A tree/crop interface design and its use for evaluating the potential of hedgerow intercropping

The paper describes a tree/crop interface (TCI) experiment designed to investigate the effects of row orientation using Leucaena leucocephala Lam. Each TCI plot consisted of a regularly pruned Leucaena hedge in the middle and 12 crop rows on either side. Eight such plots were arranged at 45° around a sole Leucaena plot with rows oriented in four compass directions viz., North-South, East-West, Northeast-Southwest and Northwest-Southeast.Results of four years from 1984 to 1987 did not show any effect of row orientation, and similarly, no effect was seen on crop rows due to their location on the windward or leeward side of the hedge. The TCI effect was positive on the first crop row in the first year because Leucaena grew slowly, but depressed the yield of the first 4 to 6 crop rows(1.8 to 2.7 m from hedge) in subsequent years. The negative effect of Leucaena was noted more on sunflower in a relatively dry year than on sorghum in other years.Results from the TCI plots were used to estimate the yield of five hedgerow intercropping (HI) systems with varying alley widths (2.7 to 9.9 m). Comparison with sole stands of Leucaena and crops indicated that HI was more productive particularly at close alley widths. For example, hedges spaced at 2.7 m and 3.6 m averaged 37% and 25% higher productivity than the respective sole stands; but this advantage may be an overestimation of the real potential.The relevance of TCI experiments for studying agroforestry systems, their merits and limitations, especially of the design employed in this study are discussed.     

Tree-crop interactions in hedgerow intercropping with different tree species and tea in Sri Lanka: 1. Production and resource competition

Incorporation of tree hedges along contours has been proposed as a means of reducing soil erosion and increasing soil fertility of tea (Camellia sinensis (L.) O. Kuntz) plantations on sloping terrain in high-rainfall zones of Sri Lanka. Tea yields in these hedgerow intercrops are determined by the balance between the positive (i.e., increased soil fertility) and negative (i.e., resource competition) effects of hedgerows. Therefore, the objectives of the present study were to determine: (a) the net effect of six different hedgerow species (Calliandra calothyrsus Meissner, Senna [Cassia] spectabilis (DC.) H. Irwin and Barneby, Euphatorium innulifolium (R.M. King and H. Rob.) H.B.K., Flemingia congesta Aiton F., Gliricidia sepium (Jacq.) (Kunth.) Walp. and Tithonia diversifolia (Hemsl.) A. Gray) on long-term tea yields; (b) the effect of incorporating hedgerow prunings as mulch; and (c) the relationship between hedgerow characteristics (i.e., biomass production, canopy and root growth) and tea yield variation. Tea yields, measured over one complete pruning cycle from October 1998 to September 2001, showed reductions relative to a sole tea crop under all hedgerow species except Euphatorium. The yield reductions ranged from 22 to 40%. Tea yields under Euphatorium showed increases up to 23% relative to the sole crop control. Addition of hedgerow prunings as mulch increased tea yields in all hedgerow intercrops. The yield increases ranged from 11 to 20%, with the highest being under Euphatorium. Tea yields showed a negative relationship (R²=0.38) with the pruned biomass of hedgerows. Limitation of environmental resources (e.g., water and light) and hedgerow characters which intensified resource competition (i.e., greater canopy lateral spread and height and greater root length densities, especially in the top soil layer) were responsible for observed tea yield reductions in hedgerow intercrops.     

A tree/crop interface orientation experiment with aGrevillea robusta hedgerow and maize

Pruned triple-row hedges ofGrevillea robusta with 11 rows of maize planted in parallel on either side were grown in 12.5×16.0 m plots on a flat, freely exposed site. There were two replicates of four orientation treatments and crop yield was recorded row-by-row over 9 seasons. After the hedges had become established there were some marked differences in maize yields between sides, especially in the first rainy seasons. The maize on the downside of prevailing winds frequently showed improved growth and yield over the non-sheltered maize, sometimes by as much as 50% over the whole subplot and up to 80% in the tree/crop interface zone. These differences were significant (p0.05) when analyzing the whole experimental period for first and for second rainy seasons separately (prevailing winds more southerly or northerly, respectively), although they varied between seasons and zones. Attributing results to shelter can only be adduced circumstantially because meteorological data were not collected at the plot level. Shading effect were not considered important over the whole sub-plot. Soil water profiles obtained from an array of gypsum resistance blocks in two of the plots (at 000° and 090°) characterized the wetting and drying patterns in this system, but expected asymmetries due to orientation effects were not apparent except, perhaps, fleetingly in the topsoil. At the end of the experiment the distribution of fine roots of both species was examined by means of root trenching. TheG. robusta root system extended almost symmetrically to the edges of the plot in the uncropped part, but only to some 4 m from either side of the hedge where cropped. Part of the upper profile to 1.3 m was shared with the maize roots, butGrevillea rooted further down to at least 2 m. as well as rooting deeper. The possible implications of such niche differentiation for below-ground resource capture are noted.     

Tree-crop interactions in hedgerow intercropping with different tree species and tea in Sri Lanka: 2. Soil and plant nutrients

The main aim of this study is to review the environmental and socioeconomic sustainability of the gum arabic farming system in central Sudan. A further aim is to analyse some of the main factors influencing production in recent decades in order to understand the future trade potential and consequently the smallholder livelihood. The study shows that end-user imports of gum arabic have increased during recent decades. Gum arabic is mainly for uses such as soft drinks, confectionary, and pharmaceuticals. However, even with this increased demand the production in Sudan, the main country of production, is declining. The producers, mainly smallholders, suffer from fluctuating prices. If the gum arabic farming system should be able to provide the environmental benefits of improved soil fertility and the socioeconomic benefits of risk spreading and dry season income opportunities, the prices paid to smallholders must be stabilized at a fair level, otherwise a shift to other crops or practices might take place.     

Contour hedgerow intercropping in the mountains of China: a review

Hedgerow intercropping systems were introduced in China in early 1990s. Achievements in research and extension of contour hedgerow intercropping in China during the past 15 years are reviewed here. Results reported in over 70 published papers have shown that hedgerow intercropping contributes to soil and water conservation, soil fertility amelioration, land productivity improvement, bio-terrace formation, and gives more options for income generation based on local resources in mountain areas. Research and demonstration works on contour hedgerow systems have achieved success by integrating local resources and needs into the system, especially in the dry valleys of the upper reaches of the Yangtze River, and the Three Gorges region. Contour hedgerow intercropping has attracted the attention of researchers, policy-makers, and farmers, and has been taken as an alternative to implementation of the Grain for Green policy, and ecological reconstruction and restoration today. To date, hedgerow intercropping has been demonstrated and applied practically on sloping land in more than six provinces of China, particularly Sichuan, Guizhou, Shanxi, Shaanxi, as well as in the Three Gorges region of Chongqing and Hubei Province. The intercropping system has also been practiced as an optimized technology for conserving farming on sloping lands, improving cash income, and reducing agricultural risks in depressed mountainous regions in southwest and northern China over recent years. Some misunderstandings and problems in studies and extension of the system in China are summarized and clarified, and some recommendations for further research and expansion of the system are also presented in this paper.     

Cost-benefit analysis of alternative forms of hedgerow intercropping in the Philippine uplands

Considerable resources have been expended promoting hedgerow intercropping with shrub legumes to farmers in the Philippine uplands. Despite the resources committed to research and extension, persistent adoption by farmers has been limited to low cost versions of the technology including natural vegetation and grass strips. In this paper, cost-benefit analysis is used to compare the economic returns from traditional open-field maize farming with returns from intercropping maize between leguminous shrub hedgerows, natural vegetation strips and grass strips. An erosion/productivity model, Soil Changes Under Agroforestry, was used to predict the effect of erosion on maize yields. Key informant surveys with experienced maize farmers were used to derive production budgets for the alternative farming methods. The economic incentives revealed by the cost-benefit analysis help to explain the adoption of maize farming methods in the Philippine uplands. Open-field farming without hedgerows has been by far the most popular method of maize production, often with two or more fields cropped in rotation. There is little persistent adoption of hedgerow intercropping with shrub legumes because sustained maize yields are not realised rapidly enough to compensate farmers for establishment and maintenance costs. Natural vegetation and grass strips are more attractive to farmers because of lower establishment costs, and provide intermediate steps to adoption. Rural finance, commodity pricing and agrarian reform policies influence the incentives for maize farmers in the Philippine uplands to adopt and maintain hedgerow intercropping. This revised version was published online in August 2006 with corrections to the Cover Date.     

Insect pests and beneficial arthropods population under different hedgerow intercropping systems in semiarid Kenya

Exploiting the principles of biological control of crop pests is an important management strategy for the resource-poor smallholder farmers of the tropics. The effects of nine hedgerow species on the abundance of major insect pests of beans and maize, and predatory/parasitic-arthropods were monitored over two cropping seasons and the intervening dry period. The tree/shrub species evaluated were Gliricidia sepium, Grevillea robusta, Senna siamea, Senna spectabilis, Flemingea congesta, Croton megalocarpus, Morus alba, Calliandra calothyrsus and Lantana camara. Arthropod abundance was monitored through counts of infested plants and using yellow pan and pitfall traps. Beanfly (Ophiomyia spp.) infestation was significantly higher in the presence of hedgerows (35%) than in their absence (25%). Hedgerows did not influence aphid (Aphis fabae) infestation of beans. In contrast, maize associated with hedgerows experienced significantly lower stalk borer (Busseola fusca and Chilo spp.) and aphid (Rhophalosiphum maidis) infestations than pure maize, the margin of difference being 13% and 11% respectively for the two pests. Ladybird beetles closely followed their prey, aphids, with significantly higher catches in sole cropped-plots than in hedgerow-plots and away from hedgerows. Activity of wasps was significantly greater close to hedgerows than away from them. Spider catches during maize season were 77% greater in the presence of hedgerows than in their absence, but catches during other seasons were similar between the two cropping systems. Differences among hedgerow species were not significant for most of the arthropods monitored, except that beanfly infestation was greater with Gliricidia hedgerows. The study indicates that the effect of hedgerows on pest infestations of crops and their role as refugia for predators cannot be generalized but depends on the specific arthropods.This revised version was published online in November 2005 with corrections to the Cover Date.     

Testing a process-based model of tree seedling growth by manipulating

A model was developed that simulated photosynthesis, growth and allocation in tree seedlings. The model was parameterized with data from experiments on seedlings of sycamore (Acer pseudoplatanus L.), Sitka spruce (Picea sitchensis (Bong) Carr.) and young birch trees (Betula pendula Roth.). In these experiments, CO2 concentration ([CO2]) and nutrient addition rate were varied. Parameters quantifying nutrient uptake, translocation and starch synthesis were fitted, based on data from control treatments. Elevated [CO2] and low-nutrient treatments were then used to test the predicted response of growth and allocation against observations. The model accurately predicted total seedling growth in the elevated [CO2] treatments. A response of growth to elevated [CO2] was seen in the birch and sycamore experiments, but not in the Sitka spruce, because of photosynthetic down-regulation. Predictions of allocation were reasonably accurate in the birch and Sitka spruce experiments, but were notably poorer in the sycamore experiments, possibly because of differences in sink strength between root and shoot. In the birch and sycamore experiments, little change in allocation with elevated [CO2] was observed or predicted. This was ascribed to the relative values of K(Tc) and K(Tn), the translocation coefficients that determine the sensitivity of allocation to carbon and nitrogen uptake rates, respectively. Growth and allocation in the low-nutrient treatments were poorly predicted by the model. In Sitka spruce, it was suspected that the photosynthetic parameters measured in August 1994 had been higher earlier in the season, before nutrients became depleted. In sycamore, the discrepancies were thought to relate to differences in sink strength between root and shoot that could not be described by the model.     

The effect of harvest tree distribution on harvesting productivity in selection cuts

Abstract

The study explored the effect of tree-selection pattern on harvesting productivity, under specific site conditions typical of mountain forestry and suitable for mechanized cut-to-length harvesting. In particular, the experiment tested two different single-tree-selection principles, namely, uniform spatial distribution across the whole site and clustered distribution, where selection trees were concentrated along presumed strip roads. The test was conducted on two different sites, representative of two different stand-development stages. Ten plots were located on each site, and randomly assigned to the two spatial distribution treatments (uniform and clustered). Great care was taken to maintain even conditions for the test. Mean tree size and removal intensity were the same on all plots from the same site; all plots were harvested by the same harvester and forwarder team; data were collected by the same researchers in all plots. Under the specific conditions of the study, the spatial distribution of harvest trees had no significant effect on harvesting productivity and cost. In contrast, harvesting performance was affected by tree size and possibly removal intensity. Anecdotal evidence about the effect of spatial distribution on harvesting cost may derive from its possible association with the other above-mentioned factors (e.g. tree size or removal intensity).     

A cost-benefit analysis of hedgerow intercropping in the Philippine uplands using the SCUAF model

Soil erosion in the Philippine uplands is severe. Hedgerow intercropping is widely advocated as an effective means of controlling soil erosion from annual cropping systems in the uplands. However, few farmers adopt hedgerow intercropping even in areas where it has been vigorously promoted. This may be because farmers find hedgerow intercropping to be uneconomic compared to traditional methods of farming. This paper reports a cost-benefit analysis comparing the economic returns from traditional maize farming with those from hedgerow intercropping in an upland community with no past adoption of hedgerows. A simple erosion/productivity model, Soil Changes Under Agroforestry (SCUAF), is used to predict maize yields over 25 years. Economic data were collected through key informant surveys with experienced maize farmers in an upland community. Traditional methods of open-field farming of maize are economically attractive to farmers in the Philippine uplands. In the short term, establishment costs are a major disincentive to the adoption of hedgerow intercropping. In the long term, higher economic returns from hedgerow intercropping compared to open-field farming are realised, but these lie beyond farmers limited planning horizons.     

A study of the potential of hedgerow intercropping in semi-arid India using a two-way systematic design

The potential of hedgerow intecrropping with Leucaena leucocephala was explored on vertic Inceptisols over 4 years at ICRISAT Center, Patancheru, India. The study was conducted using a systematic layout involving different alley widths ranging from 1.35 to 4.95 m and with varying distances between hedge and crops. The alleys were cropped with alternate rows of sorghum and pigeonpea. Hedges composed double Leucaena hedgerows 60 cm apart were periodically harvested for fodder. Sole crops of all components and a sorghum/pigeonpea intercrop were included in all four replications of the study.Starting in the second year, Leucaena was progressively more competitive to annual crops, causing substantial yield reduction. Competition (primarily for moisture) was most severe in narrow alleys and was greatest on pigeonpea.The growth of Leucaena was not sufficient to compensate for reduced crop yields. Land equivalent ratios (LERs) calculated on the basis of grain yield of crops and Leucaena fodder yields showed that hedgerow intercropping (HI) was advantageous over sole crops only during the first two years using wide alleys, but disadvantegeous in the last two years. LERs calculated on the basis of total dry matter indicated only a small advantage for HI (13–17 percent) over sole crops in wider (>4 m) alleys. Average returns per year from HI exceeded those of the most productive annual crop system (sorghum/pigeonpea intercropping) by 8 percent in 4.05 m alleys, and by 16 percent in 4.95 m alleys. Fodder production during the dry season was 40 percent of the annual total in these alley widths. Thus hedgerow intercropping at 4–5 m alley width is not very attractive for farmers in semi-arid India, which has 600–700 mm of annual rainfall. There is a need to examine the potential of HI in wider alleys. The merits and limitations of the systematic design are discussed.     

Estimation of net gain of soil carbon in a nitrogen-fixing tree and crop intercropping system in sub-Saharan Africa: results from re-examining a study

Nitrogen (N)-fixing tree and crop intercropping systems can be a sustainable agricultural practice in sub-Saharan Africa and can also contribute to resolving climate change through enhancing soil carbon (C) sequestration. A study conducted by Makumba et al. (Agric Ecosyst Environ 118:237?C243, 2007) on the N-fixing tree gliricidia and maize intercropping system in southern Malawi provides a rare dataset of both sequestered soil C and C loss as soil carbon dioxide (CO₂) emissions. However, no soil C gain and loss estimates were made so the study failed to show the net gain of soil C. Also absent from this study was potential benefit or negative impact related to the other greenhouse gas, nitrous oxide (N₂O) and methane (CH₄) emissions from the intercropping system. Using the data provided in Makumba et al. (Agric Ecosyst Environ 118:237?C243, 2007) a C loss as soil CO₂ emissions (51.2?±?0.4?Mg?C?ha^?1) was estimated, amounting to 67.4% of the sequestered soil C (76?±?8.6?Mg?C?ha^?1 in 0?C2?m soil depth) for the first 7?years in the intercropping system. An annual net gain of soil C of 3.5?Mg?C?ha^?1?year^?1 was estimated from soil C sequestered and lost. Inclusion of the potential for N₂O mitigation [0.12?C1.97?kg?N₂O?CN?ha^?1?year^?1, 0.036?C0.59?Mg CO₂ equivalents (eq.) ha^?1?year^?1] within this intercropping system mitigation as CO₂ eq. basis was estimated to be 3.5?C4.1?Mg CO₂ eq.?ha^?1?year^?1. These results suggest that reducing N₂O emission can significantly increase the overall mitigation benefit from the intercropping system. However, significant uncertainties are associated with estimating the effect of intercropping on soil N₂O and CH₄ emissions. These results stress the importance of including consideration of quantifying soil CO₂, N₂O and CH₄ emissions when quantifying the C sequestration potential in intercropping system.     

Growth characteristics of multipurpose tree species,crop productivity and soil properties in agroforestry systems under subtropical humid climate in India

Multipurpose tree species （MPTs） were studied in an agroforestry arboretum under subtropical humid climate in Northeast India. Out of 12 MPTs planted under agroforestry systems, Acacia auriculiformis in spacing of 2 m × 2 m （2500 stems·hm^-2） could have the potentiality to meet the timber/fuelwood requirement due to its high wood production of 635 m^3·hm^-2 with mean annual increment （MAI） of 2.54×10^-2 m^3.treel.a^-1 in a short rotation period of 10 years. Thus, A. auriculiformis is a short rotation forest tree species suitable to grow in subtropical humid climate. On the other hand, at 16 years of age, Eucalyptus hybrid and Michelia champaca in spacing of 3 m × 3 m （1111 stems.hm^2） produced appreciably high timber volume of 315 m^3.hm^-2 and 165 m^3.hm^-2 with MAI of 1.77×10^-2 m^3.tree^-1·a^-1 and 0.92×10.2 m^3.tree^-1.a^-1, respectively. At 16 years of age, Gmelina arborea produced a timber volume of 147 m^3.hm^-2 with MAI of 1.47×10^-2 m^3.tree^-1.a^-1 followed by Samania saman （140 m^3.hm^-2）, Albizziaprocera （113 m^3·hm^-2） and Tectona grandis （79 m3.hm^-2） with MAI of 1.40, 1.13 and 0.78 × 10^-2 m^3 .tree^-1a^-1, respectively in 4 m × 4 m spacing （625 stems.hm^-2）. Gliricidia maculata and Leucaena leucocephala could be used as live fences around the farm boundary to supply their N-rich leaves for mulch as well as manure to crops. In agroforestry arboretum, direct seeded upland rice （Oryza sativa - variety, AR-11）, groundnut （Arachis hypogaea - variety, JL-24） and sesamum （Sesamum indicum - variety, B-67） were grown during the initial period upto 8 years of tree establishment. Under other MPTs, there was a reduction in crop productivity as compared to open space. After 8 years of tree establishment, horti-silvi and silvi-pastoral systems were developed and pineapple （Ananas comosus - variety Queen）, turmeric （Curcuma longa -variety RCT -1） and cowpea （Vigna sinensis - variety Pusa Barsati） as forage crop were raised. The productivity of p     

印地安潮湿亚热带的多种用途树种的生长特性以及农林系统中作物产量和土壤特性(英文)

在印地安具有亚热带潮湿气候特点的东北地区,本文对几种多种用途的树种进行研究。选择12种多用途树种种植在农林系统中,耳果相思(Acacia auriculiformis),种植间距为2m×2m(2500 个·hm-2),有很大的生长潜力。在 10 年期的短期轮伐中,每年有很高的产量(635 m3·hm-2),平均每年增涨量为 2.54×102 m3·tree-1·a-1,可以满足薪碳材的需要。所以,耳果相思(Acacia auriculiformis)是一种短期轮伐树种,适合种植在潮湿亚热带气候地区。另一方面,16年树龄的桉树(Eucalyptus hybrid)和黄兰(Michelia champaca) 种植间距为3m×3m (1111个·hm-2),分别有较高的生长量(315 m3·hm-2和165 m3·hm-2),平均每年每棵树增长量分别为1.77×10-2 m3和0.92 × 10-2 m3。16年树龄的印度石梓(Gmelina arborea) 木材产量为147 m3·hm-2,每年每树的增长量是 1.47×10-2 m3。其次为 Samania saman(140 m3·hm-2)和柚木(Tectona grandis (79 m3·hm-2)),平均每年每树的增长量分别是 1.40,1.13和0.78×10-2 m3。毒鼠豆属植物(Gliricidia maculate) 和银合欢(Leucaena leucocephala) 可以用作农厂的围栏并提供农作物的肥料。在植物园,种植树木的最初 8 年间,可以种植高地水稻(Oryza sativa–variety, AR-11), 落地花生(Arachis hypogaea–variety, JL-24)和胡麻(Sesamum indicum–variety,B-67)。由于林下郁闭度太大,在耳果相思(Acacia auriculiformis)林下的农作物产量不高,与空旷地的农作物比较产量有所降低。树木种植 8 年后,种植草料作物如,凤梨、姜黄、豇豆,它们的产量在印楝(Azadirachta indica)林下很高。高产量的农作物和园艺植物与具有高产量多种用途树种一起种植构成了有发展前景的农林系统,并改变土壤特性,改善土壤酸度,提高土壤有机质,降低土壤腐蚀度,提高土壤含水量。     

Testing the safety-net role of hedgerow tree roots by 15N placement at different soil depths

Trees which root below crops may have a beneficial role in simultaneous agroforestry systems by intercepting and recycling nutrients which leach below the crop rooting zone. They may also compete less strongly for nutrients than trees which root mainly within the same zone as crops. To test these hypotheses we placed highly enriched ¹⁵N-labelled ammonium sulphate at three depths in the soil between mixed hedgerows of the shallow-rooting Gliricidia sepium and the deep rooting Peltophorum dasyrrhachis. A year after the isotope application most of the residual ¹⁵N in the soil remained close to the injection points due to the joint application with a carbon source which promoted ¹⁵N immobilization. Temporal ¹⁵N uptake patterns (two-weekly leaf sub-sampling) as well as total ¹⁵N recovery measurements suggested that Peltophorum obtained more N from the subsoil than Gliricidia. Despite this Gliricidia appeared to compete weakly with the crop for N as it recovered little ¹⁵N from any depth but obtained an estimated 44–58% of its N from atmospheric N₂-fixation. Gliricidia took up an estimated 21 kg N ha^–1 and Peltophorum an estimated 42 kg N ha^–1 from beneath the main crop rooting zone. The results demonstrate that direct placement of ¹⁵N can be used to identify N sourcing by trees and crops in simultaneous agroforestry systems, although the heterogeneity of tree root distributions needs to be taken into account when designing experiments. This revised version was published online in June 2006 with corrections to the Cover Date.     

石家庄市园林绿化树种的选择与配置

本文对石家庄市园林绿化树种的选择和配置进行了阐述,认为树种的选择和配置应以乡土树种、速生树种、乔木树种为主,同时注重选择环保型、彩叶型、果树型树种,并加强野生树种、引进树种的研究和开发力度,加强苗圃建设和苗木培育工作。     

杨农复合经营系统碳储量及其分布特征研究

以杨树纯林和杨农复合林2种生态系统为研究对象,对林木、农作物(林下植被)、土壤、凋落物各组分的碳储量进行比较研究。结果表明,杨树纯林和杨农复合林的碳储量分别为(101.59±8.1),(116.65±10.3)t/hm2。在2个生态系统中林木及土壤碳储量均占总量的90%以上。杨农复合林凋落物和林下植被碳储量所占比例约为6%,是杨树纯林的2倍。林木碳储量、农作物(林下植被)碳储量均为杨农复合林较高;杨农复合林土壤碳储量仅在中间层(10~25,25~50 cm)显著高于杨树纯林;凋落物碳储量2者之间差异不显著。相对纯林,农林复合系统具有较高的吸收和固碳能力。     

广州市绿化树种选择与配置的研究

根据广州市自然地理状况及目前绿化树种选择与配置上存在的问题,提出了广州市树种选择、配置的原则,探讨了树种选择、配置的思路。重点阐述了城区绿化,其中包括城区街道,公共绿地、单位附属绿地、居住区、工业区绿化和城郊绿化,其中包括道路防护林带、林区绿化的思路。     

Stand basal area as an index of tree competitiveness in timber intercropping

To derive optimal benefits from intercropping timber, farmers should make important initial decisions on tree species and planting density with a good understanding of their tradeoffs. Complex and data-intensive models used by researchers should be supplemented with simpler models based on easily measured parameters and easily understood competition functions. In experiments in the Philippine uplands, growth parameters of three popular farm-forestry species (Eucalyptus deglupta, E. torelliana, and Paraserianthes falcataria) were measured, along with intercropped and non-intercropped yields of maize and vegetables. The commonly used forestry parameter of stand basal area had a significant negative correlation with intercrop yields (as a percentage of non-intercropped yields). The slope of the regression line differed between species; in this study, percent yield loss per unit stem basal area growth was in the order E. deglupta > E. torelliana > P. falcataria. The relationship between stand basal area and intercrop-yield decline was tested on an independent data set from China. Intercrop yields had significant negative correlations with stand basal area of Paulownia elongata. We propose that adaptive tree-screening trials evaluate competitiveness in addition to evaluating growth and mortality. Stand basal area may be better suited to this task than more mechanistic indices such as leaf-area index as it is easy to measure, calculate, and understand, and it may serve as a better index of total (aboveground + belowground) competition. Basal area is also directly related to tree volume, and allows farmers to more easily evaluate the economic tradeoffs between tree growth and intercrop-yield declines. This revised version was published online in June 2006 with corrections to the Cover Date.     

Tree leafing phenology and crop productivity in semi-arid agroforestry systems in Kenya

The hypothesis that temporal separation of resource use between trees and crops minimises competition for wa ter in agroforestry systems during the cropping period and increases utilisation of annual rainfall was tested at Machakos in semi-arid Kenya. Four popular tree species were chosen to provide a range of leafing phenologies. These included Melia volkensii, which sheds its leaves twice a year, Senna spectabilis and Gliricidia sepium, which shed their leaves during the long dry season, and the evergreen Croton megalocarpus. All four species retained their foliage during the long rains, offering little scope for temporal separation of resource use. Maize (Zea mays) yields were reduced by 50–70% in the agroforestry treatments. Reductions in crop yield were strongly correlated with tree growth (r ² =0.94) and available soil moisture (r ² =0.88). G. sepium remained leafless for much of the short rains despite the presence of available soil water, and was least competitive with the bean crops (Phaseolus vulgaris) grown at this time. Reductions in crop yield in the agroforestry treatments were closely correlated with tree growth (r ² =0.99) and available moisture (r ² =0.79) during the 1996/97 short rains (158 mm), but not during the much wetter 1997/98 season (608 mm). Shading by trees or shade nets reduced crop yield, in contrast to previous studies in the semi-arid tropics. Low off-season rainfall during the study period (9% of annual rainfall compared to the long-term average of 20%) limited the potential for temporal separation of growing periods. Where the prospects for temporal or spatial separation in resource use are limited, shoot and/or root pruning may be necessary to manage competition between trees and crops. This revised version was published online in June 2006 with corrections to the Cover Date.