青海海北高寒嵩草草甸系统水分利用效率特征

系统水分利用效率(WUE)是高寒草甸生态系统碳水耦合及植被演替的重要指标,但其时间格局和主要环境因素仍不清楚。基于青藏高原东北隅高寒嵩草草甸的涡度相关仪器观测的CO_2和H_2O通量数据,分析了2015年植被生长季(4月-10月)的系统水分利用效率(总初级生产力/植被蒸腾)的变化特征和主要环境影响因子。结果表明,总初级生产力(GPP)和植被蒸腾具有显著的单峰季节趋势,与群落叶面积指数(LAI)显著相关。生长季日均WUE为3.24g C·kg H_2O~(-1),没有显著的季节动态,呈现出"保守型"的特征。增强回归树的结果表明光合光量子通量密度(PPFD)、饱和水汽压差(VPD)和净辐射(Rn)是其主要调控因素,与WUE显著指数负相关(r~2>0.40,p<0.001)。     

不同放牧率对羊草+苔草低地草甸草地再生和净初级生产力的影响

在羊草 苔草低地草甸,通过小区围栏放牧,定量研究了不同放牧率对低地草甸牧草再生性能和地上净初级生产力的影响.结果表明:在不同放牧率下各再生期间的再生草量存在着明显的季节性差异,其最大值出现在5～6月份,为136.6 g/m2.适宜的放牧率有利于增加草地的再生草量,各放牧小区的相对生长速率均大于对照小区的相对生长速率,放牧率较低的P2小区相对生长速率最大.各放牧小区地上净初级生产力均小于对照小区地上净初级生产力,即不存在超补偿性生长现象.     

青海气候变化趋势及对植被生产力影响的研究

应用青海省南部三江源区、东北祁连山地及环青海湖区气象站1961-2004年气温、降水和所在地区植被地上净初级生产力资料,分析和模拟了44年来有关气候变化特征以及植被生产力与气温、降水、地理坐标参数间的关系,模拟估算了假设未来气候温暖化情景下青海植被生产力变化的可能。结果表明:44年来青海各地气温均在升高,青海北部比南部增温明显;年降水量变化平稳,但北部比南部有所增加;土壤实际蒸发散表现出明显的升高趋势;青海南部植被地上净初级生产力(NPP)逐年降低,青海东北地区相对平稳。模拟计算表明,由于青藏高原植被的生长主要受温度条件的限制,在未来气候增暖,降水不变或增加的趋势下,植被地上NPP均有所增加。     

荒漠草原不同地形条件下土壤水分和地上生物量的时空分异

运用方差分析、多重比较、Pearson相关系数及克里格插值方法,在时间和空间尺度上分析了荒漠草原地形因素对土壤水分及草地生物量异质性的影响以及土壤水分和生物量的关系。结果表明：荒漠草原地区土壤水分垂直分布规律明显,土壤水分的变异系数在0～20 cm最小、60～80 cm最大;地上生物量在整个生长季的空间异质性较大。3种地形条件下土壤水分含量及时空分布格局各不相同,说明地形是土壤水分异质性的关键影响因子;地形对植物的生长动态规律影响很大,却对地上生物量的空间分异影响较小。不同生育期,土壤水分与地上生物量的关系不同,生长初期地上生物量与60～100 cm土壤水分显著负相关,说明生长初期的土壤水分是植物生长的关键影响因子,并且植物生长耗水主要来自60～100 cm土层;生长旺期和末期的生物量与土壤含水量相关性较弱,这可能与干旱区植物耐旱器官的建成有关。
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气候变暖对高寒地区植物生长与物候影响分析

研究高寒地区植被在气候变化下的反应与适应等过程成为众多科学家所关注的问题。植物生长与物候是对气候变化敏感且易观测的指标,为更好得了解高寒地区植被生态系统对全球气候变化的响应机制,文中基于已有的研究并结合自己的实验分析了气候变暖对高寒地区植物生长与物候的影响。分析指出:1)温度升高对高寒植被有正效应,而温度持续升高,则对植被产生负效应,但增温效应都未达到显著水平,说明短期增温对该植被影响不显著;2)较高海拔地区(4500m)的植物高度有增加趋势,而较低海拔地区(3400m)的植物可能出现矮化;3)在高寒生态系统,温度升高能够加快植物物候进程,延长生长季,但物候的改变也会给其带来生存风险。     

高寒草甸植物群落的环境特征分析

分析了高寒矮嵩草草甸植物群落的环境特征,以及环境对群落的影响,高寒矮嵩草原甸分布区域,年平均气温较低,一般〈０℃。最热月７月平均气温〈１０℃,年内≥１０℃的积温不足１００℃,持续天数〈１０天,这里年长水量约为５８０ｍｍ,多分布于牧草生长季的５－９月,占年降水总量的８０％,土壤含水量较高,且表层大于深层,年内表现春季植物返青融湿期,雨季植物生长水分波动下降期,雨季末期植物枯黄水分上升期和冬季土壤冻结     

温度条件对辽宁南部玉米生长发育和产量的影响——以庄河分期播种试验为例

利用2012年庄河5期分期播种试验资料,分析温度条件对辽宁南部春玉米生长发育及产量的影响.结果表明:在光照和水分条件适宜情况下,玉米各生育期持续时间受温度条件影响较为明显,出苗、苗期及灌浆期随温度升高而缩短,其中出苗时间对日最低气温比日均温响应更为明显.玉米生长状态在营养生长阶段受温度影响较小,而产量形成过程因不同温度条件对干物质积累影响的差异而不同.产量形成不但取决于灌浆期积温,同时还受气温日较差和日最低气温的影响.对于庄河地区而言,适当晚播将有利于玉米各生育期经历更为适宜的温度条件进而提高产量.     

不同放牧率对草甸植被特征的影响

通过对羊草+苔草草甸进行受控放牧实验,定量研究了草地植被结构对不同放牧率条件下的反应特征。结果表明:所有不同放牧小区的四种牧草牧前高度远远低于对照小区,而且留茬高度差异不大。各种牧草放牧后的高度受草地群落生物量和牧草本身适口性的影响,适口性越高,降低程度越大。在整个生长季内,羊草+苔草草甸群落地上部分生物量一直处于不断增加的状态,其中苔草和芦苇地上生物量于6月份达到峰值;羊草于7月份达到峰值;香茅在整个生长季内不断增加,于8月份达到最大值。各放牧小区群落生物量均明显低于对照小区。随着放牧率的提高,多样性指数先增高后降低,即适度放牧有利于多样性的维持。     

胡杨、灰叶胡杨蒸腾耗水规律初步研究

以塔里木盆地3年生胡杨、灰叶胡杨幼苗为试验材料,采用盆栽方法模拟不同土壤干旱条件,研究胡杨、灰叶胡杨的蒸腾耗水规律。结果表明:生长期胡杨、灰叶胡杨的蒸腾耗水量表现出适宜水分>轻度胁迫>中度胁迫>重度胁迫,月蒸腾耗水量呈单峰变化,最高值出现在7月。不同土壤水分条件下,两树种月耗水量差异均较明显,说明土壤水分含量是决定树木蒸腾耗水量的主要因素;天气状况也影响苗木耗水量,耗水峰值多出现在晴天,峰谷多是阴天。不同土壤水分条件下灰叶胡杨单株耗水量均高于胡杨,水分条件较好时灰叶胡杨的生长较胡杨好,水分利用效率高,但随水分的减少胡杨的水分利用效率提高,表明胡杨对干旱环境的适应性较灰叶胡杨强,能有效地利用有限的水资源在荒漠生境中生存。     

东祁连山河谷高寒草地植被群落特征及其与土壤性状的关系

为探究东祁连山河谷高寒草地植被群落特征与土壤性状的关系,根据研究区河谷地形地貌及河谷高寒草地分布特征,选取了分布在河谷山地阳坡上部和下部、河谷山地阴坡上部和下部及河谷水平阶地的5类不同植被类型的高寒草地为研究对象。研究草地植被群落和土壤性状特征及相关关系。结果表明:河谷中不同坡向、坡位高寒草地植被群落优势种不同,河谷阳坡高寒草地以嵩草属植物为主,阴坡以蓼属植物为主;阴坡植被总盖度高,阳坡低;河谷阴坡下部草层高度最高,而水平阶地最小;植被群落总科数、总属数和总种数均表现为坡下部高于上部,阴坡高于阳坡;地上生物量依次为:阴坡下部>阳坡上部>阴坡上部>阳坡下部>水平阶地;0~60 cm土层地下生物量依次为:阴坡下部>阳坡下部>阴坡上部>水平阶地>阳坡上部。不同高寒草地土壤性状之间有显著差异(P<0.05)。相关分析表明,土壤含水量、容重、孔隙度和全氮等指标与植被总盖度、地上生物量、总科数、草地平均高度有显著的相关关系。可见,研究区河谷高寒草地植被与土壤之间的影响主要表现为土壤含水量、容重、孔隙度和全氮与植被总盖度、地上生物量、总科数、草地平均高度间的相互作用。     

Effects of warming and clipping on plant and soil properties of an alpine meadow in the Qinghai-Tibetan Plateau,China

Climate warming and livestock grazing are known to have great influences on alpine ecosystems like those of the Qinghai-Tibetan Plateau(QTP) in China. However, it is lacking of studies on the effects of warming and grazing on plant and soil properties in these alpine ecosystems. In this study, we reported the related research from manipulative experiment in 2010–2012 in the QTP. The aim of this study was to investigate the individual and combined effects of warming and clipping on plant and soil properties in the alpine meadow ecosystem. Infrared radiators were used to simulate climate warming starting in July 2010, while clipping was performed once in October 2011 to simulate the local livestock grazing. The experiment was designed as a randomized block consisting of five replications and four treatments: control(CK), warming(W), clipping(C) and warming+clipping combination(WC). The plant and soil properties were investigated in the growing season of the alpine meadow in 2012. The results showed that W and WC treatments significantly decreased relative humidity at 20-cm height above ground as well as significantly increases air temperature at the same height, surface temperature, and soil temperature at the depth of 0–30 cm. However, the C treatment did not significantly decrease soil moisture and soil temperature at the depth of 0–60 cm. Relative to CK, vegetation height and species number increased significantly in W and WC treatment, respectively, while vegetation aboveground biomass decreased significantly in C treatment in the early growing season. However, vegetation cover, species diversity, belowground biomass and soil properties at the depth of 0–30 cm did not differ significantly in W, C and WC treatments. Soil moisture increased at the depth of 40–100 cm in W and WC treatments, while belowground biomass, soil activated carbon, organic carbon and total nitrogen increased in the 30–50 cm soil layer in W, C and WC treatments. Although the initial responses of plant and soil properties to experimental warming and clipping were slow and weak, the drought induced by the downward shift of soil moisture in the upper soil layers may induce plant belowground biomass to transfer to the deeper soil layers. This movement would modify the distributions of soil activated carbon, organic carbon and total nitrogen. However, long-term data collection is needed to further explain this interesting phenomenon.     

巴音布鲁克主要草地类型表层土壤有机碳特征及其影响因素的研究

以新疆天山中段巴音布鲁克主要草地类型为研究对象,分析了不同草地类型土壤有机碳(SOC)的分布特征及其与气候因子、植被特征和土壤特性的关系。结果表明:分布在盆地海拔最低处的高山沼泽化草甸的土壤有机碳含量最高,而分布在盆地中间海拔的高山草原最低,分布在较高海拔的高寒草甸和高山草原化草甸处于高山沼泽化草甸和高山草原之间。0-30cm土壤有机碳含量与空气相对湿度、草地生产力、植被盖度、土壤含水量呈显著正相关(P<0.05),与土壤容重、土壤紧实度、土壤pH呈显著负相关(P<0.05),土壤有机碳与温度存在一定负相关,但未达到显著水平。偏相关分析显示,影响表层土壤有机碳含量最主要的因素是土壤含水量、土壤紧实度、草地生产力、空气相对湿度。     

Warming effects on plant biomass allocation and correlations with the soil environment in an alpine meadow,China

Alpine meadow ecosystem is fragile and highly sensitive to climate change.An understanding of the allocation of above-and below-ground plant biomass and correlations with environmental factors in alpine meadow ecosystem can result in better protection and effective utilization of alpine meadow vegetation.We chose an alpine meadow in the Qinghai-Tibetan Plateau of China as the study area and designed experimental warming plots using a randomized block experimental design.We used single-tube infrared radiators as warming devices,established the warming treatments,and measured plant above- (AGB) and below-ground biomass (BGB) during the growing seasons (May to September) in 2012 and 2013.We determined the allocation of biomass and the relationship between biomass and soil environment under the warming treatment.Biomass indices including above-ground biomass,below-ground biomass and the ratio of root to shoot (R/S) ,and soil factors including soil moisture and soil temperature at different depths were measured.The results showed that (1) BGB of the alpine meadow had the most significant allometric correlation with its AGB (y=298.7x~ (0.44) ,P0.001) ,but the relationship decreased under warming treatment and the determination coefficient of the functional equation was 0.102 which was less than that of 0.188 of the unwarming treatment (control) ; (2) BGB increased,especially in the deeper soil layers under warming treatment (P0.05) .At 0–10 cm soil depth,the percentages of BGB under warming treatment were smaller than those of the control treatment with the decreases being 8.52% and 8.23% in 2012 and 2013,respectively.However,the BGB increased 2.13% and 2.06% in 2012 and 2013,respectively,at 10–50 cm soil depths; (3) BGB had significant positive correlations with soil moisture at 100 cm depth and with soil temperature at 20–100 cm depths (P0.05) ,but the mean correlation coefficient of soil temperature was 0.354,greater than the 0.245 of soil moisture.R/S ratio had a significant negative correlation with soil temperature at 20 cm depth (P0.05) .The warmer soil temperatures in shallow layers increased the biomass allocation to above-ground plant parts,which leading to the increase in AGB;whereas the enhanced thawing of frozen soil in deep layers causing by warming treatment produced more moisture that affected plant biomass allocation.     

高寒草甸退化对土壤电导率变化影响的研究

土壤电导率是表征土壤水溶性盐的一个重要指标,可反映土壤盐渍化程度.为了研究高寒草甸退化对土壤电导率的影响,以三江源区未退化高寒草甸和退化高寒草甸为研究对象,系统分析了退化高寒草甸的植被特征和土壤特征与土壤电导率的相互关系.结果表明:高寒草甸退化会对土壤电导率产生显著负影响,且土壤电导率与评价高寒草甸的退化指标植被盖度、...     

新疆巴音布鲁克高山草地物种丰富度与生产力的关系

利用野外调查的物种丰富度和生物量资料,对位于新疆天山中段巴音布鲁克草地的20个样地进行群落数量分类,揭示每种群落类型的物种丰富度和生产力特征,在此基础上,考察了群落内和群落间物种丰富度与生产力的关系,并探讨其形成机制.结果表明:利用TWINSPAN将20个样地划分为6个群落类型(亚高山草原、亚高山草甸化草原、亚高山草原化草甸、亚高山草甸、高山草甸和高山沼泽草甸).6个草地群落类型地上生物量、物种丰富度差异明显:就地上生物量而言,亚高山草原和亚高山草甸较低,高山沼泽草甸和高山草甸较高,而亚高山草甸化草原和亚高山草原化草甸处于两者之间;就物种丰富度而言,亚高山草甸和高山草甸较高,亚高山草原和高山沼泽草甸较低,而亚高山草甸化草原和亚高山草原化草甸处于两者之间.群落内物种丰富度与生产力的关系表现出多种形式:亚高山草原、亚高山草甸和高山草甸的物种丰富度与生产力呈正相关,亚高山草甸化草原的物种丰富度与生产力呈负相关,而亚高山草原化草甸和高山沼泽草甸的物种丰富度与生产力没有表现出显著的相关关系.群落间物种丰富度与生产力呈现"单峰"关系.群落内生产力范围不够大,可能是群落内未能观察到"单峰"关系的原因;环境异质性是导致群落间出现"单峰"关系的关键因素.     

围栏与不同放牧强度对东祁连山高寒草甸植被和土壤的影响

在东祁连山高寒草地,对围栏7年和不同放牧强度的草地进行了物种数、地上生物量、地下生物量、土壤理化性质等研究。结果表明,围栏7年的高寒草地鲜草产量为425.8 g·m-2,显著高于夏季中牧159.3 g·m-2和夏季重牧91.0 g·m-2,但与冬季轻牧、夏季轻牧差异不显著。围栏条件下的物种数为26.3种·16 m-2,显著低于其他放牧条件下的物种数,但显著高于夏季重牧条件下的物种数23.0种·16 m-2;轻度或重度放牧都会使物种数减少,夏季中牧下的物种数最高(33.5种·16 m-2)。在0~10 cm的表层土壤中,围栏7年的草地根系生物量显著高于其他放牧强度。随着放牧强度的增加,根系生物量在0~10 cm土壤中呈下降趋势,在30~40 cm土壤中则表现为升高趋势。围栏7年的土壤容重低于其他放牧强度下的土壤容重,但差异不显著;夏季重牧的土壤容重显著高于围栏7年和其他放牧强度的土壤容重。随着放牧强度的增加,0~10 cm土壤碱解氮增加,围栏7年草地最低。围栏封育可有效改善和恢复草地植被,但不能长时间禁牧不进行放牧利用。合理的放牧能够维护高寒草甸草地生态系统功能、促进物种丰富度和土壤营养的均衡。     

Effects of long-term warming on the aboveground biomass and species diversity in an alpine meadow on the Qinghai-Tibetan Plateau of China

Ecosystems in high-altitude regions are more sensitive and respond more rapidly than other ecosystems to global climate warming.The Qinghai-Tibet Plateau(QTP)of China is an ecologically fragile zone that is sensitive to global climate warming.It is of great importance to study the changes in aboveground biomass and species diversity of alpine meadows on the QTP under predicted future climate warming.In this study,we selected an alpine meadow on the QTP as the study object and used infrared radiators as the warming device for a simulation experiment over eight years(2011-2018).We then analyzed the dynamic changes in aboveground biomass and species diversity of the alpine meadow at different time scales,including an early stage of warming(2011-2013)and a late stage of warming(2016-2018),in order to explore the response of alpine meadows to short-term(three years)and long-term warming(eight years).The results showed that the short-term warming increased air temperature by 0.31℃and decreased relative humidity by 2.54%,resulting in the air being warmer and drier.The long-term warming increased air temperature and relative humidity by 0.19℃and 1.47%,respectively,and the air tended to be warmer and wetter.The short-term warming increased soil temperature by 2.44℃and decreased soil moisture by 12.47%,whereas the long-term warming increased soil temperature by 1.76℃and decreased soil moisture by 9.90%.This caused the shallow soil layer to become warmer and drier under both short-term and long-term warming.Furthermore,the degree of soil drought was alleviated with increased warming duration.Under the long-term warming,the importance value and aboveground biomass of plants in different families changed.The importance values of grasses and sedges decreased by 47.56%and 3.67%,respectively,while the importance value of weeds increased by 1.37%.Aboveground biomass of grasses decreased by 36.55%,while those of sedges and weeds increased by 8.09%and 15.24%,respectively.The increase in temperature had a non-significant effect on species diversity.The species diversity indices increased at the early stage of warming and decreased at the late stage of warming,but none of them reached significant levels(P>0.05).Species diversity had no significant correlation with soil temperature and soil moisture under both short-term and long-term warming.Soil temperature and aboveground biomass were positively correlated in the control plots(P=0.014),but negatively correlated under the long-term warming(P=0.013).Therefore,eight years of warming aggravated drought in the shallow soil layer,which is beneficial for the growth of weeds but not for the growth of grasses.Warming changed the structure of alpine meadow communities and had a certain impact on the community species diversity.Our studies have great significance for the protection and effective utilization of alpine vegetation,as well as for the prevention of grassland degradation or desertification in high-altitude regions.     

高寒草甸不同类型草地土壤机械组成及肥力比较

研究了青藏高原高寒草甸不同类型草地土壤机械组成和土壤养分变化特征,并用相关分析探讨了土壤理化特征、土壤机械组成对不同草地类型群落物种组成、生物量变化的响应。结果表明:不同草地类型土壤机械组成分布大致是矮嵩草草甸:粉粒>细砂粒>粘粒>粗砂粒;高山嵩草草甸:细砂粒>粉粒>粘粒>粗砂粒;藏嵩草沼泽化草甸:细砂粒>粉粒>粘粒>粗砂粒;金露梅灌丛:粉粒>粘粒≥细砂粒>粗砂粒。矮嵩草草甸、高山嵩草草甸为粉砂质粘壤土,藏嵩草沼泽化草甸为壤土,金露梅灌丛为壤质粘土。矮嵩草草甸、高山嵩草草甸和金露梅灌丛土壤颗粒分布相对比较均匀(除藏嵩草沼泽化草甸外),主要集中在<0.5mm的范围内,土壤粘粒含量普遍大于20%。土壤全量养分和速效养分以及土壤物理特征均影响着高寒草甸不同草地类型土壤质量和土壤结构。土壤结构和养分状况是判断高寒草甸生态系统生态功能维持的关键指标之一。     

Effects of soil nutrients and climate factors on belowground biomass in an alpine meadow in the source region of the Yangtze-Yellow rivers,Tibetan Plateau of China

Improving our knowledge of the effects of environmental factors(e.g.soil conditions,precipitation and temperature) on belowground biomass in an alpine grassland is essential for understanding the consequences of carbon storage in this biome.The object of this study is to investigate the relative importance of soil nutrients and climate factors on belowground biomass in an alpine meadow in the source region of the Yangtze and Yellow rivers,Tibetan Plateau.Soil organic carbon(SOC),total nitrogen(TN) and total phosphorous(TP) contents and belowground biomass were measured at 22 sampling sites across an alpine meadow on the Tibetan Plateau.We analyzed the data by using the redundancy analysis to determine the main environmental factors affecting the belowground biomass and the contribution of each factor.The results showed that SOC,TN and TP were the main factors that influenced belowground biomass,and the contribution of SOC,TN and TP on biomass was in the range of 47.87%–72.06% at soil depths of 0–30 cm.Moreover,the combined contribution of annual mean temperature(AMT) and mean annual precipitation(MAP) on belowground biomass ranged from 0.92% to 4.10%.A potential mechanism for the differences in belowground biomass was caused by the variations in soil nitrogen and phosphorous,which were coupled with SOC.A significant correlation was observed between MAP and soil nutrients(SOC,TN and TP) at the soil depth of 0–10 cm(P0.05).We concluded that precipitation is an important driving force in regulating ecosystem functioning as reflected in variations of soil nutrients(SOC,TN and TP) and dynamics of belowground biomass in alpine grassland ecosystems.     

应用MODIS数据估算草地生物量

利用新一代EOS/MODIS卫星遥感数据,针对MODIS的应用特点,借鉴国内外基于遥感手段监测植被的方法以及植被指数的研究进展.以天山北坡乌鲁木齐南郊的草地为研究区,通过野外实地采样得到与MODIS影像资料时相一致的草地地上生物量数据.并利用ENVI软件,提取典型区各样点的植被指数,分析遥感植被指数与植被生物量的相关关系,从而建立植被指数在不同季节草甸和草原的生物量估测模型.