, although mean actual heights per phase were slightly higher. Net herbage accumulation (NHA) was measured within mobile exclosure areas over successive two-week intervals. Gross photosynthesis was measured in circular turves removed from the trial area representing a range of LAIs with an at least reasonable clover content. Despite wide differences in mean sward height and herbage mass, NHA and net clover accumulation for a given phase were not generally affected by treatments. Positive effects of grazing at 5 cm in Phase 1 on NHA and clover accumulation later in the year, and of grazing at 7 cm in Phase 2 on NHA in the following spring were sometimes apparent. Gross canopy photosynthesis (g CO₂ m^?2 h^?1) at 1500 μE m^?2 s^?1 and 18–21°C was linearly related to LAI described by 1·003 + 1·165 LAI over the LAI range 0·7 to 4·5. Total herbage and clover growth, interpreted from NHA by a previously described model, was predicted to be marginally lower in shorter swards. Similarity in NHA and clover accumulation between treatments was considered to be because of lower senescene and decomposition, and a higher proportion of new tissue being assigned to lamina growth, despite lower LAI and gross photosynthesis in the shorter swards. It was concluded that stocking intensity in swards continuously stocked with cattle did not have a strong influence on net total herbage and clover growth.     

The effect of some pre-treatments on proteolysis during the ensiling of herbage

Two experiments were carried out to examine the nitrogenous changes occurring in herbage during harvesting and ensiling. In the first, ryegrass-clover was wilted rapidly in the laboratory (6 h) and in the field under good (48 h) and poor (48 h and 144 h) weather conditions. Protein breakdown and ammonia formation were negligible in herbage wilted rapidly and it was only when the crop was exposed to a prolonged wilt in humid conditions that appreciable proteolysis occurred. In the second experiment ryegrass-clover was ensiled in laboratory silos after treatment with varying levels of formic and sulphuric acids. An additional treatment included herbage inoculated with a mixture of lactic acid bacteria and glucose. The silos were opened after 4 and 50 d and samples were analysed for protein-N, ammonia-N, water-soluble carbohydrates and organic acids. There was a high negative correlation between level of addition of either acid and the degree of proteolysis and deamination. Even at the highest levels of formic acid (7·7 g per kg) and sulphuric acid (4·0 g per kg) additions, however, about 45% of the original herbage protein was degraded after 50 d in the silo. The inoculum treatment was also effective in reducing proteolysis, the effect after 50 d being comparable with formic acid applied at a rate of 4·1 g per kg.     

The effect of herbage allowance on daily intake by Creole heifers tethered on natural Dichanthium spp. pasture

Two experiments were carried out in Guadeloupe to estimate the organic matter intake (OMI) and digestibility (OMD) of a Dichanthium spp. sward, grazed by tethered Creole heifers [mean live weight (LW) 202 ± 2·0 kg], at three daily herbage allowances. Experiment 1 examined herbage allowances of 16, 25 and 31 kg of dry matter (DM) d^–1 on a fertilized sward at 21 days of regrowth whereas, in experiment 2, lower allowances of 11, 15 and 19 kg DM d^–1 were examined on the same sward, which was unfertilized and grazed at 14 days of regrowth. In each experiment, the herbage was grazed with three groups of two heifers in a 3 × 3 Latin square design. Sward characteristics were described before grazing. OMI was calculated from total faecal output, and OMD was predicted from the crude protein (CP) content of the faeces. The amount of herbage defoliated by the heifers was also estimated on tillers selected at random.
Organic matter intakes were on average 26 g and 19 g OM kg^–1 LW, and OMD values were 0·740 and 0·665 for Experiments 1 and 2, respectively, and were not affected by allowance. In Experiment 1, the herbage quality was high [0·50 of leaf and 116 g CP kg^–1 organic matter (OM)] for a tropical forage, whereas in Experiment 2, the quality of the herbage (0·27 of leaf and 73 g CP kg^–1 OM) was lower. These differences were reflected in differences in intake and digestibility in the two experiments.
The experimental tropical Dichanthium spp. swards can have intake characteristics similar to those of a temperate sward.     

The effect of herbage allowance on intake and performance of ewes suckling twin lambs

The effect of herbage allowance on the herbage intake and performance of ewes and their twin lambs at pasture was investigated. Daily herbage allowances of 40,80.120 and 160 g organic matter (OM) kg^-1 ewe live weight, based on herbage mass measured to ground level, were offered during the first 12 weeks of lactation.
The sheep were grazed rotationally around four paddocks of a perennial ryegrass pasture for 7-day periods and herbage mass, extended tiller length and digestibility of the herbage consumed by the animals were estimated. Herbage intake by the ewes was estimated during weeks two to twelve and live weights were recorded weekly: during the last two weeks of the experiment grazing behaviour of one ewe on each treatment was recorded continuously.
Herbage intakes by the ewes (164, 1–81, 2–42 and 268 ± 0153 kg d^-1) and live weight gains of the lambs (202, 245, 274 and 300 ± 7–3 g d^-1) increased with increasing herbage allowances. Herbage intakes by the ewes and growth rates of their lambs increased up to a herbage allowance which was over five times the amount of herbage eaten by the ewes.
As the animals reduced herbage mass and sward height, biting rates by the ewes during grazing increased by 4 (± 0·08) bites min^-1 cm^-1 and masticating rates decreased. Mastications reached a maximum of 90 (± 3·5) min^-1 at a sward surface height of 9 cm.     

The effect of cutting systems on the yield of herbage from a small-plot experiment

Plots of an established perennial ryegrass sward (cv. Talbot) fertilized with 100 or 300 kg N ha⁻¹ were harvested with one of three plot mowers; an Agria with 38-mm finger spacing and an Allen Mayfield with 38-mm finger spacing or 78-mm finger spacing. The choice of mower was arranged factorially with four post-harvest treatments: no post treatment; a second (cleaning) cut using the same mower and cutting in the same direction as the first (sampling) cut; a cleaning cut using the same mower, but in the opposite direction to the sampling cut; and a cleaning cut using a rotary mower. Sward productivity as organic matter (OM) was measured over four harvests at 6- to 7-week intervals. The Agria mower resulted in 12·3 and 15-9% more OM yield than the Allen Mayfield fitted either with knife-bars at a similar spacing, or with wider-spaced fingers, respectively. Yield of OM in the subsequent cut was reduced when a cleaning cut was taken; particularly when it was taken in the opposite direction to the sampling cut or when using a rotary mower.     

The effect of daily herbage allowance and sward characteristics upon the ingestive behaviour and herbage intake of calves under strip-grazing management

Measurements of herbage intake and ingestive behaviour were made on British Friesian calves 4–9 months old, which were strip-grazed on swards of S24 perennial ryegrass at three (Experiment A) or four (Experiment B) levels of daily herbage allowance in three (A) or four (B) 12-d periods. Herbage intake declined by approx. 18% as daily herbage allowance was reduced from 90 to 30 g DM per kg LW in both experiments, but the relationship between intake and allowance was not consistent between periods. Bite size and rate of biting were greater in calves on low than on high allowances after entry to a new strip of herbage but the positions were reversed towards the end of grazing. Reduced herbage intakes at low allowance resulted from reductions of approx. equal magnitude in bite size, rate of biting and grazing time, though not all of these effects were significant. It is suggested that these responses result from the increasing difficulty of prehending and ingesting herbage as swards are grazed down. The calves at low herbage allowances did not increase grazing time, probably because their behaviour was conditioned by anticipation of a new allowance of herbage. Bite size and daily herbage intake were greater on a spring sward than on an autumn sward, but within experiments herbage intake was not consistently related to any of the sward characteristics measured.     

The effect of tractor wheel passes on herbage production from diploid and tetraploid ryegrass swards

Diploid and tetraploid perennial ryegrass swards were managed under an intensive silage system over 3 years. The swards were subjected to five tractor wheel pass treatments to simulate combinations of silage operations in the field from cutting, tedding, lifting and fertilizing. Wheel tracking was applied uniformly over the swards after June, July, August and October cuts each year.
Tractor wheel passes caused mean reductions in herbage dry matter (DM) yield during successive years of 14%, 6% and 9% compared with their absence, and frequent or delayed wheel passes affected yield more adversely than infrequent or undelayed wheel passes. Nitrogen and mineral concentrations in herbage and nutrient offtakes were also generally reduced but organic matter digestibility (OMD) was unaffected. Soil assessments each autumn showed that soil bulk density increased in the wheel-pass treatments, particularly when frequently applied. The two sward types responded in a similar manner to the wheel-tracking treatments, probably because they had similar cushioning capability, the disadvantage of the tetraploid in terms of fewer tillers in relation to the diploid being compensated for by larger tiller size. Soil compaction, with its adverse effect on sward growth vigour, and direct wheel damage to young regrowth will be lessened by minimizing wheel traffic on silage swards and undertaking operations over short rather than long periods.     

The influence of surface and sub-surface application methods for pig slurry on herbage yields and nitrogen recovery

Experiments were conducted on a grassland site at Wrest Park, Silsoe, Bedfordshire between 1987 and 1989, to compare herbage yields from slurry applied by deep and shallow injection, low trajectory and conventional vacuum tanker methods. Slurry application rates for all spreaders were calibrated at 86 ± 5 t ha^-1, an equivalent of c. 200 kg NH₄⁺-N ha^-1 applied in autumn or in spring.
As expected, herbage yields following spring applications were higher than from autumn applications, with average mineral fertilizer equivalents of 122 and 89 kg N ha^-1 respectively. Yields from the conventional and low trajectory spreaders showed no consistent differences. However, in both years, first cut yields from plots were significantly lower ( P < 0·05) where slurry had been injected than where surface applications had been used by an average of 0·7 t DM ha^-1. Subsequent cuts in 1988 demonstrated higher residual effects from injection so that annual total yields were similar from all slurry applications irrespective of spreader type.
Analysis of N content revealed high N levels in herbage from deep injection plots. Mean concentrations of N in the herbage, expressed as a percentage of the dry matter, were 1·43 for surface treatments and 1·79 for deep injection in 1988, and 1·84 for surface treatments, 2·13 for shallow injection and 2·68 for deep injection in 1989.     

The herbage productivity of tall fescue in the Pampas region of Argentina is correlated to its ecological niche

Since its introduction in the 1950s, tall fescue has spread throughout the Pampas region of Argentina to become naturalized in much of this region. Its annual forage mass productivity shows significant differences among sites. In this work, we hypothesized that, under dryland conditions, the annual forage mass productivity is related to the probability that the corresponding site is within the ecological niche of the species. To test this hypothesis, the spontaneous occurrence of tall fescue was surveyed in an area of 440 000 km². The abundance of tall fescue was verified on roadsides of national and provincial roads, in systematic sampling points spaced every 50 km. Subsequently, the ecological niche of the species was estimated with the Maxent software, based on the environmental information available. Additionally, the bioclimatic variables with greatest influence in determining the ecological niche were identified. Based on the information of comparative yield trials of tall fescue cultivars, a correlation analysis between annual forage mass productivity and the likelihood of ecological niche was performed. We found that the realized ecological niche of tall fescue in the Pampas region covers an area of 237 000 km² where the average minimum temperature of the coldest month is between 2·6 and 3·2°C and the rainfall in the driest quarter is 100 mm or higher. We also found that there were significant correlations between annual forage mass productivity and the probability of realized ecological niche. Therefore, in areas with low probability of realized ecological niche, other forage species could be a better option than tall fescue to obtain a high and stable annual forage mass productivity.     

The effects of change in herbage mass on rates of herbage growth and senescence in mixed swards

A study was conducted to determine the influence of a change in herbage mass on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in swards of Lolium perenne L., Poa annua L. and Trifolium repens L. under continuous stocking with sheep. Swards were maintained at either high (1700–1900 kg OM ha⁻¹; H) or low (700–900 kg OM ha⁻¹; L) herbage mass from April to 20 August. From 20 August to 13 September half of the L sward was permitted to grow until a herbage mass similar to that of the H sward was achieved (LH), and at the same time half of the H sward was grazed down until a herbage mass similar to that of the L sward was achieved (HL). The other half of each sward was held at constant mass (LL and HH). Estimates of G, S and NP were obtained for each plant species over a 2-week period from 17 September to 1 October and estimates of species population densities were also made.
The combined species rates of G and S per unit area were highest in treatment LH and lowest in treatment HL, whilst the rates for the LL and HH swards were intermediate. The rate of NP was similar in the LL, HH and LH treatments (34.5, 29.3, 33.6 ± 6.2 kg DM ha⁻¹d⁻¹) but was lower in the HL sward (100 ± 6.2). The effects of alteration of herbage mass on individual tiller or stolon G and S rates were rapid but population density changes were slower.
Within the limited conditions of this trial it was not possible to increase NP by manipulation of herbage mass under continuous stocking management but the results indicate that NP can be reduced in the short term if a sward of high herbage mass and low population density is grazed hard.     

The effect of sward height and its direction of change on the herbage intake, diet selection and performance of weaned lambs grazing ryegrass swards

Abstract Four sward height treatments were imposed by continuous variable stocking using at least ten Suffolk × Greyface lambs per plot from late July to late August: constant 3·5 cm. constant 6·0 cm. 3·5cm increasing to 6·0 cm and 6·0 cm decreasing to 3·5 cm. The treatments were established on two swards given fertilizer N applications over the season of 97 and 160 kg N ha^?1 respectively. Animal density was greater on the high fertilizer treatment, at the lower sward height and especially on the decreasing height treatments. Liveweight change of lambs was higher (P <0·001) on the 6-cm than on the 3·5-cm treatments (+159 vs-13g d^?1 and was also higher (P <0·001) on the increasing than on the decreasing sward height treatments (+92 vs-26 g d^?1). Herbage organic matter intake (OMI), measured on two occasions in the experiment, was greater (P <0·001) on the 6·0 cm than on the 3·5-cm sward heights whereas values for the increasing sward height treatments were much greater than those for the decreasing sward height treatments. There was little difference in the organic matter digestibility of the diet between treatments. Diets were composed largely of lamina, although there was more pseudostem and dead herbage in the diets of Iambs grazing the decreasing than the increasing sward height treatments at the end of the experiment. Bite mass was closely related to OMI but the treatment and period differences were relatively greater than for OMI. Bite mass was more closely related to the depth of the lamina layer (sward height-pseudostem height) than it was to sward height. There was evidence that pseudostem acted as a barrier to defoliation on these short swards and also that the proportion of youngest leaf in the diet was positively related to sward height and to increases in sward height. Sward height and especially the direction of change in sward height, together with associated stock density, were potent influences on lamb growth rate. This was a consequence of differences in herbage intake, which was strongly influenced by bite mass.     

Effect of humic substances on nutrient uptake by herbage and on production and nutritive value of herbage from sown grass pastures

The effect of humic substances on the nutrient uptake, herbage production and nutritive value of herbage from sown grass pastures was studied in six field experiments. Commercial humic substances were applied in combination with mineral fertilizer or slurry, either as a solution (HF liquid; 8·3 kg humic substances ha⁻¹) or incorporated into the mineral fertilizer (HF incorporated; 3·6 to 6·4 kg humic substances ha⁻¹). A series of cuts, ranging from two to five cuts, was taken during the growing season. The general response in herbage production to application of humic substances was an increase in herbage mass of dry matter (DM) at the first cut although this was only significant in two experiments for the HF incorporated treatment. Total herbage production of DM over the growing season, however, was similar for treatments with or without application of humic substances. The overall effect of HF incorporated and HF liquid on the herbage mass of DM at the first cut across the experiments was calculated using a meta-analysis technique and it was shown that there was a significant proportional increase of 0·14 ( P  <   0·05) with the HF incorporated treatment and a non-significant increase of 0·08 with the HF liquid treatment compared to the control treatment. The nutritive value of the herbage at the first cut was similar across all treatments. In general nitrogen, phosphorus and potassium uptake at the first grass cut was higher after application of humic substances but only in one experiment was this increase statistically significant.     

Interactions of slope and canopy of herbage of three herbage species on transport of faecal indicator bacteria by rain splash

The movement of faecal pathogens from land to surface and ground‐water are of great interest because of the public and livestock health implications. Knowledge of canopy structure and how it might be managed to help mitigate nutrient and pathogen movement in pasture is needed to create management practices that balance livestock production with environmental benefits. An experiment was conducted using a rainfall‐simulating device to test whether canopy structure of species common to pastures in Appalachia, USA could be managed to influence dispersion of faecal pathogens. Seven pots (30‐cm diameter) of white clover, orchardgrass and perennial ryegrass were lined up on horizontal and sloping surfaces under a rainfall simulator. The centre pot was inoculated at the soil surface with 4 × 10¹⁰ faecal coliform bacteria (FC) just before rainfall simulation started. The species were maintained under short, moderate and tall canopy management treatments. White clover exhibited the greatest rates of lateral and vertical dispersion of FC into the canopy, especially in the short canopy management treatment following 30 min of rainfall (about 40 mm). Low concentrations of faecal coliform bacteria also dispersed into the canopies of the grass species but the differences in concentration of FC between the grass species were not different. When the proportion of white clover in a pasture is high, the canopy should be relatively taller to reduce the likelihood of infection associated with faecal coliform‐contaminated herbage.     

The effect of inoculation with Lactobacillus plantarum on fermentation in laboratory silos of herbage low in water-soluble carbohydrate

In experiment 1, a mixture of perennial ryegrass and white clover (176 g DM kg^?1) was ensiled with no addition, or with the application of Lactobacillus plantarum, 3 × 10⁶(g fresh weight of herbage)^?1 or enterobacteria, 3 × 10⁶ (g fresh weight of herbage)^?1 or both of these inoculants. Silos were incubated at either 18°C or 26°C for 7 d and sealing of half of the silos was delayed for 48 h. In experiment 2, cocksfoot (247 g DM kg^?1) was ensiled with no additive, with the application of L. plantarum, 4 × 10⁶ (g fresh weight of herbage)^?1 enterobacteria, 4 × 10⁶ (g fresh weight of herbage)^?1 or ammonium nitrate, 5 g (kg herbage)^?1. Silos were incubated at 18°C and sealing was delayed for 48 h. In neither experiment were the untreated silages of satisfactory quality after 35 d ensilage, having pH's of 4·3 and 6·4. and ammonia-N concentrations of 139 and 209 g N (kg total N)^?1 in experiments 1 and 2 respectively. The poorer fermentation in experiment 2 reflected the lower water-soluble carbohydrate (WSC) content of the ensiled herbage (136 and 53 g WSC (kg DM)^?1 in experiments 1 and 2 respectively). Inoculation with L. plantarum significantly reduced silage pH and the concentrations of ammonia-N (experiment 1 only) and ethanol and increased lactic acid concentrations. Inoculation with enterobacteria had only minor effects in each experiment. In experiment 1, incubation at 26°C increased the rate of fermentation in the silos as pH was significantly lower after 7d incubation at 26°C than at 18°C, but this effect had largely disappeared after 35 d incubation. When sealing of the silos was delayed for 48 h, the resulting silages had significantly higher pH and ammonia-N contents than silages from silos that had been sealed immediately after filling. In experiment 2, addition of ammonium nitrate significantly reduced the concentrations of ethanol and acetic acid in silages opened after 7 d. After 35 d the concentrations of ethanol were significantly reduced and those of ammonia-N increased by the addition of ammonium nitrate. There were marked increases in silage pH between 7 and 35 d ensilage and in the concentrations of ammonia-N and acetic acid in the silages that had been inoculated with ammonium nitrate at ensilage.     

The effect of cell wall degrading enzymes or formic acid on fermentation quality and on digestion of grass silage by cattle

A first cut of timothy, treated with water (untreated), formic acid (FA), cellulase + lactic acid bacteria (CB), cellulase + hemicellulase (CH) or cellulase + hemicellulase + a lignin-modifying enzyme (CHL), was ensiled in pilot-scale silos. Silages, except CB, were fed to four male cattle, each equipped with a rumen and duodenal cannula, in a digestibility trial designed as a 4 × 4 Latin square. The animals were fed a diet of 400 g of concentrate and 600 g of silage at a level of 70 g DM kg^?1 live weight (LW^0·75). All enzyme-treated silages were well-preserved with a more extensive fermentation than in FA silage. The quality of untreated silage was poorer as indicated by higher pH and ammonia-N content. The amount of effluent from enzyme-treated silages ranged from 116 to 127 g kg^?1; for FA and untreated silages values were 101 g kg^?1 and 80 g kg^?1, respectively. Total DM losses from enzyme-treated silages were higher than from FA silage (P < 0·05). No significant differences were noticed between silages in the apparent digestibility of organic matter (OM), neutral-detergent fibre (NDF), acid-detergent fibre (ADF) or nitrogen (N). The apparent digestibility of cellulose was higher with enzyme-treated silages than with FA silage (P < 0·05). The values for microbial N flow at the duodenum were 80·0, 91·9, 80·7 and 70·5g N d^?1, and for the efficiency of rumen microbial N synthesis 38·6, 47·6, 36·9 and 32·5 g N kg^?1 OM apparently digested in the rumen for untreated, FA, CH and CHL silages, respectively. In the rumen the molar proportion of propionate was higher (P < 0·01) and that of butyrate lower (P < 0·01) with enzyme-treated silages when compared with FA silage. The proportion of butyrate was also lower with untreated than with other silages (P < 0·01). The rumen residence time of NDF and ADF was longer (P < 0·05) with enzyme-treated silages than with FA silage.     

Effect of herbage mass per unit area and conditioning treatment on herbage drying rate

The rates of drying of herbage, cut from perennial ryegrass (Lolium perenne L.) – dominant swards and subjected to different treatments, were investigated under field conditions by changes in weight of herbage in wire mesh trays in 1995 and 1996. A series of replicated factorial experiments studied the effects, in different combinations, of intensity of conditioning achieved by passing the cut herbage through a laboratory‐scale macerator zero (0C), once (1C), three (3C) or six (6C) times; weight of herbage per unit area equivalent to 450, 675 and 900 g dry matter (DM) m^?2. In one experiment, pressing the herbage to form a mat was incorporated into the experimental design. A further experiment investigated the effect of varying the proportion of conditioned herbage in the herbage mass from 0·00, 0·25, 0·50, 0·75 and 1·00 on drying rate. On each occasion the trays plus herbage were weighed at hourly intervals over an ≈6‐h period and the DM content of the herbage estimated from the change in weight. On all occasions, conditioning and weight of herbage per unit area significantly (P < 0·001) influenced herbage drying rate. Lowering the weight per unit area of both unconditioned and conditioned herbage increased the rate of moisture loss. Unconditioned herbage at the equivalent of a herbage mass of 450 g DM m^?2 had a total moisture loss that was on average 1·5–1·8 times greater than unconditioned herbage at the equivalent of a herbage mass of 900 g DM m^?2. Similarly, conditioned herbage at the equivalent of a herbage mass of 450 g DM m^?2 had a total moisture loss that was 1·8–2·3 times greater than unconditioned herbage at the equivalent of a herbage mass of 900 g DM m^?2. Increasing the level of conditioning produced a non‐linear response in rate of moisture loss, consequently 3 passes through the macerator produced >0·95 of the total moisture loss that was produced by 6 passes through the macerator. Increasing the proportion of conditioned herbage in the herbage mass increased rate of moisture loss and consequently final DM content (P < 0·001) although there was little effect from increasing the proportion of conditioned herbage above 0·75. The effects of conditioning and weight of herbage per unit area treatments on total nitrogen , water‐soluble carbohydrate and acid‐detergent fibre concentration of the herbage were small.     

The size of the root system of tetraploid red clover and its relation to the chemical composition of the herbage produced

The size of the root system was measured by means of electrical capacitance in relation to the soil in field experiments with three varieties of tetraploid red clover over a period of 3 years. The root system was greatest in the first year, smaller in the second year and least in the third year. The plants that gave an above-average yield in any cut always had above average-sized root systems in the preceding period, but usually after cutting, the size of their root systems decreased, an effect which was accentuated by plants drying out. The plants that dried out during the experimental period always had below average-sized root systems in preceding periods. It was found in four cuts that the herbage dry matter of the plants with larger roots contained a smaller proportion of nitrogenous materials and ash, and more fibre, but the yield of all measured nutrients and elements increased. The ash of the herbage DM of these plants contained more potassium, calcium and magnesium. The quantity of mineral materials absorbed per unit of size of root systems was higher in the plants with larger root systems.