THE USE OF LIVEWEIGHT-GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

The problems of designing, conducting and interpreting the results of grazing trials to evaluate tropical pasture mixtures are discussed in relation to an experiment wbich measured the effect of three fixed stocking rates upon two grass/legume pastures. A uniformity trial prior to grazing indicated considerable soil variability, which necessitated ground replication. Sequential grazing of three pasture replicates at a fixed stocking rate and a fixed interval of grazing resulted in sites of low fertility being heavily penalized. Such a form of replication was of value for measuring the effect of treatments upon sward composition, and, to a lesser extent, variation in animal production, within the experiment area. Differences in animal production between pasture species were small when compared with the effect of various stocking rates, emphasizing the need for an objective assessment of stocking rate when evaluating pasture species. Fixed-stocking-rate designs have more general application in the tropics because they are easier to conduct, and the results are repeatable and can be extrapolated to farm practice.     

THE INFLUENCE OF SEVERAL FACTORS IN DETERMINING THE GRAZING BEHAVIOUR OF BORDER LEICESTER × MERINO SHEEP

Over a period of 20 months twenty continuous, 24-hour, observations were made on the grazing behaviour of Border Leicester × Merino sheep. Distinct seasonal changes were found in the pattern of grazing. Grazing usually began during the dawn period and decreased at dusk. In spring and summer there were two peaks of grazing, but there was only one peak in autumn and winter. The percentage of grazing taking place between 6 a.m. and 6 p.m. varied with season, and was highest in spring and early summer. These seasonal changes in the pattern of grazing appeared to be independent of grazing pressure, or of the nutritional requirements of the sheep.
The mean grazing time, for all observations, was 8·2 hours with a standard deviation of 0·97. Decreasing pasture availability increased grazing time, as shown by an increased intensity of grazing.
Ruminating times were found to be considerably lower than have been reported for cattle. There was a significant negative correlation between ruminating time and the nitrogen content of available pasture.
There was no clear evidence that the number of lambs carried before parturition, or suckled after parturition, had any effect on grazing time.
There was no evidence that the behaviour of one sheep influenced that of another, even of a twin, grazing nearby.
Consistent differences between sheep were found for ruminating time. The variation in grazing time between sheep, at any one time, was considerably greater than previously reported and differences between sheep were not consistent.
Variation in grazing time appeared to be less in flocks of less than 6 sheep than in flocks of 6 to 12 sheep.     

THE EFFECT OF WINTER GRAZING BY SHEEP ON SPRING AND EARLY SUMMER PASTURE PRODUCTION

Studies were made in S.W. Scotland of the effect of winter grazing (October to March) by sheep on subsequent spring and early summer pasture production from a sward in its first harvest year and from a permanent pasture. Averaged over the two swards, DM yields in April, May and June were reduced by 38%, 8% and 5%, respectively, as a result of various times and frequencies of grazing, compared with no grazing. Reductions were greatest after grazings in the January to March period, especially grazings involving the month of March. In a trial in which fertilizer N was used to compensate for March grazing, 50 kg NJha raised DM yields in April to the level of those from no grazing, while only 15 kg NJha were needed to raise June yields to parity. Controlled winter grazing of sheep and the use of fertilizer N to restore production losses after early spring grazing are justified.     

THE EFFECT OF NITROGEN, STOCKING RATE AND GRAZING METHOD ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in 1971 and 1972 to study the effects of two levels of fertilizer N (50 and 504 kg/ha) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and three at the higher. In addition to rotational grazing, set-stocking and an integrated grazing-conservation system were included. The responses/kg N were approximately 1 kg live-weight gain, 19–24 Meal ME and 8–9 kg DM. Maximum yields of 1200–1300 kg live-weight gain/ha and 25,000–27,000 Meal ME/ha were recorded. Productivity of set-stocked pasture was similar to rotational grazing at the high N level, but lower at low N. The integrated system yielded comparable results to high-N rotational grazing. Performances and herbage intakes per animal reflected the stocking rates imposed.     

THE EFFECT OF NITROGEN, STOCKING RATE AND FREQUENCY OF GRAZING BY BEEF CATTLE ON THE OUTPUT OF PASTURE

Two successive grazing experiments were conducted over 12 weeks on perennial ryegrass pastures with 50 and 44 young cattle to study the effect of N fertilizer when applied at a daily rate of 1 or 3 kg N/ha. At each level of N two stocking rates differing hy 20% were imposed. At the liigher N level and stocking rate, three frequencies of grazing were imposed. At the stocking rates imposed N tended to reduce the daily liveweight gain per head, but increased the total liveweight gain per ha by from 0.79 to 1.58 kg/kg N. A 20% increase in stocking rate depressed gain per head in both years. In the first year it did not improve gain/ha, hut in the second year gain/ha was increased by 12–17%. There was a tendency for performance per animal and per ha to increase as the grazing cycle was lengthened. In 1969 the highest yield in the whole season was 1880 grazing days and 1260 kg gain/ha. The overall response to N fertilizer was similar to that recorded in other reports, but it is possible that a lack of K had limited pasture growth.     

THE SIMULATION OF ANIMAL RETURNS IN GRASSLAND EXPERIMENTATION

An experiment was carried out on a grass/white-clover sward at North Wyke to evaluate the use of mixtures of inorganic and organic nutrient sources to simulate the nutrient effect of animal excreta, both with and without overall PK fertilizer. Grazing with sheep and the return of excreta from caged sheep both increased herbage yields, as did artificial return treatments. Artificial return treatments giving 75% of the nutrient level returned by excreta gave yields comparable with grazing. The effect of the simulated return treatments was not influenced by PK application. Grazing gave a very different botanical composition to that from the treatments in wbich natural or simulated excreta were returned to cut swards. Mixtures simulating excreta may replace the nutrient effect of the grazing animal, but not its physical effect, in some forms of grassland experimentation. They are not recommended for fertilizer trials.     

THE EFFECT OF DUNG AND URINE AND ITS INTERACTIONS WITH APPLIED NITROGEN, PHOSPHORUS AND POTASSIUM ON THE CHEMICAL COMPOSITION OF PASTURE

The importance of the grass/clover balance of a pasture in determining the effects of manurial returns and fertiliser applications on the chemical composition of the sward is shown.
The response to applied nitrogen was very dependent on the presence or absence of the animal excrements. For example, the N recovery from an application of 18 cwt. per acre of nitro-chalk was only 2% in the absence but 68% in the presence of the grazing animal.
Urine caused a marked increase in the N and K content and the yield of pasture, particularly when returned in quantity to a high-producing grass-dominant sward, and at the same time tended to depress the Ca, P and Mg content. Urine K was superior to fertiliser K, per pound of K returned or applied, in raising the K content of the pasture.
By comparison, dung had little direct effect on chemical content; even when returned in large amounts it affected the Ca and K content of the pasture to a relatively small extent. The P returned in the dung had little effect on the P content of the pasture and was inferior to superphosphate in this respect.
Together, as in normal grazing, dung and urine tended to counter or enhance their individual effects.
There was a positive correlation between the Mg and P content of the pasture and a negative correlation between Mg and K. Brief mention is made of the possible importance of the nutrient balance in pasture as a factor in animal health.     

THE USE OF LIVEWEIGHT-GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

An experiment is described in which the liveweight gains of cattle, grazing different pasture species, were measured when grazing sequences were arranged in the form of a Latin Square. The average coefficient of variation for the 6 "animal" Latin squares used was 28·5%, with a range of 15·1 to 42·6%. Significant improvements in liveweight gain were demonstrated arising from the inclusion of Stylosanthes gracilis in swards of Chloris gayana and Panicum maximum .
The problems of conducting and interpreting trials of this type are discussed in relation to the need for data on the economic value of pasture species in the tropics. Latin square grazing designs offer a useful means of measuring animal production from pasture, with the advantage of requiring relatively small numbers of animals in each trial, but trials should be repeated at different sites.     

A NOTE ON THE ZERO-GRAZING OF BEEF CATTLE

Groups of 8 steers weighing 300–400 kg (660–880 Ib) were rotationally grazed on a ryegrass-dominant pasture for 115 days. The systems compared were zero grazing (Z), field grazing conducted at the same stocking rate (FC), and field grazing conducted at a stocking rate varied with the intention of giving the same liveweight gain per uiimal as zero grazing (FV). Mean daily liveweight gains were: Z, 0.98; FC, 0.78; FV, 0.90 kg/animal (2.2, 1.7 and 20 Ib/animal), and liveweight gains/unit area were in the ratio 100:78:85. Organic-matter intake, measured on four occasions, was, on average, Z, 6.54; FC, 6.18; FV, 687 kg/head daily (14.4, 13.6 and 15.1Ib). From these results it appears that a comparison of zero grazing and field grazing made at the same stocking rate is likely to underestimate the potential of field grazing for beef production from grass.     

THE USE OF LIVEWEIGHT–GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

A grazing experiment is described in which the liveweight gains of small East African Zebu heifers were measured from Hyparrhenia rufa/Stylosanthes gracilis and H. rufa/Centrosema pubescens swards. The design of this trial enabled animal production to be assessed from variations in liveweight gain since stocking rate was maintained at the same level on both pastures, but the overall stocking rate was varied according to seasonal changes in herbage availability.
Production was measured both by using individual animals within a group as independent units, and by true animal replication. Gains from individual animal blocks were shown to be of value for comparing pasture species, provided that the composition of the grazing group was varied. Problems of conducting and interpreting small–scale grazing trials as means for evaluating pastures are discussed.     

The potential for control of the soft rush (Juncus effusus) in grass pasture by grazing goats

Two experiments were conducted to examine the effect of inter-tussock grass height and stocking rate on the utilization of the rush ( Juncus effusus ) by grazing goats. In the first experiment, on rush-infested Festuca rubralTrifolium repens pasture, the utilization of rushes by goats grazing at a sward height of 3–4cm or 5–6cm was compared with that occurring on plots grazed by sheep at a sward height of 3–4 cm. Sheep grazed minimal amounts of rush. In contrast, it was estimated that 90% and 75% of current seasons growth of rush was grazed by goats at sward heights of 3–4 and 5–6 cm respectively during the first year. With continued goat grazing at 5–6cm there was a dramatic reduction in the cover and vigour of the rushes, and at a sward height of 3–4cm established tussocks were eliminated from the pasture.
The second experiment compared the utilization of rushes invading predominantly Agrostis swards stocked with goals at 10, 20 or 30 ha^-1 and in which inter-tussock sward height was maintained at 4–5cm on all treatments, by adding or subtracting sheep. On one site rush utilization increased with the increase in stocking rate of goats and rush tussocks were eliminated within 3 years at 30 goats ha^-1. On another, there was no difference between plots stocked at 20 or 30 goats ha^-1 and viable tussocks remained. The influence of the composition and productivity of the inter-tussock herbage and the proportion of rush in the biomass are discussed.
Goats can be used lo control rushes in grassland but high stocking levels and low inter-tussock pasture heights are required to promote adequate levels of utilization.     

THE INFLUENCE ON THE FOLLOWING ARABLE CROP OF SHEEP EXCRETA AND FERTILIZER TREATMENTS APPLIED TO A LEY

The differences in soil fertility in a ley as a result of treatments applied for four years in a sheep-grazing trial were measured by growing crops of wheat and kale. The pasture treatments had been: control, dung, urine and dung plus urine, each combined with four levels of fertilizer nitrogen.
Yields of winter wheat harvested in 1956 ranged from 27.1 to 38.6 cwt. per acre. Plots to which faeces had been allowed to return during the pasture phase outyielded (p <0.001) those from which it was withheld. Urine was relatively ineffective except in combination with nitrogen. In the absence of animal returns, nitrogenous fertilizer depressed grain and straw yields. A top-dressing of K applied to the wheat in spring as a sub-treatment had no significant effect. The percentages of N and K in the grain were unaffected by the former pasture treatments, or by the K top-dressing.
A subsidiary small-plot experiment in which the above pasture treatments had been combined with P and K, each at two levels, was cropped with marrow-stem kale. In general, N applied to the pasture increased kale yields except where dung and urine had been withheld. Neither P nor K had a significant effect, except when combined. The leaf/stem ratio of the kale was reduced by applied N and by P (both p <0.05).
The pasture/arable-crop relationship is considered.     

Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

HERBAGE AND BEEF PRODUCTIVITY OF FERTILIZED PASTURES IN NORTHERN ONTARIO

The productivity of 2 fertilized sown pasture swards and of a 30-year-old pasture sod was evaluated under rotational and continuous grazing in 1956–62.The application of NP fertilizer (448 kg 11–48–0/ha every 3 years with 112 kg ammonium sulphate in the other years) gave the highest yields of DM and of beef, followed by the P treatment (1120 kg superphosphate/ha). Beef production showed only small responses to either the type of pasture sward, or the method of grazing. Seasonal distribution of beef production was slightly more even on the old sod. For the peak period of June-July, 68% of the gain was made on the old sod against 74% on the cultivated swards under rotational grazing and 79% under continuous grazing. DM/gain ratios were generally higher in wet summers than in dry ones. Conversely, estimated DM digestibility was lowest in wet summers.     

THE FREQUENCY OF DEFOLIATION OF INDIVIDUAL TILLERS IN A SET-STOCKED SWARD

A method designed to study the defoliation of individual tillers in a set-stocked sward is described. This was used on S23 ryegrass swards carrying 19 (medium-stocked) and 30 (heavily-stocked) sheep/ac.
The tillers in the heavily-stocked sward were defoliated, on average, every 7–8 days, and those in the medium-stocked sward every 11–14 days. The total green leaf length (GLL) of tillers grazed by the sheep was generally greater than that of tillers not grazed. On average 27 and 40% of the GLL of tillers in the medium- and heavily-stocked swards respectively was removed at each grazing. Older leaves on the grazed tillers were defoliated much less frequently than were younger leaves.
It is suggested that a rational analysis of the relationship between the grazing animal and the grazed sward cannot be made without more work of this nature.     

A PORTABLE-CORRAL TECHNIQUE FOR MEASURING THE EFFECT OF GRAZING INTENSITY ON YIELD, QUALITY AND INTAKE OF HERBAGE

A grazing experiment using four stocking rates of sheep, equivalent to 34, 45, 57 and 68 sheep/ha on an assumed 200-day grazing season, was conducted using a portable grazing corral technique. Plots were subjected to fotir grazing periods between mid May and early September. The effect of stocking rate on herbage yield and quality and the influence of these factors and feed intake on liveweights of the sheep were recorded. The mean daily herbage organic matter available over the trial was 9.6, 5.8, 3.3 and 23% of the total liveweight of the sheep at each of the four stocking rates. The highest grazing pressure led to the production of high-quality herbage, but also led to reduced productivity, low feed intake and liveweight losses. The most lenient grazing pressure failed to provide adequate herbage utilization. Even at the most intensive stocking pressure, only 66% of the herbage available to ground level was utilized in grazings after July. Intake results suggested that sheep of 45 kg liveweight required 1000–1200 g digestible organic matter per day to maintain body weight. Despite the higher in vitro digestibility of herbage on offer at the higher stocking rates, intake was limited through lack of herbage; a high degree of correlation existed between herbage availability and herbage intake over the four stocking rates and at all grazings. It is concluded that the portable corral technique is well suited for grazing studies and the assessment of sward response to varying stocking rates and is particularly useful where facilities for more extensive studies are limited.     

THE GRAZING ANIMAL AND SWARD PRODUCTIVITY

A trial designed to show the effects of animal excreta on sward productivity is described.
It was estimated that the grazing sheep returns in a season nutrients capable of immediate uptake by the plant equivalent to 5–6 cwt. nitro-chalk and 2–21/2 cwt. 60% muriate of potash per acre.
A large part of sward yield depended on animal return. Clover depression from the effects of animal excreta and, under mowing treatment, from the effects of fertilizer nitrogen, did not result in marked loss of transferred clover nitrogen. There appeared to be a gain in clover soil-nitrogen from a reduction of the clover stand.
A comparison is made between the restilts obtained from sheep excreta on the sward with those obtained from mowing treatment.
The interaction between animal or fertilizer nitrogen and the clover plant is discussed.     

REMOVAL OF THE EFFECTS OF UNEVEN GRAZING IN PASTURE EXPERIMENTS

An experiment is described in which uneven grazing by sheep introduced a bias in a white clover variety trial. The data were adjusted by covariance, using as the independent variable the position of the plots within the paddocks.
Attention is drawn to the errors that may result from uneven grazing and a comparison is made with errors due to transference of fertility. Means of avoiding similar occurrences are suggested.     

Sodium fluorescein as an internal tracer on the location of bovine urine patches in pastures

Sodium fluorescein (SF) was evaluated and validated as an internal marker in cattle for the location of urine patches in pastures. Three trials were carried out aiming at the following: evaluating the effect of dilution and volume of application on SF fluorescence on the environment; testing dosages and duration of excretion of SF administered orally to cattle; and validating the methodology for SF use on the location of cattle urine patches in pastures. The marker was tested in beef cattle kept on pastures under three grazing systems: degraded pasture under extensive management (De), intensively managed dryland pasture with high stocking rate (Id) and intensively managed irrigated pasture with high stocking rate (Ii). Besides the localization of urination sites, the number of urinations/animal/d and the area covered by urine were also determined. The residue of SF remained fluorescent in the pasture foliage up to 34 h after urination events, allowing the location of urine patches by a black‐light flashlight. There was no difference between grazing systems in the number of urine patches/animal/d during the rainy and dry seasons. The average number of urine patches was approximately 11/animal/d. As expected, the estimated volume and area covered by urine varied according to the stocking density. The chosen SF dose (50 mg kg^?1 LW) did not adversely affect the animals when administered once daily during 2 d. However, the same SF dose administered during four consecutive days caused urinary disorder in the animal. The distribution of urine patches was spatially dependent on specific characteristics of the paddock.     

THE USE OF LIVEWEIGHT-GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

A 21-acre grazing trial which ran continuously for three years to evaluate grass and grass/legume pasture with and without inorganic fertilizers is described. Techniques for conducting grazing trials where large treatment differences are expected and the interpretation of results from these trials are discussed. Good practical management of stock and pastures was shown to be necessary in these grazing trials and where it was only possible to establish one true replicate some measure of statistical control was achieved by comparing regression coefficients of cumulative animal production.
The results showed that up to 475 1b liveweight gain/acre per annum (532.48 kg/ha) could be obtained from pastures grazed by small East African Zebu steers. The addition of the legumes Stylosanthes gracilis and Centrosema pubescens to grass mixtures gave significant and economic increases in liveweight gain. Grass/legume swards receiving phosphorus and sulphur produced liveweight gains equivalent to the animai production from grass swards receiving 140 1b N/ac per annum (156.94 kg/ha).