The relation between canopy spread cover and the aboveground available phytomass of Nama Karoo subshrubs and grasses

The canopy spread cover of Karoo bushes and grasses in the Nama Karoo was measured and related to the aboveground phytomass available to the grazing animal. A correlation coefficient of r = 0.7456, calculated for Karoo bushes, indicates that this method can be used to non-destructively estimate available phytomass. This in turn can be used for the prediction of current grazing capacity in areas where Karoo bushes are dominant in the vegetation. In the case of the Karoo grasses, a rather loose definition and interpretation of the canopy spread cover, led to a low correlation coefficient, r = 0.5662. This renders the present method largely unsuited for available phytomass estimations to be used in further predictions of grazing capacity in the mountainous areas where the studied grasses are dominant in the vegetation. In the more arid areas, however, the method can be used to advantage.     

Effect of continuous grazing in the dohne sourveld on species composition and basal cover

Abstract

In a grazing trial conducted at the Dohne Agricultural Research Station, natural grassland was grazed continuously for 36 years with cattle. In a second investigation conducted on similar grassland, paddocks were grazed or rested, in all combinations of the four seasons of the year, over a ten year period. Changes in botanical composition and cover of the sward were monitored at regular intervals. The results of these investigations indicate that under a regime of non‐selective grazing the composition of Dohne Sourveld is remarkably stable. After 36 years of continuous grazing the species composition of the sward was largely maintained. Floristically Dohne Sourveld is dominated by a small number of grass species of which Andropogon appendiculatus, Elionurus muticus, Heteropogon contortus, Sporobolus africanus, Themeda triandra and Tristachya leucothrix are the most important. These species react differently to under‐utilization or increasingly severe defoliation, enabling them to be classified into decreaser or increaser categories. Severe defoliation resulted in a marked decrease in basal cover but only small changes in relative species composition. As the intensity of defoliation increases, decrease in grass cover is associated with an increase of forbs, such as Senecio retrorsus.     

温性典型草原羊草光合特性对模拟放牧因素分解的响应

放牧是一种复合胁迫，包括采食、践踏、粪尿返还等因素。为明晰放牧对草原影响的过程与机制，通过刈割、人工模拟踩踏、人工粪尿添加来模拟家畜放牧过程，以羊草为例，研究了株高、盖度、干重以及羊草叶片的光合特性对放牧分解因子的响应。结果表明，采食及包含采食因子的处理显著降低了羊草的盖度、高度及干重，但提升了羊草叶片的净光合速率（P_n）、蒸腾速率（T_r）、气孔导度（G_s）、胞间CO₂浓度（C_i），降低了羊草叶片的水分利用效率（WUE）。羊草株高与羊草叶片净光合速率、气孔导度、胞间CO₂浓度、蒸腾速率显著负相关，与水分利用效率显著正相关；土壤含水量与气孔导度、胞间CO₂浓度、蒸腾速率显著正相关。相对而言，践踏和粪便以及它们的交互作用对羊草的影响比采食弱。综合可得，家畜的采食是放牧对温性典型草原羊草光合特性影响的主要因子。     

Effects of selective defoliation and height of defoliation on tiller dynamics and herbage yield of Themeda triandra in a semi-arid rangeland in Zimbabwe

Understanding the impacts of intensity and selection of grazing on the performance of grasses is important in the management of grazing areas. It is especially important in semi-arid environments where, apart from moisture, the levels at which grasses are utilized has a major influence on their persistence in the environment. The effects of selective defoliation and height of defoliation (5 cm and 10 cm stubble heights) on the performance of the grass, Themeda triandra, were investigated in a field experiment for two growing seasons. Performance was measured as tiller production, rate of production, tiller mortality, herbage yield and quality. Tiller production was greater (81 tillers per plant) under non-selective defoliation than under selective defoliation (49 tillers per plant) in the 1995/1996 season. Tiller mortality was higher (66.45%) under heavy selective defoliation than under nonselective defoliation (21.98%). Herbage yield, apart from the control treatment, was high (13.6 g per plant) under light non-selective defoliation. Heavy selective defoliation reduced the nutrient levels (e.g. levels of soluble carbohydrates under heavy selective defoliation were 6 g kg^?1 glucose compared to 20 g kg^?1 glucose under light non-selective defoliation).     

Measuring the impact of indigenous herbivores: reply to Kerley andLandman

Pteronia paniculata is an indigenous, unpalatable shrub that invades mismanaged Karooveld, resulting in degraded rangelands with low species diversity and grazing potential. We conducted a series of trials in the Succulent Karoo Randteveld near Barrydale to determine if the uniform defoliation of P. paniculata dominated vegetation at two heights (0.05 m and 0.20 m above ground level) using a brush-cutter and, in one trial, application of a second cut will improve the plant species composition, productivity and grazing capacity of the veld. Brush-cutting treatments and the uncut control all resulted in a change in species composition towards greater species diversity and more palatable species and an average increase of 540 kg ha^?1 (28%) in above-ground biomass over four years. It appears that there was a pervasive improvement in species composition associated with a general decline in the cover and abundance of P. paniculata over the time-scale of the present study that was not influenced by the defoliation treatments, except for the 1996-cut treatment where the cover of P. paniculata increased. The absence of propagules of palatable species in the soil seed bank and competition from P. paniculata (a long-lived, perennial shrub) are assumed to be among the main reasons for the lack of response of the vegetation community to the defoliation treatments. Brush-cutting (in the absence of reseeding), aimed at reducing the dominance of unpalatable karoo shrubs, was more costly but not significantly better than long-term resting in improving veld composition or forage production.     

Vegetation gradients around cattleposts in the eastern mountains of Lesotho

The grasslands of the mountains in eastern Lesotho support large numbers of sheep, angora goats and other livestock during the summer, during which they are corralled nightly by herders at cattleposts. Few studies, however, have quantified the impact of grazing on the composition and structure of the mountain grasslands. Gradients of grazing and trampling intensity related to distance from 23 cattleposts in the subalpine and alpine belts (2 650–3 175 m above sea level) were examined to describe the long-term effects of livestock on unpalatable dwarf shrubs, bare area and plant species composition. The potential for shrub invasion and bare areas (low herbaceous cover) to develop was highest near cattleposts. Species composition also changed significantly along the grazing gradient, accompanied by a shift in dominants from grazing-sensitive to grazing-resistant grasses and shrubs. Species either decreased or increased monotonically along the grazing gradient or had an intermediate, unimodal response. The relative positions of species’ centroids along the grazing gradient were used to quantify their grazing sensitivity on a scale from 10 (most sensitive) to 0 (most resistant), which confirmed and contradicted previous classifications of their grazing responses. Basic ecological research on this important mountain grazing and catchment system is recommended.     

Drafting a Conservation Blueprint: A Practitioner's Guide to Planning for Biodiversity — Craig R Groves 2003

Individual forage species were appraised in varying distances along transects radiating from the water points of the ranch and traditional cattle post management systems. Measurements of plant height were used to assess the degree of forage species utilisation by livestock around water points. Livestock use forage plants more heavily near water points and the degree of use is more pronounced in the traditional cattle post areas, compared to ranch grazing. Utilisation of annual plants occurring at the 0m zone from water was greater than that of the same plant species growing at further distances. Periodic closure of water points aimed at reducing grazing pressure has been indicated as a method to promote production of forage around water points. With the exception of Panicum maximum, the abundance of which was low or non-existent in both management systems, Digitaria eriantha was the most utilised grass by livestock, followed by Schmidtia pappophoroides and Eragrostis lehmanniana. Stipagrostis uniplumis, one of the dominant grasses in the Kalahari Sandveld, exhibited a weak tendency to decline as distance from water increased. It was observed that high forage utilisation rates coincide with drought periods, implying that correct stocking rate adjustments should always match the available forage. Forage utilisation from water points in the traditional cattle post areas extended beyond 4 000m, especially during the dry periods.     

环县典型草原放牧家畜践踏的模拟研究

践踏是放牧家畜对草地的三大影响因素之一,也是导致草原全面退化的主要原因。但是由于技术和方法的限制,相关报道较少,极大地削弱了对草地农业生态系统的主体-草畜界面生态动力机制的整体认识。基于此,在我国干旱半干旱的黄土高原典型草原区兰州大学环县草畜生产系统野外实验点,通过滩羊夏季轮牧试验践踏强度的跟群观测、践踏同质性试验、模拟降水与试验践踏双因子试验进行了为期2年的系列研究。     

The grazing index method of range condition assessment

Abstract

Owing to the difficulty of examining succession theory in the Karoo, it is suggested the ecological index method (EIM), be replaced by the grazing index method (GIM), through the introduction of grazing index values (GIV) for Karoo plant species. The GIM may provide more acceptable range condition scores and more realistic estimates of the current grazing capacity. Using GIVs, arrived at by scoring the agronomic attributes of the plant species (productivity, forage value, perenniality), the estimated current grazing capacities are found to be closely similar to the grazing capacity norms established for the different agro‐ecological areas in the Karoo. The manner in which the range condition survey is conducted and the application of the grazing index method, ensures the computation of a figure representative of the agronomic potential of the area to support livestock. The cover figure and the range condition score determined for a sample site, can be employed in monitoring the trend in changes in the vegetation, brought about by climatic and biotic variables. Grazing capacity norms are closely related to median annual precipitation (r² = 0.98). The GIM is more closely related to grazing capacities than the EIM (r² = 0.96 and 0.68 respectively).     

Web spider abundance is affected by sheep farming in the Karoo§

Impacts of livestock farming include effects on arthropods and a range of related ecological processes. Grazing by sheep may indirectly affect web spider populations by changing vegetation structures available for web construction, or directly by trampling the spiders or their webs. We tested these two potential impacts by surveying spiders along transects and found that web spider abundance was 52% lower in a sheep grazing area compared with inside the adjacent Tierberg-LTER (Long Term Ecological Research) sheep exclosure. This reduction in web-spider abundance could be due to the 10% lower shrub cover in the sheep pasture than in the exclosure. Further support for the hypothesis that web spider abundance is affected by shrub cover came from spider abundance being higher on densely vegetated mounds, termed heuweltjies, than in the surrounding shrub matrix in both the exclosure and the sheep pasture, with heuweltjies showing cross-fence differences, consistent with grazing impacts. In addition, there may be a direct effect of trampling, as suggested by experimental removal of webs. By depressing spider populations, sheep grazing may affect a number of ecological factors, such as trophic relationships or nutrient cycling.     

山地牧道与植被空间异质性研究评述

山地牧道和沿牧道放牧改变植被空间异质性。综合国内外研究进展,将山地牧道的涵义扩展为家畜在山地垂直空间移动的固定路线、放牧往返小道、相对固定的采食路径以及践踏小径等,并总结了山地牧道的形成与分布规律,重点评述了山地牧道与植被空间异质性相互作用机制。山地牧道作为一种线状廊道,为物种扩散提供通道、生境,分割斑块景观,影响植被边界动态,在不同尺度上影响植被空间异质性;家畜沿牧道的选择采食与植被空间异质性相互影响,家畜还通过践踏和排泄改变植被空间异质性;家畜采食、践踏和排泄综合作用于土壤,通过影响土壤异质性间接影响植被的空间分布。山地牧道在动物行为学、放牧生态学研究中具有理论和实践价值,可以为监测植被退化、控制载畜量提供依据。     

Livestock grazing has minimal effect on the species richness and diversity of semi-arid shrubland in the Western Little Karoo,South Africa

The Succulent Karoo, one of two arid biodiversity hotspots in the world, is known for its high plant species richness, but little is known about the influence of topography and how it mediates the potentially deleterious effects of grazing. Changes in vegetation species composition, cover and species diversity were examined along piosphere gradients on northerly slopes, bottomlands and low-lying plains on 45 farms. Landscapes differed in the plant speciesand life-form composition, species richness and cover, with the more species-rich, heterogeneous grassy northerly slopes distinct from the plains and bottomlands, which were dominated by Pteronia pallens, Psilocaulon junceum and Drosanthenum spp. with more annual and ephemeral species. Overall, species richness declined linearly with decreasing rangeland condition. A weak grazing effect was detected only on the north-facing slopes, where shrubs and grass decreased with intense grazing. Years of overgrazing have resulted in the widespread dominance of P. pallens, especially on the plains and bottomlands, rendering them insensitive to grazing. The current condition of the northern slopes should be maintained and managed such that palatable species can spread to more degraded areas of the landscape.     

Influence on a North‐Eastern Sandy Highveld sward of winter utilization by sheep

North‐Eastern Sandy Highveld was utilized selectively by sheep grazing alone in winter with a supplementary lick. Winter utilization in four consecutive years had no detrimental effect on production, cover or seed production but care should be exercised in recommending this practice because of the tendency to start “winter” grazing in the critical autumn months and to continue grazing into spring.     

Remote Sensing of Spatial and Temporal Vegetation Patterns in Two Grazing Systems

One constraint that range scientists must face in grazing studies is the lack of accurate and repeatable techniques for discriminating grazing effects from both temporal variability and spatial heterogeneity of vegetation. Both forms of variability contribute to inconsistent grazing system effects on vegetation response and forage production in semiarid ecosystems. Remote sensing may be an efficient tool for detecting differences in spatial and temporal patterns of grazing impact on vegetation. The purpose of this study was to evaluate the spectral data derived from satellite images as a tool for comparing grazing system impacts on spatial and temporal vegetation patterns. We evaluated the effect of two grazing systems, “Continuous” (C) and “Two-Paddocks Rest-Rotation” (TPRR), on vegetation cover from 1996 to 2006 in a semiarid ecosystem of Argentina. We compared grazing effects on vegetation cover using two indices derived from the Normalized Difference of Vegetation Index (NDVI) data from Landsat Thematic Mapper images. We observed a slight advantage in NDVI improvement for the TPRR over the C. Even though, in both grazing systems, an upward vegetation trend occurred only in areas located far from the watering points, TPRR showed higher relative vegetation cover near the watering point than C. We consider this methodology an important step for monitoring vegetation changes and making management decisions in livestock systems of semiarid regions because grazing system impacts may be compared for both spatial and temporal vegetation patterns. However, we think that the key next step is to develop procedures that discriminate between forage and nonforage components.     

Long-term changes in land use,land cover and vegetation in the Karoo drylands of South Africa: implications for degradation monitoring§

We used several large data sets at a range of temporal and spatial scales to document the land-use/land-cover change (LULCC) dynamics of the semi-arid Succulent Karoo and Nama-Karoo biomes of South Africa. More than 95% of the Karoo is comprised of land classified as Natural, which has been relatively stable since 1990. Over the last 100 years cultivation, as well as the number of domestic livestock, has declined significantly in both biomes. Protected areas have increased since 1980 to comprise nearly 8% of the Succulent Karoo biome, although they only cover 1.6% of the Nama-Karoo biome. There has been a significant recent increase in renewable energy installation applications, which cover 4% of the Karoo drylands. The trend in vegetation productivity (NDVI; 1982–2015) is unchanged over 90% of both biomes, while nearly 10% of the Karoo has shown a significant increase in NDVI trend. An analysis of repeat photographs shows that vegetation cover has either remained unchanged or has increased at most locations. Although the Karoo drylands appear less degraded than they were in the mid-twentieth century, on-going monitoring at different temporal and spatial scales is essential to evaluate the future impact of LULCC on these semi-arid environments.     

Grazing and Burning Japanese Brome (Bromus Japonicus) on Mixed Grass Rangelands

Japanese brome (Bromus japonicus Thunb. ex Murr.) is an introduced, annual cool-season grass adapted to the central and northern Great Plains. Japanese brome has negatively impacted perennial grasses and decreased seasonal animal gains. Prescribed spring burning and defoliation have been effective in reducing brome density or cover, but little information directly compares the two common strategies. The objectives of this study were to 1) compare annual spring burning and grazing to reduce Japanese brome populations; and 2) evaluate trends of vegetative composition and biomass in burned, grazed, and unburned rangelands infested with Japanese brome. Paddocks with Japanese brome were assigned to one of four treatments: 1) annual prescribed spring burning, 2) spring grazing, 3) a combination of annual spring burning and grazing, and 4) an idle control. Treatments were applied annually from 2000 to 2004. Japanese brome density was greatest in the idle control in all years, even when low winter and spring precipitation limited Japanese brome recruitment. Late spring Japanese brome density was similar in all treatments with grazing or burning in four of the five seasons. Spring burning resulted in less than 65% litter cover the last 3 years, whereas the idle control and spring grazing had over 80% litter cover the last 4 years. Western wheatgrass (Pascopyrum smithii [Rydb.] A. Löve) decreased with spring grazing in burned and unburned paddocks. Buffalograss (Bouteloua dactyloides [Nutt] J. T. Columbus) composition decreased in the idle control treatment. Blue grama (Bouteloua gracilis [Willd. ex Kunth] Lag. ex Griffiths) and sideoats grama (Bouteloua curtipendula [Michx.] Torr.) composition varied by year. Even though annual burning and spring grazing were equally effective in limiting Japanese brome density and biomass compared to the idle control, Japanese brome was still present after 5 years, which indicates the difficulty of eradicating Japanese brome from ecosystems where it has become naturalized.     

A study of animal production under different grazing regimes in the False Upper Karoo at Grootfontein

Three systems of animal production on the fragile Karoo veld, i.e. multi-paddock rotational grazing, pauci-paddock rotational grazing and continuous grazing were studied. Of these systems, multi-paddock rotational grazing gave the best results with continuous stocking resulting in the worst. All three systems studied had long histories of both continuous stocking- and of pauci-paddock rotational grazing. The veld, under both these previous systems of management has stabilised, but at different seral stages. The continuously stocked range stabilised in the early developmental seral stage, consisting mainly of annual grasses. The veld under pauci-paddock rotational management stabilised in the early to mid-developmental seral stage, consisting mainly of perennial grasses. Experimentation on veld in a stable condition is a prerequisite if valid conclusions in respect of rotational grazing and continuous stocking are to be drawn. Rotational grazing on overgrazed veld will result in poor animal performance, while it is generally accepted that continuous stocking on pristine veld will result in excellent animal performance for a number of years. Multi-paddock rotational grazing outperformed both pauci-paddock rotational grazing as well as continuous stocking on both types of veld. This is surprising taking into account the mid-seral stage condition of the vegetation. This supports the argument that with improved veld management, an improved production potential is possible on veld even in a degraded or current condition.     

Development of veld management research in the karoo region

The development of veld research in the semi‐arid Karoo areas since 1934 is outlined. It entailed key experiments on different types of veld, and the synthesis of grazing systems and their testing in experimental camps and co‐operatively on a practical farming scale. The experiments were primarily conducted with Merino sheep but Angora goats, Boer goats, cattle and Persian sheep were also employed. A large number of rotational grazing systems, special treatments and methods have been set out for the entire Karoo Region. The group camp approach serves as a basis for complete veld management planning in the Karoo Region.

Supporting research covered plant physiological, phenological and ecological studies; the determination of selective grazing habits of livestock; production of karoo bushes; and the development of research techniques especially in connection with quantitative and qualitative vegetation surveys.     

陇东黄土高原典型草原滩羊践踏强度与放牧强度关系的研究

通过在环县典型草原不同放牧强度下轮牧试验的跟群观测,发现放牧强度增加3倍,其践踏强度却增加5倍多,说明践踏作用对放牧的敏感性高于以家畜采食作用为基础的适宜载畜量和放牧强度,获得了家畜践踏强度随放牧强度幂函数式增大的定量结论,说明调控放牧家畜的践踏效应应是放牧管理的一项重要的指标。