THE EFFECT OF TIME OF CUTTING ON EAR PRODUCTION AND SEED YIELD IN S.48 TIMOTHY

After uniform defoliation in February, spaced plants of S.48 timothy were cut on one of 12 fortnightly dates, starting at the end of March.
Analysis of the herbage removed at each cut showed an increase in plant and unit tiller weight throughout the season, but a decline in the number of tillers, especially before the beginning of shooting.
The date of ear emergence was not affected by cutting up to 5 May and only slightly delayed in plants cut on 19 May. Plants treated subsequently flowered much later but after a fairly constant interval following defoliation.
Cutting on and after 7 April depressed fertile tiller formation with increasing severity, until only vegetative tillers appeared in the recovery growth following treatment at the beginning of August.
Seed yield per plant declined with the decrease in fertile tiller numbers, but after the early cuts some compensation by production of slightly more seed per ear was recorded.
The results are discussed in terms of the physiological transition of the plant towards flowering.     

THE EFFECT OF DEFOLIATION ON THE SEED-PRODUCING CAPACITY OF BRED VARIETIES OF GRASSES

Leafy grass varieties S24 perennial ryegrass, S26 cocksfoot, S215 meadow fescue and S51 timothy were cut in October and at various growth stages in spring; S53 meadow fescue was grazed at similar periods. Sub-plots were given different applications of N. The control plots had similar manurial treatments but were not defoliated. The seed yield of S24 following October cutting was significantly lower than the control in the first harvest year and significantly higher in the second; the mean yields over 2 years, as a result of cutting at this stage and before ear formation in March, were very similar to those of the control treatment. The mean yields from 3 harvest years for S26, S215 and S51 were generally improved by cutting in October and before ear formation, but cutting after ear formation reduced the number of heads and the seed yields. Extra N applied in conjunction with the defoliation had no significant effect on the number of seed heads or on seed yield. Double grazing of S53 in Sept. and Dec. reduced significantly its seed production in both harvest years. Application of N in spring gave a greater response than a similar application in autumn.     

温光敏核不育小麦C338S的育性表现及不同蘖位和花位的育性差异

为了解温光敏核不育小麦不同播期及不同蘖位、花位的育性变化,通过多年分期播种试验,研究了小麦温光敏细胞核雄性不育系C338S不同播期下的育性表现及不同蘖位和花位的育性差异。结果表明,随着播期的调整,C338S分别表现完全不育、高度不育、半不育、低不育、可育的不同育性状态,具有育性转换特性。主茎穗与分蘖穗之间、主茎穗不同部位的小穗之间以及不同位次的小花之间的育性存在差异。发育较晚的后生分蘖的育性高于主茎穗和早生分蘖;主茎穗的下部小穗育性>中部小穗>上部小穗;早播不育植株发育较晚的3、4位小花的育性较好,而晚播可育植株发育较早的1、2位小花的育性较高。     

THE EFFECT OF CUTTING COCKSFOOT (DACTYLIS GLOMERATA L.) GROWN FOR PRODUCTION OF SEED

An experiment is described in which cocksfoot, grown for production of seed, was cut annually for 3 years in either autumn, winter or spring. No significant increases in yield of seed/ac were obtained from any treatment compared with an uncut control, and a cut in late April in one year seriously reduced the yield of seed. Additional N applied after a defoliation in October did not increase the yield of seed compared with that from plots which were cut at the same time but not fertilized. The production of tillers was not affected by cutting on any date. The percentage fertility of tillers was increased by cutting in the first year, when the number of tillers was smallest; the fertility was less consistently affected in the second year, and was generally unaffected in the third year. The weight of seed produced per ear was usually reduced by the cutting treatments. In the third year of the experiment it was shown that the penetration of light to the base of the shoots was increased substantially by defoliation, but it is deduced that maximum interception of light by the whole plant is more important for the production of seed than increasing by cutting that proportion which penetrates to the base of the plant.     

THE EFFECT OF CUTTING TIMOTHY (PHLEUM PRATENSE L.) GROWN FOR PRODUCTION OF SEED

An experiment is described in which timothy, grown for production of seed, was cut once annually on dates ranging from 23 October to 24 May. New tillers in selected plants were labelled at monthly intervals during autumn and winter for 2 years, to determine the pattern of production of tillers and to assess the reproductive development of categories of tillers. The majority of tillers were produced in autumn, but production was continuous throughout the winter. There was a big increase from the first to the second year in the total number of tillers produced, but neither the pattern nor amount of tiller production was affected by cutting. The % of fertile tillers was highest in the oldest tillers, and was affected little by the increase in the number of tillers from the first to the second year; cutting had little effect on the % fertility except where floral primordia were removed by the cut on 24 May. In all treatments at least 90% of the ears at harvest were produced by tillers initiated the previous autumn. The penetration of light to the base of the plants was increased considerably by cutting, but it is concluded that interception of light by the whole plant is more beneficial than an increase in the penetration of light into the plant and a concomitant reduction in the total interception of light by the plant.     

FERTILE TILLER PRODUCTION AND SEED YIELD IN MEADOW FESCUE (FESTUCA PRATENSIS L.)

The fertile tiller production and seed yields of spaced plants of meadow fescue varieties were found to vary widely with the time of sowing, particularly in the first harvest year. The benefit of early spring sowing is indicated in the total productivity over two harvest years.
Varieties differed in the amount of heading in the seeding year; the American type was the most prolific. This had a bearing on the relative yields in the first harvest year, since many of the potentially productive tillers of this variety were wasted through the plants becoming ripe to flower in the seeding year and, therefore, the rate of tiller production was checked. Competition within the plant appeared to be the cause of tillers remaining vegetative when conditions otherwise seemed suitable for generative development. The smaller-sized plants gave a higher proportional increase of fertile tillers at harvest than would be expected from the vegetative tillers counted in April, while at the same time there was a negative relationship at harvest from plants which had a very high number of vegetative tillers.
Infertility as reflected in a poor seed set at the time of harvest is a serious wastage of potential production in meadow fescue and, in selecting for superior seed output, breeders need to take this factor into consideration.
The 1000-seed weight varied between varieties, the pasture type being inferior to the two hay types.     

Spring defoliation of white clover seed crops. 2. Potential harvestable seed yield and seed yield components of contrasting white clover cultivars

The effects of different spring defoliation managements on potential harvestable seed yield and seed yield components of three contrasting white clover cultivars were assessed. The small-leaved cv. S184 produced more but smaller inflorescences than the large-leaved cv. Olwen and Menna, a medium-leaved cultivar. Cultivar Olwen, however, produced more ripe and brown (nearly ripe) inflorescences with more florets, seeds per floret and a higher seed yield per ten inflorescences than the other cultivars. Potential harvestable seed yield and individual seed yield components were only influenced by defoliation after bud emergence, as defoliation before bud emergence had no effect on seed yield components. Defoliation after bud emergence had a similar effect on all cultivars: the number of ripe inflorescences was unaffected by defoliation but the number of brown and therefore harvestable (ripe + brown) inflorescences was highest following defoliation three weeks after bud emergence. Florets per inflorescence, seed per floret, 1000 seed weight, seed yield per ten inflorescences and potential harvestable seed yield were not influenced by defoliation after bud emergence. Season had a significant effect on seed yield components and influenced the effect of defoliation treatments, emphasizing the importance of climate in white clover seed production. The results are discussed in relation to the spring defoliation of white clover seed crops, harvesting techniques and the provision of guidelines for optimizing seed yield.     

THE FREQUENCY OF DEFOLIATION OF INDIVIDUAL TILLERS IN A SET-STOCKED SWARD

A method designed to study the defoliation of individual tillers in a set-stocked sward is described. This was used on S23 ryegrass swards carrying 19 (medium-stocked) and 30 (heavily-stocked) sheep/ac.
The tillers in the heavily-stocked sward were defoliated, on average, every 7–8 days, and those in the medium-stocked sward every 11–14 days. The total green leaf length (GLL) of tillers grazed by the sheep was generally greater than that of tillers not grazed. On average 27 and 40% of the GLL of tillers in the medium- and heavily-stocked swards respectively was removed at each grazing. Older leaves on the grazed tillers were defoliated much less frequently than were younger leaves.
It is suggested that a rational analysis of the relationship between the grazing animal and the grazed sward cannot be made without more work of this nature.     

Nitrogen studies in Lolium perenne grown for seed III. The effect of nitrogen and water stress

The effects of water and nitrogen (N) availabilities and their possible interactions on the seed yield and yield components of S23 perennial ryegrass grown in pots in the glasshouse were investigated. Increasing the N fertilizer from 0 to 5 g per pot more than doubled the seed yield but no further increase in seed yield was found by increasing N fertilizer from 5 to 10 g per pot. The main effect of increasing water availability was to increase both seed and straw yield. When N had been applied, accumulated evapotranspiration was linearly related to seed and straw yield. The component of yield having the greatest effect on yield was the number of fertile tillers and, although both N and water availability affected this component, N had the larger effect. Implications of these responses are discussed in relation to the potential yield of the crop and to the field situation.     

Pattern of interspecific tiller defoliation in a mixed-grass prairie grazed by cattle

Permanently marked tillers of five perennial grasses, native to the mixed-grass prairie of North America, were monitored to determine patterns of defoliation, architectural attributes influencing probabilities of defoliation. and post-defoliation responses. Frequency of tiller defoliation was greatest for one of the dominant midgrasses, intermediate for the remaining midgrass and two dominant shortgrasses. and least for the subdominant midgrass. Midgrass species, including the infrequently grazed subordinate species, were consistently grazed more intensively than the shortgrass species. However, the relative intensity of defoliation did not vary among species or grazing periods indicating that intensity of defoliation was primarily a function of pre-defoliation tiller height. Tiller architecture, including height, lamina number and the presence of reproductive culms, did not significantly influence frequency or intensity of defoliation within a species. The lack of evidence supporting tiller architecture as a selection criterion within a species suggests that animals were selecting on vegetation parameters at higher levels of vegetation organization than individual tillers. Grazed tillers of the three midgrasses exhibited greater relative rates of tiller elongation between grazing periods than non-grazed tillers, particularly early in the season when environmental conditions for growth were most favourable. The patterns of interspecific tiller defoliation observed in this study parallel the long-term patterns of grazing-induced species replacement observed in this grassland.     

SEED PRODUCTION STUDIES ON PERENNIAL RYEGRASS, TIMOTHY AND PRAIRIE GRASS

A study over two years involving tiller identifica tion according to month of origin enabled the contribution of individual tillers of perennial rye grass Grasslands Ruanui, timothy Grasslands Kathu and a local New Zealand strain of prairie grass (Bromus unioloides) to be determined at seed harvest. In addition, the distribution of seed-heads in different arbitrary emergence groups was detemined so as to allow analysis of individual seedhead components within each group. In all species, tillers formed during the first 4 months following an autumn sowing in the first year, and also in the immediate post-harvest period and through the autumn in the second year, made a major contribution to seedhead numbers and seed yield at harvest. These tillers had a lower vegetative mortality rate than those formed in the spring. Primary tillers formed after sowing were highly persistent and became almost exclusively repro ductive. Many vegetative tillers died before har vest, behaving as annual tillers. Spring-formed tillers made a major contribution to the vegetative growth of the plant over the summer and early autumn. These tillers generally died before the following winter. Tillers formed in winter and early spring by ryegrass and prairie grass showed a high mortality, particularly in the second year. In timothy, mortality was highest amongst tillers formed in late spring. The reduced contribution of spring-formed tillers to the seed head population occurred as a result of fewer heads numerically compared with older tillers. In all species, most seedhead components varied according to time of ear emergence. Early emerged heads were generally longer, bore more spikelets and more florets per head and had a greater culm length than late-emerged heads. The major trend in all species was a general depression in head length, spikelet and floret number and culm length with grazing. These effects were in most cases partly or completely overcome by N application. The major effect of applied N was to increase floret number per head.     

EAR-BEARING CAPACITY OF TILLERS ARISING AT DIFFERENT TIMES IN HERBAGE GRASSES GROWN FOR SEED

Three experiments are described, two with S215 meadow fescue and one with S37 cocksfoot, all grown for seed, during which all new tillers, soon after their appearance, were labelled at intervals of a month—a total of nearly 12,000 tillers. Whatever the first date of observation, the earliest tillers were always found to have contributed the largest proportion of the ears present at harvest. In one experiment applied nitrogen caused an increase in the number of tillers and ears, without however affecting the percentage contribution of the earliest tillers to the final crop of ears. The importance of the autumn period of tillering is stressed.     

THE EFFECT OF DEFOLIATION ON THE SEED-PRODUCING CAPACITY OF BRED STRAINS OF GRASSES

Two leafy strains of cocksfoot, S143 and S37 were subjected to winter, spring and winter + spring defoliation; 2 cwt. per acre of nitro-chalk was applied after each defoliation, in addition to the standard manuring for all plots.
Winter grazing of leafy cocksfoot does not appear to depress the yield of seed appreciably when extra nitrogen is applied.
Grazing of cocksfoot in mid-April is likely to depress the seed yield substantially, especially if followed by a very dry summer.
The reaction to hard defoliation by sheep may vary with the growth form of the cocksfoot strain.
Grazing appeared to be more detrimental to seed yield than defoliation by cutting.
During this investigation, prolonged frost which occurred during the early part of the year appeared to have an adverse effect on cocksfoot seed yield.     

FERTILE TILLER PRODUCTION AND SEED YIELD IN MEADOW FESCUE (FESTUCA PRATENSIS L.)

The seed-producing capacity of two varieties of meadow fescue, S215 and S53, sown at three different spacings, was compared. One level of nitrogen was applied on three different dates in the spring. Tiller counts were made periodically during the growth of the crop. Fertile-tiller production was found to be related to vegetative growth.
The varieties differed not only in yield, but also in growth dynamics. A larger percentage of overwintering shoots became fertile in the hay variety, than in the pasture variety.
Drill spacings had significant effects on plant establishment and on vegetative-and fertile-tiller numbers. In both years N applied in March gave the highest production of fertile tillers. Date of N application did not produce marked trends in yield. The influence of spacing and date of N application on the number and density of spikelets indicated that the plant adjusted itself to environmental factors. Thus spikelet numbers were reduced in early heads with narrower drill spacing and with late applications of N. Head emergence was earlier in narrow drills, but time of N application in the spring did not produce significant differences in this respect.     

THE EFFECT OF DEFOLIATION ON THE SEED-PRODUCING CAPACITY OF BRED STRAINS OF GRASSES

A leafy strain of timothy and of perennial ryegrass showed no significant reductions in seed yields following grazing at different dates from October to January, when extra nitrogen was applied.
Spring and winter + spring grazing, together with extra nitrogen, significantly reduced the seed yield of S50 timothy for three successive harvest years.
Perennial ryegrass S101 showed no reduction after spring or winter + spring grazing when defoliation was made not later than mid-April, but when grazing was delayed until May and drought occurred there was a significant reduction in seed yield.     

Nitrogen studies in Lolium perenne grown for seed. II. Timing of nitrogen application

The effects of applying nitrogen at different growth stages to S24 and S23 perennial ryegrass grown for seed were investigated in a series of field experiments from 1971 to 1976. These varieties of ryegrass were found to be insensitive to timing of nitrogen application from apex initiation to the stage when ears first emerged. However, if nitrogen application was delayed until about 30% or more of the ears had emerged, yields were lower compared with earlier applications, this effect being significant when nitrogen was delayed until 70–80% ear emergence because of a decrease in both numbers of fertile tillers and number of seeds per unit area. No advantages were found for splitting nitrogen applications between apex initiation and ear emergence.     

Defoliation of ryegrass-dominated swards by sheep

Patterns of defoliation in perennial ryegrass-dominated swards were examined in two experiments. In Experiment I measurements were made on four swards after a single overnight grazing. In Experiment 2 observations were made repeatedly on marked tillers in two continuously stocked swards which were grazed down over a 24-d period. In both experiments the chance of defoliation was related to lamina height both within and between age classes. Lamina angle was measured in Experiment 2 and within age classes the chance of defoliation was also independent of lamina angle. In Experiment 1 tillers most frequently had both of the two youngest laminae grazed; in Experiment 2 tillers with only lamina 1 grazed occurred most frequently, but defoliation of the two youngest laminae was again common.
The different frequencies of grazing of the different classes of laminae is interpreted as being a function of their frequency of occurrence within the grazed layer at the top of the sward. The pseudostem apparently restricted the depth of the grazed layer in the very short swards of Experiment 2.
Grazing behaviour in relation to sward structure and the patterns of defoliation and their implications for herbage production are discussed.     

The influence of sward condition on rates of herbage growth and senescence in mixed swards under continuous stocking management

A study was conducted to determine the influence of herbage mass under continuous stocking management with sheep on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in a mixed-species sward of Lolium perenne. Poa annua and Trifolium repens. Plots were maintained in as nearly constant herbage mass as possible at 500, 700, 1000 and 1700 kg organic matter (OM) ha⁻¹ by continuous but variable stocking with sheep for the period May to July inclusive. Estimates of G, S and NP were obtained for each species from repeated measurements over two 2-week periods on individually identified grass tillers or clover stolons. Estimates of species population densities enabled estimates of G. S and NP per unit area to be made.
Rates of G were higher and of S were lower for Lolium tillers and Trifolium stolons than for Poa tillers. The laminae of Poa tillers occupied inferior positions in the canopy of the swards maintained at higher herbage masses and were less accessible to the grazing animals, so that Poa tissue was less well utilized.
The total population densities of tillers and stolons declined at herbage masses both above and below 700 kg OM ha⁻¹. The rate of G for the three species combined increased with herbage     

THE EFFECTS OF SEED RATE, CLOVER AND APPLIED NITROGEN ON BROADCAST SEED CROPS OF PASTURE RYEGRASS

This investigation was carried out with leafy varieties of perennial ryegrass at four centres near Aberystwyth over a period of three years. Nitrogen applied in spring increased lodging, the number of fertile tillers and also seed yield, except where sowing was made without a cereal cover crop. While reasonable yields were obtained from badly laid crops, it is suggested that lodging prior to anthesis has an adverse effect on seed yield. Clover included as a companion species tended to depress the yield of seed and increase the degree of lodging.
Seed rate had less infiuence on lodging and seed yield, but the data show that similar yields were obtained from light seed rates of 4 lb. per acre as from the heavier rates. On the other hand, there was a tendency for the low rate of seeding to produce weedy plots, especially when sown in the absence of a cover crop.     

Effect of seed crop management on the potential seed yield of contrasting white clover varieties.

Three contrasting white clover varieties, Olwen (large-leaved), Menna (medium-leaved) and S184 (small-leaved) were sown with and without a grass companion in 30 cm drills. Two grazing and one mechanical spring defoliation managements were imposed. Counts of the total number of inflorescences and of the percentage in various ripeness categories were made throughout the period of seed crop development to determine the pattern of inflorescence development and optimum harvest date. Inflorescence numbers were greatest in cv. S184 and least in cv. Olwen and, in general, varieties produced both more inflorescences and a greater proportion of ripe inflorescences when grown under spring defoliation managements similar to those for which they were bred. Thus, cv. Olwen produced more ripe inflorescences, and a lower proportion of brown inflorescences (semi-ripe), after mechanical defoliation while cv. Menna produced more ripe inflorescences following mechanical defoliation and rotational grazing. However, cv. S184 produced more ripe inflorescences under both grazing managements than under mechanical defoliation. Optimum harvest date was not affected by management or variety, total inflorescence number and maximum ripe inflorescences having reached a peak on 11 September. However, varieties differed in the proportion of ripe and brown (semi-ripe) inflorescences on this date, with cv. Menna and cv. S184 containing a significantly higher proportion of brown inflorescences than cv. Olwen, The implications of these differences in inflorescence development, the proportions of inflorescences in the various ripeness categories and their contribution to seed yield are discussed in relation to the management of white clover seed crops and harvesting method under UK climatic conditions.