Plant and sward response to patch grazing in the Highland Sourveld

Abstract

Selective grazing by livestock inevitably results in patch grazing. This study was initiated to determine the effects of patch grazing on (1) the vigour of a single grass species, Themeda triandra, and of the sward, and (2) sward species composition. The generally held idea that a full season's rest followed by an early spring burn would prevent preferential grazing of patch grazed areas which had developed in the seasons before the rest was also tested. The vigour of 71 triandra was estimated from etiolated growth of marked tufts while sward vigour was indexed by above‐ground herbage production (AGHP). Etiolated growth of T. triandra and AGHP of the sward within patches were negatively affected by three seasons of grazing, but a full season's rest appeared sufficient to restore both the etiolated growth of T. triandra and the AGHP of the sward to a level similar to that in the non‐patches. A full season's rest followed by spring burning did not, however, prevent preferential grazing of grazed patches which had developed in the seasons prior to the rest. Species composition within patches (characterised by Increaser II species) differed significantly from the species composition of non‐patches (characterised by Increaser I species). Patch grazing may therefore initiate the rangeland degradation process in Highland Sourveld and patch grazing may be the focus from which rangeland degradation proceeds.     

Patterns of plant selection by grazing cattle in two savanna grasslands: A plant's eye view

The amount of grazing experienced by individual tufts of the most abundant perennial grasses of two Transvaal savannas was monitored at regular intervals for two growing seasons. In general, Setaria incrassata and Themeda triandra were heavily grazed, Bothriochloa insculpla and Aristida bipartita were lightly grazed, and Heteropogon conlortus and Digitaria eriantha were intermediate. Tufts with a greater amount of moribund material were less heavily grazed. The smallest tufts of all species were the most lightly grazed, although the most heavily‐grazed tufts were on occasion intermediate in size rather than being the largest. A tuft had an increased likelihood of being heavily grazed if it had previously been heavily grazed, both within a growing season and for two successive growing seasons. A spatial pattern of grazing was evident, as individual species were less heavily grazed at different locations within the paddocks. The influence of species identity on the amount a tuft was grazed was not absolute but rather quantitative and conditional on the level of other variables. Tuft size and the amount a tuft had been previously grazed generally had a greater influence on the amount a tuft was grazed during an individual grazing event than species identity, location or moribund material. Species selection is possibly, in part, a consequence of selection for plant structure which optimizes intake.     

Grass species selection patterns on rotationally‐grazed Dohne Sourveld during autumn and early winter

Herbaceous species preference was studied during autumn and winter periods of occupation, on rotationally‐grazed Dohne Sourveld, at four different stocking rates. Only the most abundant grass species in the sward were monitored. Animals (cattle and sheep) grazed selectively (for plant species) during both autumn and winter, but the degree of selectivity increased in winter. Although animals could be forced to graze less selectively by increasing the stocking rate in autumn, stocking rate did not affect the sequence of species selection, but merely accelerated the process. Regrazing of tillers during grazing periods was a common occurrence in all but the heavy stocking rate treatment. Although Tristachya leucothrix was the most preferred species, the amount (mass) of this species that was grazed per unit basal tuft area was less than that of Themeda triandra.     

Forage production of some grass species in the false Thornveld of the Eastern Cape

Abstract

The most commonly occurring grass species in the False Thornveld (Acocks, 1953; veld type no. 21) were clipped before and after grazing on veld in good and poor condition at different times of the year. Herbage yield per species was expressed per unit basal tuft area, and the difference in yield before and after grazing was considered to reflect the production of available forage. Themeda triandra produced more forage than any other species at all times of the year. Sporobolus fimbriatus generally produced the second highest amount of available forage. Cymbopogon plurinodis, despite having the second highest herbage yield before grazing, was not grazed to a significant extent. On the basis of the results, Themeda triandra and Sporobolus fimbriatus were identified as key grass species in the False Thornveld of the Eastern Cape.     

Growth and tiller production of Themeda triandra as affected by NPK fertilisation

Themeda triandra Forssk. is a common perennial grass in southern African grasslands that tolerates low nutrient soils. Establishment of the species into degraded or transformed areas is difficult, but there have been some advances with clonal propagation using tillers. To generate propagules (tillers), tiller production of plants must be maximised. This research focused on the growth response of T. triandra to fertilisation with nitrogen (N), phosphorus (P), and potassium (K) each at three levels in a fully factorial pot experiment. Fertilisation increased growth. The number of tillers and aerial shoot mass were controlled by an N × P interaction, and total mass and root mass were controlled by N and by P, with N having a larger effect. Mass per tiller was not related to treatment. Shoot:root ratio and root mass per tiller were controlled by N. Potassium had no effect on any response. For maximum tiller production of T. triandra, plants should be fertilised with high levels of N and P.     

Evaluation of the grazing potential of grass species in Tarchonanthus veld of the northern cape 1

Abstract

The grazing value of the seven most prominent grass species in Tarchonanthus‐limeveld was investigated from 1977 to 1981.

The frequency of utilization of Chrysopogon serrulatus was exceptionally high (93%), while Sporobolus fimbriatus, Themeda triandra and Digitaria seriata were intermediate (50–60%) followed by Cymbopogon plurinodis (34%). Eragrotis lehmanniana and Fingerhuthia africana were poorly grazed (12 to 16%). The utilization of species was influenced by stocking rate. Statistical differences between species indicated a relatively high biomass production (600 to 740 g DM/m² per year) for E. lehmanniana, C. plurinodis and T. triandra, while F. africana, S. fimbriatus and D. seriata were intermediate (320 to 420 g DM/m² per year). E. lehmanniana, C. plurinodis and T. triandra produced the highest above‐ground phytomass, but had a lower crude protein content (2,9 to 7,4%) and digestibility (46,1 to 61,3%) than the other four species (3,8 to 9,7% crude protein and 50,5 to 68,4% digestibility).     

Effect of continuous grazing in the dohne sourveld on species composition and basal cover

Abstract

In a grazing trial conducted at the Dohne Agricultural Research Station, natural grassland was grazed continuously for 36 years with cattle. In a second investigation conducted on similar grassland, paddocks were grazed or rested, in all combinations of the four seasons of the year, over a ten year period. Changes in botanical composition and cover of the sward were monitored at regular intervals. The results of these investigations indicate that under a regime of non‐selective grazing the composition of Dohne Sourveld is remarkably stable. After 36 years of continuous grazing the species composition of the sward was largely maintained. Floristically Dohne Sourveld is dominated by a small number of grass species of which Andropogon appendiculatus, Elionurus muticus, Heteropogon contortus, Sporobolus africanus, Themeda triandra and Tristachya leucothrix are the most important. These species react differently to under‐utilization or increasingly severe defoliation, enabling them to be classified into decreaser or increaser categories. Severe defoliation resulted in a marked decrease in basal cover but only small changes in relative species composition. As the intensity of defoliation increases, decrease in grass cover is associated with an increase of forbs, such as Senecio retrorsus.     

Die invloed van die beskikbaarheid van nutriënte en ontblaringsbestuur op die produksie van digitar1a Eriantha variëteit Eriantha

The influence of heavy grazing by cattle for three years, and of rainfall interception for one year, on the herbaceous composition of a sandveld savanna in Gazankulu was investigated. The composition and the abundance of the predominant species changed substantially, owing mainly to rainfall. Pogonarthria squarrosa, considered to increase under grazing, was initially the most abundant species, but declined most rapidly on the heavily‐grazed, rainfall interception treatment, owing to a high mortality. Digitaria eriantha increased markedly on the heavily‐grazed treatment, despite up to a 24% tuft mortality, because of extensive vegetative recruitment from stoloniferous growth. Aristida congesta subsp. barbicollis and Aristida stipitata, considered to increase under heavy grazing, increased on the lightly‐grazed treatment. There was an explosion of annual forbs, predominantly Tephrosia purpurea, Indigofera costata and Cassia absus, in the third year. The trend of rainfall during the study, initially wet and successively drier, apparently had a greater influence on compositional trends than the imposed treatments.     

The influence of tiller age and time of year on the growth of Themeda tri andra and Sporobolus fimbriatus in semi‐arid grassveld

Abstract

Sequential growth measurements were made on successive populations of Themeda triandra and Sporobolus fimbriatus tillers initiated at different times of the year. The independent influences of tiller age and time of year on tiller growth were isolated by deriving growth indices for tillers of the same age at different times of the year and for tillers of all ages at each time of the year respectively.

Isolating the effect of plant age on growth showed that T. triandra is most productive under a regime of long periods of undisturbed growth, while S. fimbriatus is better suited to frequent interruptions in growth. Isolating the effect of time of year shows that both species are particularly sensitive to perturbations in spring. However, the photosynthetic efficiency of T. triandra is strongly influenced by time of year while S. fimbriatus seems to be a more opportunistic assimilator of sunlight.     

The importance of the ecological view of agriculture

Abstract

Marked populations of Themeda triandra and Sporobolus fimbriatus tillers were sequentially harvested at fortnightly intervals both under irrigation and under unamended moisture conditions. Sampling covered a period from tiller initiation through to senescence after flowering.

Tiller masses and green leaf areas were used to derive absolute growth rates, relative growth rates, net assimilation rates and leaf area ratios respectively. Growth rates of Themeda triandra were higher, or were maintained at high levels for longer than those of Sporobolus fimbriatus. Moisture stress resulted in considerably depressed growth rates. Photosynthetic efficiency of spring emerging tillers was potentially greatest in midsummer, but this seasonal influence was masked by moisture stress on non‐irrigated treatments.     

Effect of three grazing intensities on selected soil properties in semi‐arid west Texas

Abstract

Soils in the study, carried out in the Texas Trans‐Pecos region, were fine‐loamy, mixed, thermic Aridic Calciustolls; fine‐loamy, mixed, thermic Aridic Ustochrepts; and fine, mixed, thermic Typic Chromusterts. Our primary objective was to determine the effect of three grazing intensities within an intensive, short‐duration grazing system on selected soil physical properties. Maximum compaction occurred in the A2/AB horizons, which has significantly higher (P<0.05) penetrometer readings than the overlying A I/A horizons under all grazing intensities. Heavily grazed areas also have significantly higher (P<0.05) penetrometer reading than the other two areas in both horizons. Heavily grazed areas also have significantly lower (P<0.05) infiltration rates, saturated hydraulic conductivities, and water‐stable aggregates than soils grazed at the other two levels. Aggregates in the A2/AB horizon were more stable than aggregates in the Al and Bw horizons in every pedon under all grazing intensities. When the effect of grazing intensity was analysed for each horizon separately, the lightly grazed areas had significantly lower (P<0.05) bulk density and higher total porosity in the A2/AB horizons than comparable horizons from the other two sites. However, when all horizons were analysed together, no significant differences in bulk density were found among treatments and horizons. No significant differences were observed in percentage organic carbon among treatments.     

The dynamics of Themeda Triandra tillers in relation to burning in the natal Drakensberg

Abstract

Marked tillers of Themeda triandra were sampled regularly over a two year period in Highland Sourveld burnt annually in winter, biennially in spring and after a single summer and early winter burning treatment. Parameters recorded were height of shoot apex, tiller mass, number of new lateral tillers and time of flowering. Monitoring of marked tillers before and after each burn enabled the life history of populations of tillers to be followed.

There were no significant differences in survivorship between tillers in the annual winter and biennial spring burn treatments but winter burning stimulated the development of a greater number of daughters per parent. Both populations exhibited a constant death rate over the two year period. Only 8% of the marked T. triandra tillers survived the single summer burning treatment.     

The effect of leaf age on the photosynthetic rate of themeda triandra forsk 1

Abstract

The apparent photosynthesis of progressively older leaves of Themeda triandra was recorded in a controlled environment by means of gas exchange measurements. Photosynthetic rate increased with increasing leaf age and was maximal in the third fully expanded leaf. Thereafter, photosynthesis dropped sharply, but the photosynthetic rate of the oldest green leaf was still 55 % of that of the most efficient leaf. The implication is that old green leaves can probably still play an important role in initiating regrowth after defoliation of younger leaves.     

A flexible system of management for sourveld in Natal

The conventional systems of veld management have a number of shortcomings, the main one being that the period of absence from grazing during the growing period is normally too short. This is due to the fact that a particular group of animals is allocated to a specific set of camps for the whole growing season.

A more flexible system of veld management would allow the grazing procedure to be manipulated in such a way that the period of absence could be varied in relation to the rate of regrowth of the grass.

The application of such a flexible system would involve the following: (a) the minimum number of camps required is calculated on the basis of at least four to five camps for each group of animals; (b) one quarter of the area of veld is rested; (c) the remaining camps are grazed whenever sufficient regrowth has taken place, and no group of animals is confined to particular camps or set of camps; (d) each camp is well grazed down and the number of camps grazed for the season will depend on weather conditions; during a dry year, more camps will be grazed and fewer camps during a wet year; (e) the camps grazed most selectively, are rested during the following season; and (f) burning of the veld is confined to those camps that have been rested during the previous year.     

Ontogeny of Themeda triandra tillers in the false thornveld of the eastern cape

Abstract

Marked populations of Themeda triandra tillers were sequentially sampled at fortnightly intervals under normal conditions and without moisture stress respectively, covering a period from tiller initiation through to senescence after flowering. Parameters recorded were height of shoot apex, tiller mass, leaf area, leaf number, secondary tiller development and premature tiller mortality. Tillers of irrigated plants had a significantly higher mass, more elevated shoot apices, and produced a greater total number of leaves than those of non‐irrigated plants. Fewer unstressed tillers senesced prematurely than those experiencing moisture stress.     

Effects of selective defoliation and height of defoliation on tiller dynamics and herbage yield of Themeda triandra in a semi-arid rangeland in Zimbabwe

Understanding the impacts of intensity and selection of grazing on the performance of grasses is important in the management of grazing areas. It is especially important in semi-arid environments where, apart from moisture, the levels at which grasses are utilized has a major influence on their persistence in the environment. The effects of selective defoliation and height of defoliation (5 cm and 10 cm stubble heights) on the performance of the grass, Themeda triandra, were investigated in a field experiment for two growing seasons. Performance was measured as tiller production, rate of production, tiller mortality, herbage yield and quality. Tiller production was greater (81 tillers per plant) under non-selective defoliation than under selective defoliation (49 tillers per plant) in the 1995/1996 season. Tiller mortality was higher (66.45%) under heavy selective defoliation than under nonselective defoliation (21.98%). Herbage yield, apart from the control treatment, was high (13.6 g per plant) under light non-selective defoliation. Heavy selective defoliation reduced the nutrient levels (e.g. levels of soluble carbohydrates under heavy selective defoliation were 6 g kg^?1 glucose compared to 20 g kg^?1 glucose under light non-selective defoliation).     

The effect of defoliation and competition on the regrowth of Themeda triandra and Aristida junciformis subsp. junciformis

Abstract

The effects of a cutting trial, conducted to study the combined influence of defoliation and competition from surrounding vegetation on the vigour of Themeda triandra and Aristida junciformis, were evident after the single season of treatment implementation. Aristida junciformis was more sensitive to defoliation than T. triandra, whereas competition reduced yield to a greater extent in T. triandra than in A. junciformis. The two species differed in their ability to tolerate defoliation under competition. Defoliation reduced the yield of A. junciformis, at all levels of competition, to a greater extent than T. triandra. Aristida junciformis was particularly sensitive to a combination of severe defoliation and full competition. It is suggested that manipulation of grazing patterns may shift the competitive balance between palatable (e.g. T. triandra) and unpalatable (e.g. A. junciformis) components of the sward.     

Herbage intake and ruminal digestion of dairy cows grazed on perennial ryegrass pasture either in the morning or evening

This study aimed to clarify diurnal fluctuations of herbage intake, ruminal fermentation of herbage carbohydrates and proteins, and digesta particulate weight in the rumen of grazing dairy cows. Six ruminally cannulated, non‐lactating dairy cows were grazed on perennial ryegrass/white clover pasture either in the morning (04.00 to 08.00 hours) or the evening (16.00 to 20.00 hours). Cows grazed in the evening spent more time (P < 0.01) and consumed more herbage (P < 0.01) compared with cows grazed in the morning. Higher (P < 0.05) daily mean concentrations of total volatile fatty acid, propionate and n‐butyrate in rumen fluid were observed for cows grazed in the evening compared with cows grazed in the morning. Although cows grazed in the evening ingested more crude protein compared with cows grazed in the morning, no significant difference in NH₃‐N concentration in rumen fluid was observed between them. The ratio of purine‐derivative concentration to creatinine concentrations was higher (P < 0.01) in the urine of cows grazed in the evening than in cows grazed in the morning. These results clearly indicated that evening grazing was advantageous for dairy cows compared with morning grazing, in terms of ruminal fermentable energy intake and nitrogen utilization efficiency.     

Pathways of Grazing Effects on Soil Organic Carbon and Nitrogen

Grazing modifies the structure and function of ecosystems, affecting soil organic carbon (SOC) storage. Although grazing effects on some ecosystem attributes have been thoroughly reviewed, current literature on grazing effects on SOC needs to be synthesized. Our objective was to synthesize the effects of grazing on SOC stocks in grasslands, establishing the major mechanistic pathways involved. Additionally, and because of its importance for carbon (C) biogeochemistry, we discuss the controls of soil organic nitrogen (N) stocks. We reviewed articles analyzing grazing effects on soil organic matter (SOM) stocks by comparing grazed vs. ungrazed sites, including 67 paired comparisons. SOC increased, decreased, or remained unchanged under contrasting grazing conditions across temperature and precipitation gradients, which suggests that grazing influences the factors that control SOC accumulation in a complex way. However, our review also revealed some general patterns such as 1) root contents (a primary control of SOC formation) were higher in grazed than in their ungrazed counterparts at the driest and wettest sites, but were lower at sites with intermediate precipitation (~400 mm to 850 mm); 2) SOM C:N ratios frequently increased under grazing conditions, which suggests potential N limitations for SOM formation under grazing; and 3) bulk density either increased or did not change in grazed sites. Nearly all sites located in the intermediate precipitation range showed decreases or no changes in SOC. We grouped previously proposed mechanisms of grazing control over SOC into three major pathways that can operate simultaneously: 1) changes in net primary production (NPP pathway), 2) changes in nitrogen stocks (nitrogen pathway), and 3) changes in organic matter decomposition (decomposition pathway). The relative importance of the three pathways may generate variable responses of SOC to grazing. Our conceptual model suggests that rangeland productivity and soil carbon sequestration can be simultaneously increased by management practices aimed at increasing N retention at the landscape level.     

Effects of eight grazing systems on carcass mass gains of steers grazing irrigated Cynodon aethiopicus pastures

Abstract

Eight grazing systems were compared on irrigated Cynodon aethiopicus pastures which received 650 kg N, 36 kg P and 200 kg dolomitic limestone/ha, annually. There were either one, two, three, six or 18 paddocks in the systems, the single paddock system being continuously grazed. The others were rotational systems with various periods of grazing and of resting.

Using the comparative slaughter technique carcass mass gains of crossbred Hereford/Afrikaner steers were calculated. Three‐year mean carcass mass gains over stocking rates from 12 to 18 steers/ha ranged from 872 to 997 kg/ha. None of the rotational systems were better than continuous grazing.

Estimates of amounts of herbage available for grazing in each system showed that there was least in the continuous grazed and most in the multi‐paddock systems where there were long periods of rest between successive grazing.