Competition for light between hedgerows and maize in an alley cropping system in Hawaii, USA

Successful agroforestry systems depend on minimizing tree-cropcompetition. In this study, field experiments and a simulation model were usedto distinguish between tree-crop competition for light and belowgroundcompetition in an alley cropping system. Maize (Zea maysL.) was harvested periodically in three treatments: between vertical barriers ofshade cloth, hedgerows of Flemingia macrophylla (Willd.)Merr., and sole maize. Radiation intercepted by the maize was calculated using asimulation model based on measured values for direct and diffuse light, hedgerowdimensions and leaf area, and solar trajectory. Radiation use efficiency wascalculated as biomass production per unit of intercepted radiation. Maizebiomass and yield in both the alley crop and the shade cloth treatment weregreatest in the center of the alleys. Grain yield between hedgerows was 3.5Mg ha⁻¹ (averaged across the alley), significantlyless than in the shade cloth (7.4 Mg ha⁻¹) or thesole maize (7.7 Mg ha⁻¹) treatments. Lightintercepted by the maize in the alley crop was about half that intercepted bythe maize in the sole crop. The shade cloth intercepted less light than thehedgerows because it did not have an appreciable width. Radiation use efficiencyin the three treatments was 0.75 g mol⁻¹ PAR anddid not differ significantly among treatments. Tree-crop competition wasoverwhelmingly for light. This revised version was published online in June 2006 with corrections to the Cover Date.     

Competition for water in a pecan (Carya illinoensis K. Koch) – cotton (Gossypium hirsutum L.) alley cropping system in the southern United States

Understanding the belowground interactions between trees and crops is critical to successful management of agroforestry systems. In a study of competition for water in an alley cropping system consisting of pecan (Carya illinoensis) and cotton (Gossypium hirsutum) in a sandy loam soil (Rhodic Paleudult) in Jay, Florida, root systems of the two species were separated by trenching to 120 cm depth. A polyethylene barrier was installed in half of the plots. Spatial and temporal variations in soil water content, root distribution and water uptake by both species, and leaf area development and height of cotton were measured. Interspecific competition for water was greater in the non-barrier treatment near tree rows than at the alley center. Competition became evident 3 to 4 weeks after emergence of cotton and increased during the following 7 to 8 weeks. Compared with the non-barrier treatment, the barrier treatment had higher soil water content and better growth of cotton (height, leaf area, and fine root biomass). Cotton lint yield in the barrier treatment (677 kg ha^–1) was similar to that in a sole-crop stand, but higher than in the non-barrier (502 kg ha^–1) treatment. Lint production efficiency of plants was higher in the interior rows in the non-barrier treatment (0.197 kg lint per square meter of leaf area, compared to 0.117 kg in the barrier treatment). The results suggest that trenching or even deep disking parallel to the tree row may reduce competition for water, but the impact on tree growth cannot be established from this study. This revised version was published online in June 2006 with corrections to the Cover Date.     

Leucaena + maize alley cropping in Malawi. Part 1: Effects of N, P, and leaf application on maize yields and soil properties

Yields under alley cropping might be improved if the most limiting nutrients not adequately supplied or cycled by the leaves could be added as an inorganic fertilizer supplement. Three historic leaf management strategies had been in effect for 3 years ina Leucaena leucocephala alley cropping trial on the Lilongwe Plain of central Malawi : 1) leaves returned; 2) leaves removed; and 3) leaves removed, with 100 kg inorganic N ha⁻¹ added. An initial soil analysis showed P status to be suboptimal under all strategies. A confounded 3⁴ factorial experiment was conducted with the following treatments: leaf management strategy (as above), N fertilizer rate (0, 30, and 60 kg N ha⁻¹), P fertilizer rate (0, 18, and 35 kg P ha⁻¹), and maize population (14,800, 29,600, and 44,400 plants ha⁻¹). Both N and P were yield limiting, and interacted positively to improve yields. The addition of 30 kg N and 18 kg P ha⁻¹ improved yields similarly under all leaf management strategies by an average of 2440 kg ha⁻¹. Increasing the rates to 60 kg N and 35 kg P ha⁻¹ improved yields an additional 1990 kg ha⁻¹ in the ‘leaves returned’ and leaves removed + N’ strategies, but did not improve yields under the ‘leaves removed’ strategy. Lower yields were related to lack of P response at the highest P rate in this treatment, which may have induced Zn deficiency. Plots receiving leaves had higher organic C, total N, pH, exchangeable Ca, Mg, K, and S, and lower C/N ratios in the 0–15 cm soil layer than did plots where leaves had been removed. Leaf removal with N addition was similar to leaf removal alone for all soil factors measured except for organic C and total N, which were higher where N had been added. The results show that N and P were the primary yield-limiting nutrients. Historic N application maintained the soil's ability to respond to N and P on par with leaf additions.     

Pasture production and tree growth in a young pine plantation fertilized with inorganic fertilizers and milk sewage in northwestern Spain

The use of organic waste materials such as milk sewage as an organic fertilizer could have the dual advantages of organic-waste disposal and reduced dependence on inorganic fertilizers. The effects of fertilization with (1) conventional mineral fertilization, (2) milk sewage sludge at 40 kg N ha⁻¹ target rate and (3) no fertilization on pasture production and tree growth were examined in an experiment consisting of two pasture mixtures under a one-year-old Pinus radiata plantation with a density of 2500 trees ha⁻¹. The two pasture mixtures were: (1) Dactylis glomerata L. var. saborto (25 kg ha⁻¹) + Trifolium repens L. group Ladino (4 kg ha⁻¹) + Trifolium pratense L. var. Marino (1 kg ha⁻¹); (2) Lolium perenne L. var. Tove (25 kg ha⁻¹) + Trifolium repens L. group Ladino (4 kg ha⁻¹) + Trifolium pratense L. var. Marino (1 kg ha⁻¹). The experiment began in the spring of 1995 using a randomized block design with three replicates in Castro Riberas de Lea (Lugo, Galicia, north-western Spain). Plot size was 12 × 8 m², with a 1 m buffer strip between plots. Two-year data showed that fertilization with either material had a positive effect on pasture production, with no significant difference between the two fertilization treatments. Tree growth in the milk sewage sludge plot was significantly higher than in the control plots. Inorganic fertilization increased pasture production, but affected tree growth negatively. The results show that milk sewage sludge could be used as a fertilizer in silvo-pastoral systems. This revised version was published online in June 2006 with corrections to the Cover Date.     

Relative effects of irrigation and intense shade on productivity of alley-cropped tall fescue herbage

The relative effects of irradiance and soil water on alley-cropped herbage are poorly understood. Our objective was to determine effects of irrigation on herbage productivity when tall fescue [Lolium arundinaceum (Schreb.) Darbysh.] was grown in two sites, a meadow and a loblolly pine (Pinus taeda L.) alley (620 trees ha⁻¹), near Booneville, Arkansas. Three tall fescue entries were space planted in the meadow and pine alley with or without irrigation. Herbage mass and nutritive value were determined at 8-week intervals for 2 years. Mean daily PAR was 33.9 (2004) and 37.5 mol m⁻² d⁻¹ (2005) in the meadow, while the pine alley received 5.6 mol m⁻² d⁻¹ (17% of the meadow) in 2004 and 4.3 mol m⁻² d⁻¹ (11% of meadow) in 2005. Effect of tall fescue entry tended to be small relative to site and irrigation. Irrigation compensated for evapotranspiration in the meadow, but not in the pine alley when summer rainfall was about normal (2004) or low (2005). Nevertheless, site (PAR) had a greater effect on herbage specific leaf weight, leaf elongation rate, tillers plant⁻¹, mass tiller⁻¹, and total nonstructural carbohydrate concentration than soil water. Irrigation might have had greater impact on herbage productivity if more water had been applied or if canopy cover had been less extreme. Silvicultural practices should be imposed to improve penetration of solar irradiance to the alley crop.     

Growth and yield of maize alley cropped with Leucaena leucocephala and Faidherbia albida in Morogoro, Tanzania

This study examined the effect of alley cropping of Leucaena leucocephala and Faidherbia albida on wood biomass, maize grain yield and soil nitrogen status. The treatments were: trees planted alone at 1 × 5 m spacing; trees intercropped with maize and a sole maize crop. Mulch biomass averaged 6.18 and 0.97 t ha⁻¹ for L. leucocephala and F. albida, respectively. Corresponding wood production was 1.71 and 1.11 t ha⁻¹. Both total N and inorganic N (NO ₃ ⁻ –N plus ₄ ⁺ –N) were higher under F. albida and lowest under L. leucocephala. Similarly, foliar N concentration in maize was higher in plots intercropped with F. albida and least in L. leucocephala intercropping. Maize grain yield was little affected by the tree intercrop as competition for resources was reduced through periodic pruning and clean weeding. There was no gain in maize grain yield due to the presence of L. leucocephala and F. albida. These results suggest that alley cropping in Gario is justified for wood production but not for increasing maize grain yield. This revised version was published online in June 2006 with corrections to the Cover Date.     

Biomass production, foliar and root characteristics and nutrient accumulation in young silver birch (Betula pendula Roth.) stand growing on abandoned agricultural land

The above-ground biomass and production, below-ground biomass, nutrient (NPK) accumulation, fine roots and foliar characteristics of a 8-year-old silver birch (Betula pendula) natural stand, growing on abandoned agricultural land in Estonia, were investigated. Total above-ground biomass and current annual production after eight growing seasons was 31.2 and 11.9 t DM ha⁻¹, respectively. The production of stems accounted for 62.4% and below-ground biomass accounted for 19.2% of the total biomass of the stand. Carbon sequestration in tree biomass reaches roughly 17.5 t C ha⁻¹ during the first 8 years. The biomass of the fine roots (d < 2 mm) was 1.7 ± 0.2 t DM ha⁻¹ and 76.2% of it was located in the 20 cm topsoil layer. The leaf area index (LAI) of the birch stand was estimated as 3.7 m² m⁻² and specific leaf area (SLA) 15.0 ± 0.1 m² kg⁻¹. The impact of the crown layer on SLA was significant as the leaves are markedly thicker in the upper part of the crown compared with the lower part. The short-root specific area (SRA) in the 30 cm topsoil was 182.9 ± 9.5 m² kg⁻¹, specific root length (SRL), root tissue density (RTD) and the number of short-root tips (>95% ectomycorrhizal) per dry mass unit of short roots were 145.3 ± 8.6 m g⁻¹, 58.6 ± 3.0 kg m⁻³ and 103.7 ± 5.5 tips mg⁻¹, respectively. In August the amount of nitrogen, phosphorus and potassium, accumulated in above ground biomass, was 192.6, 25.0 and 56.6 kg ha⁻¹, respectively. The annual flux of N and P retranslocation from the leaves to the other tree parts was 57.2 and 3.7 kg ha⁻¹ yr⁻¹ (55 and 27%), respectively, of which 29.1 kg ha⁻¹ N and 2.8 kg ha⁻¹ P were accumulated in the above-ground part of the stand.     

Intercropping field crops between rows of Leucaena leucocephala under rainfed conditions in northern India

A field study was conducted for six years (1981–1986) on sandy loam soil on intercropping hedgerows of Leucaena leucocephala (Lam) de Wit. with three field crops viz. maize (Zea mays L), black gram (Vigna mungo L) and cluster bean (Cyamopsis tetragonoloba L Taub.). In treatments 1 and 2 Leucaena hedges were planted as pure crops at close (25 cm × 75 cm) and wide (25 cm × 375 cm) spacings. In treatments 3, 4 and 5 the three field crops were intercropped between the hedgerows of Leucaena at the wide spacing, and in treatments 6, 7 and 8 the field crops were raised as pure crops. Leucaena was topped to 75 cm each time it attained a height of 175 cm. The pure crop of Leucaena at close spacing produced an average, over the six years, of 34 t ha⁻¹a⁻¹ of green fodder and 9.4 t ha⁻¹a⁻¹ of air dry fuelwood. The Leucaena at wide spacing produced 18.9 t ha⁻¹a⁻¹ of green fodder and 6.3 t ha⁻¹a⁻¹ of fuelwood. Intercropping with field crops decreased the yield of green fodder and fuelwood. The yield of all the field crops was less when raised as intercrops than as pure crops. Mean maximum net returns were obtained from intercrops of Leucaena and cluster bean (Rs 3540 ha⁻¹a⁻¹) which were significantly higher than the returns from pure crop of Leucaena at wide spacing but similar to the returns from pure crops of cluster bean. Leucaena with maize (Rs 3273 ha⁻¹a⁻¹) and black gram (Rs 3125 ha⁻¹a⁻¹) gave significantly higher net returns over pure crops of Leucaena at wide spacing, maize and black gram. ICRISAT = International Crops Research Institute for the Semi-Arid Tropics- Hyderabad, India. CIAT = Centro International de Agricultura Tropical - Cali - Columbia     

Above- and belowground biomass,nutrient and carbon stocks contrasting an open-grown and a shaded coffee plantation

Coffee (Coffea canephora var robusta) is grown in Southwestern Togo under shade of native Albizia adianthifolia as a low input cropping system. However, there is no information on carbon and nutrient cycling in these shaded coffee systems. Hence, a study was conducted in a mature coffee plantation in Southwestern Togo to determine carbon and nutrient stocks in shaded versus open-grown coffee systems. Biomass of Albizia trees was predicted by allometry, whereas biomass of coffee bushes was estimated through destructive sampling. Above- and belowground biomass estimates were respectively, 140 Mg ha⁻¹ and 32 Mg ha⁻¹ in the coffee–Albizia association, and 29.7 Mg ha⁻¹ and 18.7 Mg ha⁻¹ in the open-grown system. Albizia trees contributed 87% of total aboveground biomass and 55% of total root biomass in the shaded coffee system. Individual coffee bushes consistently had higher biomass in the open-grown than in the shaded coffee system. Total C stock was 81 Mg ha⁻¹ in the shaded coffee system and only 22.9 Mg ha⁻¹ for coffee grown in the open. Apart from P and Mg, considerable amounts of major nutrients were stored in the shade tree biomass in non-easily recyclable fractions. Plant tissues in the shaded coffee system had higher N concentration, suggesting possible N fixation. Given the potential for competition between the shade trees and coffee for nutrients, particularly in low soil fertility conditions, it is suggested that the shade trees be periodically pruned in order to increase organic matter addition and nutrient return to the soil. An erratum to this article can be found at     

Alley cropping on an Ultisol in subhumid Benin. Part 1: Long-term effect on maize, cassava and tree productivity

In southern Benin, West Africa, two alley cropping systems were studied from 1986 to 1992. Yield development was followed in a maize and cassava crop rotation vs. intercropping system, with alleys of Leucaena leucocephala (Lam.) de Wit and Cajanus cajan (L.) Millsp. vs. a no-tree control, with and without NPK fertiliser. Without alleys, NPK fertilisation maintained high yield levels of 2–3 t maize dry grain plus 4–6 t ha^–1 cassava root DM in intercropping, 3–4 t ha^–1 maize and 6–10 t ha^–1 cassava in solercropping. Without NPK, final yields seemed to stabilise at about 1 t maize plus 2 t cassava in intercropping and twice as much in each solecrop. Alley cropping induced significant yield increases by about 50% with both tree species in unfertilised, intercropped maize, and with Cajanus in fertilised, solecropped cassava. In monetary terms, the NPK-fertiliser response of stabilised yields was significant for all treatments except the solecropped Leucaena alleys. It is concluded that on Ultisols with low nutrient status in the upper rooting zone, alley cropping with low-competitive tree species may improve food crop yields but the greatest monetary output is achieved by intercropping with mineral fertiliser independent of the presence or absence of an agroforestry component.     

Fine root dynamics in three central Himalayan high elevation forests ranging from closed canopied to open-canopied treeline vegetation

Fine root biomass, rates of dry matter production and nutrients dynamics were estimated for 1 year in three high elevation forests of the Indian central Himalaya. Fine root biomass and productivity were higher in closed canopied cappadocian maple forest (9.92 Mg ha⁻¹ and 6.34 Mg ha⁻¹ year⁻¹, respectively), followed by Himalayan birch forest (6.35 Mg ha⁻¹ and 4.44 Mg ha⁻¹ year⁻¹) and Bell rhododendron forest (6.23 Mg ha⁻¹ and 2.94 Mg ha⁻¹ year⁻¹). Both fine root biomass and productivity declined with an increase in elevation. Across the sites, fine root biomass was maximal in fall and minimal in summer. In all sites, maximum nutrient concentration in fine roots was in the rainy season and minimum in winter. Fine root biomass per unit basal area was positively related with elevation, Bell rhododendron forest having the largest fine root biomass per unit of basal area (0.53 Mg m⁻²) and cappadocian maple the least (0.18 Mg m⁻²). The production efficiency of fine roots per unit of leaf biomass also increased with elevation and ranged from 1.13 g g⁻¹ leaf mass year⁻¹ in cappadocian maple forest to 1.28 g g⁻¹ leaf mass year⁻¹ in Bell rhododendron forest. Present fine root turnover estimates showed a decline towards higher elevations (0.72 year⁻¹ in cappadocian maple and 0.58 year⁻¹ in Bell rhododendron forest) and are higher than global estimates (0.52).     

Crop residue effect on crop performance,soil N<Subscript>2</Subscript>O and CO<Subscript>2</Subscript> emissions in alley cropping systems in subtropical China

Land management practices that simultaneously improve soil properties are crucial to high crop production and minimize detrimental impact on the environment. We examined the effects of crop residues on crop performance, the fluxes of soil N₂O and CO₂ under wheat-maize (WM) and/or faba bean-maize (FM) rotations in Amorpha fruticosa (A) and Vetiveria zizanioides (V) intercropping systems on a loamy clay soil, in subtropical China. Crop performance, soil N₂O and CO₂ as well as some potential factors such as soil water content, soil carbon, soil nitrogen, microbial biomass and N mineralization were recorded during 2006 maize crop cultivation. Soil N₂O and CO₂ fluxes are determined using a closed-based chamber. Maize yield was greater after faba bean than after wheat may be due to differences in supply of N from residues. The presence of hedgerow significantly improved maize grain yields. N₂O emissions from soils with maize were considerably greater after faba bean (345 g N₂O–N ha⁻¹) than after wheat (289 g N₂O–N ha⁻¹). However, the cumulated N₂O emissions did not differ significantly between WM and FM. The difference in N₂O emissions between WM and FM was mostly due to the amounts of crop residues. Hedgerow alley cropping tended to emit more N₂O than WM and FM, in particular A. fruticosa intercropping systems. Over the entire 118 days of measurement, the N₂O fluxes represented 534 g N₂O–N ha⁻¹ (AWM) and 512 g N₂O–N ha⁻¹ (AFM) under A. fruticosa species, 403 g N₂O–N ha⁻¹ (VWM) and 423 g N₂O–N ha⁻¹ (VFM) under Vetiver grass. We observed significantly higher CO₂ emission in AFM (5,335 kg CO₂–C ha⁻¹) from June to October, whereas no significant difference was observed among WM (3,480 kg CO₂–C ha⁻¹), FM (3,302 kg CO₂–C ha⁻¹), AWM (3,877 kg CO₂–C ha⁻¹), VWM (3,124 kg CO₂–C ha⁻¹) and VFM (3,309 kg CO₂–C ha⁻¹), indicating the importance of A. fruticosa along with faba bean residue on CO₂ fluxes. As a result, crop residues and land conversion from agricultural to agroforestry can, in turn, influence microbial biomass, N mineralization, soil C and N content, which can further alter the magnitude of crop growth, soil N₂O and CO₂ emissions in the present environmental conditions.     

Alley cropping of maize with calliandra and leucaena in the subhumid highlands of Kenya: Part 2. Biomass decomposition, N mineralization, and N uptake by maize

A major challenge in developing agroforestry approaches that utilize tree-leaf biomass for provision of N to crops is to ensure synchrony between the N released from decomposing prunings and N demand by crops. A study was conducted in the subhumid highlands of Kenya to assess the rate of decomposition and mineralization of soil-incorporated Calliandra calothyrsus Meissner (calliandra) and Leucaena leucocephala (Lam.) de Wit (leucaena) tree biomass and maize roots (Zea mays L.) both in an alley cropping and a sole cropping system. The amount of mineralized N peaked four weeks after planting (WAP) maize in all the treatments during both seasons of 1995. Cumulative mineralized N at week 20 ranged from 114 to 364 kg N ha⁻¹ season⁻¹, the absolute control treatment giving the lowest and the prunings-incorporated treatments giving the highest amounts in the two seasons. Total N uptake by maize, ranging from 42 to 157 kg ha⁻¹ season⁻¹, was lowest in the 'alley-cropped, prunings-removed' treatments, and highest in the 'non alley-cropped-prunings-incorporated' treatments. The apparent N recovery rate by maize was highest in the fertilizer applied treatments in the two seasons. Decomposition rate constants (kD) ranged from 0.07 to 0.21 week⁻¹, and the rates among the different plant residues were as follows: leucaena < calliandra < maize roots. Nitrogen release rate constants (kN), ranging from 0.04 to 0.25 week⁻¹, followed a similar pattern as the rate of decomposition with leucaena releasing the highest amount of N followed by calliandra and lastly by maize roots. This revised version was published online in June 2006 with corrections to the Cover Date.     

Leucaena + maize alley cropping in Malawi. Part 2: Residual P and leaf management effects on maize nutrition and soil properties

Agroforestry systems involving leaf removal for animal fodder may result in rapid depletion of soil fertility. The purpose of this research was to determine if the effects of leaf removal on soil fertility parameters and maize yield in a Leucaena leucocephala alley cropping system could be reversed. Three leaf management strategies in a Leucaena alley cropping trial that had been in effect from 1987 to 1991 were investigated: 1) leaves returned, 2) leaves removed, and 3) leaves removed, with 100 kg inorganic N ha⁻¹ added. In the 1990/91 season, a 3⁴ confounded factorial design was utilized to investigate the effects of leaf management strategy, N rate (0, 30 and 60 kg N ha⁻¹); maize plant population (14,800, 29,600, and 44,400 plants ha⁻¹); and P rate (0, 18, and 35 kg P ha⁻¹). In the 1991/92 and 1992/93 seasons, leaves were applied equally to all plots, and no P was applied. The N rate and plant population treatments were continued, and the same confounded factorial design was implemented to investigate residual leaf management strategy, residual P rate, n rate, and plant population. The yield gap between the plots where leaves had been returned vs. removed narrowed each season due to uniform leaf application. Application of N improved yields during both seasons. Residual effects of the initial P application decreased to only 10% of the total yield in 1992/93. Plant population affected yields only during the season of very good rainfall. Leaf additions resulted in a relative increase in soil pH, total N,and exchangeable Ca, Mg, and K and a decrease C/N ratio in plots that had not previously received leaves. Leaves supplied more K and Zn to the upper 15 cm of soil than were being extracted by the maize crop, but uniform leaf additions eliminated differences in K and Zn uptake. Plant Zn uptake decreased with increasing P rate and plant population, and increased with increasing N rate and a history of leaf return. The results show that applying leaves equilibrated yields within two seasons, and resulted in a relative improvement of several soil properties. The residual effect from P applications was not adequate to maximize yields.     

Morphological plasticity of cotton roots in response to interspecific competition with pecan in an alleycropping system in the southern United States

A study was conducted in northwest Florida, USA, to investigate root development and morphology of cotton (Gossypium hirsutum L.) under pecan (Carya illinoensis K. Koch) trees in an alleycropping experiment. Root:shoot ratio, root biomass, total root length and root length density were examined under three treatments: (1) barrier (separating belowground interspecific competition by trenching to a depth of 120 cm and installing polyethylene barrier), (2) non-barrier (root systems were free to interact), and (3) monoculture of cotton (without above and belowground interspecific competition with trees). Results indicated that plants in the barrier and non-barrier treatments had lower root:shoot ratios compared to the monoculture treatment. Belowground competition for resources between pecan and cotton in the non-barrier treatment resulted in 25 and 33% reduction of total root length (359 cm) when compared to that of the barrier (477 cm) and monoculture (539 cm) treatments, respectively. The non-barrier plants also exhibited the lowest root length density. Specific root length was highest for the monoculture (179 cm g⁻¹) and lowest for the non-barrier treatment (146 cm g⁻¹) with the barrier treatment being intermediate (165 cm g⁻¹). Interspecific competition with pecan significantly altered root development and morphology of cotton plants. Research in agroforestry should take into account the developmental differences in root systems of the associated crop species so that better models incorporating nutrient and water uptake can be developed.     

Volunteer biomass production between multipurpose tree hedgerows after two years of fallow in southern Cameroon

The ability of multipurpose hedgerow tree species to out-compete undesired regrowth during fallow phases was examined. Biomass and spatial distribution of grass and broad leaf volunteers was measured after two years of fallow, in two alley cropping systems planted at six m interrow distance, at the Humid Forest Ecoregional Centre Research Station, Mbalmayo, southern Cameroon. The two experiments had been continuously cropped for five and six years previously. In the experiment cropped for six years, the presence of Senna spectabilis [(DC.) Irwin and Barnaby] hedgerows reduced the biomass of the volunteer regrowth from 9.2 to 4.3 Mg ha⁻¹. Tillage during the previous cropping phase increased the broad leaf biomass from 3.0 to 4.4 Mg ha⁻¹, reduced the biomass of grasses from 3.4 to 2.7 Mg ha⁻¹ but had no effect on the total volunteer biomass. Volunteer biomass was significantly lower within 1.5 m of the S. spectabilis hedgerows than at positions further away. In the experiment cropped for five years, S. spectabilis reduced the volunteer regrowth biomass significantly. Two other hedgerow species, Dactyladenia barteri [(Hook f ex Oliv.) Engl.] and Flemingia macrophylla [(Willd.) Merrill] had no effect on the total amount of volunteer regrowth but did reduce volunteer biomass within 0.5 m of the hedgerows. S. spectabilis caused a stronger reduction of volunteer biomass than D. barteri and F. macrophylla at almost all distances from the hedgerows. The competitiveness of D. barteri and F. macrophylla is insufficient and their growth habit is unsuitable to out-compete undesired species in this alley cropping system. This revised version was published online in June 2006 with corrections to the Cover Date.     

Reduced soil nutrient leaching following the establishment of tree-based intercropping systems in eastern Canada

Tree-based intercropping (TBI) systems, combining agricultural alley crops with rows of hardwood trees, are largely absent in Canada. We tested the hypothesis that the roots of 5–8 years old hybrid poplars, growing in two TBI systems in southern Québec, would play a “safety-net” role of capturing nutrients leaching below the rooting zone of alley crops. TBI research plots at each site were trenched to a depth of 1 m on each side of an alley. Control plots were left with tree roots intact. In each treatment at each site, leachate at 70 cm soil depth was repeatedly sampled over two growing seasons using porous cup tension lysimeters, and analyzed for nutrient concentrations. Daily water percolation rates were estimated with the forest hydrology model ForHyM. Average nutrient concentrations for all days between consecutive sampling dates were multiplied by water percolation rates, yielding daily nutrient leaching loss estimates for each sampling step. We estimated that tree roots in the TBI system established on clay loam soil decreased subsoil NO₃ ⁻ leaching by 227 kg N ha⁻¹ and 30 kg N ha⁻¹ over two consecutive years, and decreased dissolved organic N (DON) leaching by 156 kg N ha⁻¹ year⁻¹ in the second year of the study. NH₄ ⁺ leaching losses at the same site were higher when roots were present, but were 1–2 orders of magnitude lower than NO₃ ⁻ or DON leaching. At the sandy textured site, the safety net role of poplar roots with respect to N leaching was not as effective, perhaps because N leaching rates exceeded root N uptake by a wider margin than at the clay loam site. At the sandy textured site, significant and substantial reductions of sodium leaching were observed where tree roots were present. At both sites, tree roots reduced DON concentrations and the ratio of DON to inorganic N, perhaps by promoting microbial acquisition of DON through rhizodeposition. This study demonstrated a potential safety-net role by poplar roots in 5–8 year-old TBI systems in cold temperate regions.     

Nitrogen and fine root length dynamics in a tropical agroforestry system with periodically pruned Erythrina poeppigiana

The effect of pruning all branches (complete pruning) or retaining one branch (partial pruning) on the dynamics of nitrogen cycling in aboveground biomass, nitrogen supplying power of an amended Eutric Cambisol, and fine root length, was studied in an Erythrina poeppigiana (Walp.) O.F. Cook—tomato (Lycopersicon esculentum Mill.) alley cropping practice in Turrialba, Costa Rica during 1999–2000. Over the 1 year pruning cycle, in which trees were completely or partially pruned four times, respective aboveground biomass production was 4.4 Mg or 7 Mg ha⁻¹ (2-year-old trees) and 5.5 Mg or 9 Mg ha⁻¹ (8-year-old trees); N cycled in aboveground biomass was 123 kg or 187 kg ha⁻¹ (2-year-old trees) and 160 kg or 256 kg N ha⁻¹ (8-year-old trees); mean fine root length was 489 or 821 m (2-year-old-trees), 184 or 364 m per tree (8-year-old-trees). Pruning intensity did not significantly affect net N mineralisation and net nitrification rates during the tomato-cropping season. For the tomato crop, pre-plant mean net N mineralisation rate of 2.5 mg N kg⁻¹ soil day⁻¹ was significantly lower than 16.7 or 11.6 mg N kg⁻¹ soil day⁻¹ at the end of vegetative development and flowering, respectively. Mean net nitrification rates of 3.5, and 4.3 mg N kg⁻¹ soil day⁻¹, at pre-plant and end of vegetative development, respectively, were significantly higher than 0.3 mg N kg⁻¹ soil day⁻¹ at end of flowering. In humid tropical low-input agroforestry practices that depend on organic inputs from trees for crop nutrition, retention of a branch on the pruned tree stump appears to be a good alternative to removal of all branches for reducing N losses through higher N cycling in aboveground biomass, and for conserving fine root length for higher N uptake, although it might enhance competition for associated crops.     

Production and phenology of the fruit shrub Eugenia stipitata in agroforestry systems in Costa Rica

Arazá (Eugenia stipitata), a fruit shrub originating from Western Amazonia, was evaluated growing in association with timber shade trees (Acacia mangium or Cordia alliodora) or with plantain (Musa sp.) as a potential commercial species for the tropical moist lowlands of Talamanca, Costa Rica. Height and crown width of the four-year-old shrubs varied between 2.7–2.8 m and 2.9–3.1 m, respectively. Flowering was positively correlated with initial fruit formation 1 month later and initial fruit formation with fully developed fruits a subsequent month after that. Three to four-year-old plants produced 20.0, 20.0, and 24.5 t ha⁻¹ yr⁻¹ (fresh fruits) in associations with A. mangium, C. alliodora and plantain, respectively, with higher production in rainy months. Fruit production in later years under C. alliodora (six to eight-year-old shrubs) was 26.5 t ha⁻¹ yr⁻¹. Where markets exist for E. stipitata fruit, the association can be recommended for tropical humid lowlands of Central America.     

The effects of between-row (alley widths) and within-row spacings ofGliricidia sepium on alley-cropped maize in Sierra Leone

The effects of spacings between hedgerows (alley widths) and the spacings of trees within hedgerows ofGliricidia sepium on growth and grain yield of maize were investigated at Senehun in southern Sierra Leone. Four between-row spacings (2, 4, 6 and 8 m) were combined with three within-row spacings (0.25, 0.50 and 1.00 m) in a split block design. Maize, at densities of 20,000, 40,000 and 53,333 plants ha^–1, was established in the alleys and also as pure crops. N, P and K fertilizers were applied to all plots before pruning of the trees began. When pruning started, only the pure maize plots received fertilizer; prunings from the hedgerows were returned to the appropriate alleys in the other plots.Plots with the highest maize populations consistently gave the best yields before pruning started, but lower populations gave improved yields after pruning. Yields of maize increased with increasing alley widths before the start of pruning, after which the narrower alleys of 2 and 4 m outyielded the wider ones by almost double, probably because of the large amount of nutrients applied in prunings. Lack of light limited grain yields before the start of pruning, when there was some shading by the hedgerows. Alleys of 2–4 m wide, planted no closer than 0.50 m within rows, resulted in more than twice the yields of maize than in the 8-m alleys planted at 0.25 m within rows, once the hedgerows were well established and were being managed.