Forage selectivity by cattle and sheep co‐grazing swards differing in plant species diversity

Interactions between sward diversity and forage selectivity of cattle and sheep managed within mixed grazing systems are not fully understood. The objective of this study was to investigate the effect of either mono‐ or co‐grazing of sheep and cattle on swards differing in botanical composition (either diverse or grass‐dominated) on the intake choices of six target forage species. Jacobs' selection index (JSI) was employed to quantify the preference for single target species in relation to their proportion in the sward. Results revealed distinct intake preferences of sheep and cattle; as expected, sheep were more selective than cattle. To a lesser extent, the sward composition had an effect on intake preferences: cattle responded to botanical composition by changing their intake preferences, but to a lesser extent than sheep. Phleum pratense was the most preferred target forage species (JSI = 0.62). Lolium perenne, Taraxacum sect. Ruderale and Trifolium repens were also highly preferred regardless of sward composition or type of grazing (JSI = 0.47, 0.32 and 0.27 respectively). Dactylis glomerata and Festuca pratensis tended to increase in abundance after grazing events in both diverse and grass‐dominated swards. Co‐grazing of cattle and sheep facilitated a more homogeneous consumption of the target forage species evaluated.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

Effect of sheep, deer and cattle dung on herbage production and soil nutrient content

A field experiment compared the effects of sheep, deer and cattle dung on pasture growth and soil nutrient status. Sheep and deer dung pellets degraded rapidly within three to eight weeks, while it was 12 months before all visible cattle dung had gone from the soil surface. The cattle dung initially reduced herbage yield owing to smothering. However, after 40 days, herbage around the edges of the dung patch responded positively to the dung and more dry matter was produced in this patch than in the control patch during the first 12 months. There was no measurable herbage response to sheep dung, but deer dung significantly increased herbage production. While the rate of nutrient application per unit area was similar for sheep and deer dung, the latter was more evenly spread within the patch area. Consequently, a greater percentage of pasture plants in the patch received nutrient additions from the deer dung and a measurable response was recorded. Twelve months after dung application, soil nitrate, phosphate and organic C were higher in dung patches than in the control patches. In cattle dung patches, this response was accompanied by higher microbial biomass C levels, arylsulphatase and arylphosphatase activities. Some residual effect on organic C levels was still evident three years after cattle dung application.     

The utilization of sown and indigenous plant species by sheep and goats grazing hill pastures

The utilization of sown and indigenous plant species was studied in three experiments in which plots were stocked with similar live weights of sheep and goats. In the first experiment the animals grazed plots containing 0–5 ha of rush ( Juncus effusus )- infested reseeded pasture and 0–5 ha of unimproved blanket bog. The second and third experiments took place on old rush-infested improved pasture; in one experiment two levels of herbage mass of grass were provided while in the other the rushes were cut in spring or remained uncut.
The goats grazed the rushes readily in all three experiments. Reduction in herbage mass of grass increased utilization of rushes by goats although these animals still grazed rushes when grass supply was plentiful. Sheep scarcely grazed J. effusus even when hard-pressed by shortage of grass. Both sheep and goats grazed J. acutiflorus.
In Experiment I sheep utilized reseeded pasture more heavily than did goats. Sheep grazed similar proportions of the leaves of grass and clover while goats grazed a lower proportion of clover leaves as compared with grass. Among the grasses sheep discriminated in favour of Lolium perenne whereas goats did not. On the blanket bog vegetation, Eriophorum vaginatum and Calluna vulgaris were grazed both earlier in the season and more heavily by goats than by sheep.
The relationships between sward structure and grazing height in accounting for differences in species selection by sheep and goats are briefly discussed. The possibility of using goals to control coarse weeds in hill pasture and for strategic grazing to manipulate floristic composition is outlined.     

The use of a fluorescent pigment to record the distribution by cattle of traces of faeces from dung pats

The extent of post-defaecation redistribution of faeces by cattle in a grazed pasture was determined by adding to dung pats a pigment which is fluorescent in ultra-violet light. Traces of faeces which would normally not be detected were located under ultra-violet irradiation at night. More traces were redistributed from dung pats situated in areas where stock congregated than from dung pats in other areas of paddocks. Most traces occurred at distances of up to 2 m from each indicator dung pat treated with the pigment. The area of sward affected by faeces was over 100 times the area of individual dung pats. It is suggested that the distribution of infective parasitic larvae will follow the distribution of the traces rather than simply that of the dung pats. Grazing behaviour may also be affected by contamination of herbage by traces of faeces.     

Modelling phosphorus leaching to watercourses from extended autumn grazing by cattle

A modelling approach was undertaken to investigate the effect of grazing animals on phosphorus pollution of water draining from grazed fields. Following a similar study in relation to slurry spreading in winter, the dual‐porosity hydrological and contaminant transport model, MACRO, was calibrated to represent transport of phosphorus from faeces through the soil to field drains. Rapid flows, through water‐filled macropores in wet soil, of phosphorus sorbed onto colloidal particles in the faeces of cattle appeared to be the dominant transport mechanism. The outputs of the model supported experimental evidence that levels of phosphorus pollution of water draining from grazed fields can rise substantially if grazing is extended into late autumn, particularly if grazing is extended until soil has wetted up to around the field capacity water content. The critical housing date, to avoid phosphorus losses rising to an unacceptable level, varied considerably between years. The outputs of the model suggest that phosphorus pollution does not occur during grazing under dry conditions where soil macropores do not contain water, so that losses would remain low if animals are housed before soil substantially wets up in the autumn.     

The prediction of the in vivodigestibility of the diet of sheep and cattle grazing indigenous hill plant communities by in vitrodigestion, faecal nitrogen concentration or indigestible' acid-detergent fibre

Herbage and faeces samples were retained from an intake and in vivo digestibility trial using material harvested at three different stages from five indigenous hill plant communities (those dominated by Agrostis-Festuca., Nardus stricta., Molinia caerulea., Eriophorum vaginatum and Tri-chophorum caespitosum) and from sown swards of ryegrass or white clover. Samples of the herbages and of their separate components, together with extrusa samples of the same herbages recovered from pen-fed sheep and cattle oesophageal fistulates, were digested in vitro using rumen liquor. Measurements were made also of faecal nitrogen concentration (FN) and of indigestible acid-detergent fibre (IADF) using the samples from the original trial.
The range of in vitro disappearance values of the herbages (IVOMD; 0809-0278) was slightly wider than those of in vivo digestibility (OMD; 0-796-0-37I). Differences between OMD and IVOMD were greatest at low quality and relationships were best described by two separate linear regressions of OMD on IVOMD for (a) ryegrass, white clover, Agrostis-Festuca and Nardus (RSD; 0 0185) and, (b) Molinia, Tricho-phorum and Eriophorum (RSD; 00246). In vitro disappearance values of extrusa were higher than those of herbages offered by 00503 and 00156 units for sheep and cattle respectively, partly because of greater levels of ensalivation, especially in the sheep. The relationships of OMD to IADF and to FN were poor, though inclusion of faeces output and fitting parallel lines for communities in the regression of OMD on FN reduced the RSD to 0020. We conclude that in vitro digestion of samples of extrusa, using appropriate standards, is the best method of diet digestibility prediction for sheep and cattle grazing these communities.     

The effect of the spatial distribution of plant species within the sward on the searching success of sheep when grazing

The ability of grazing sheep to exploit preferred patches of grass species according to their spatial distribution was studied. The choices made by groups of four ewes in three cocksfoot ( Dactylis glomerata ) plots (80 × 80 m), in which 136 patches (30 × 30 cm) of perennial ryegrass ( Lolium perenne ) were either randomly distributed (plot Rand ) or grouped (plots Ag.1 and Ag.2 ) to create preferred feeding sites (four rich sites of twenty-five patches and four poorer sites of nine patches) were compared. On measurement days, each group was allocated to one of the three plots, and grazed it in at least four successive 30-min tests. At the end of the experiment the ewes were allowed to graze the plots continuously for 3 months to assess the medium-term effect of grazing sheep on the depletion rate of patches. Behavioural observations showed that, once the ewes had discovered a preferred feeding site in plots Ag.1 or Ag.2 , they frequently returned to it within a test and between tests, whereas they failed to learn the precise location of many isolated patches in plot Rand . Thus in 2 h of grazing, the ewes spent 0·40 more time feeding on perennial ryegrass when the patches were grouped than when they were randomly distributed, and perennial ryegrass depletion was greater in plots Ag.1 and Ag.2 than in plot Rand (0·193 vs. 0·105 of initial height). The richer sites were also more depleted than the poorer ones (0·209 vs. 0·152 of initial height). When the plots were continuously grazed, perennial ryegrass depletion remained greater in plots Ag.1 and Ag.2 than in plot Rand for at least 3 weeks. Hence the spatial distribution of patches can affect diet selection by sheep, and so probably their impact on grasslands.     

Effects of sward type and rest periods from sheep grazing on white clover presence in perennial ryegrass/white clover associations

The effects of the imposition and timing of a rest period from continuous sheep stocking, for a conservation cut, on white clover presence in forty perennial ryegrass/white clover associations were studied over two full grazing seasons. Each association consisted of one grass variety along with one white clover variety, the grasses being diploid and tetraploid ryegrasses from each of five maturity groups and the white clovers from each of four leaf size categories. The presence of white clover within each association was assessed at the beginning and end of both seasons by means of a 0–64-m² quadrat subdivided into 100 squares, each 80 mm × 80 mm, the number of squares in which any part of a white clover plant was visible being recorded. Complementary point quadrat data were also collected. Although continuous sheep stocking did not necessarily have an adverse effect on white clover presence, a July to mid-August rest period increased white clover proportions in the swards (means: unrested, 48–1; April to late May rest, 32.7; July to mid-August rest, 67.3 - s.e.d. 7.59; P < 0.05) the benefit increasing with increasing white clover leaf size. The early rest period (April to late May) reduced white clover presence and the late rest period (July to mid-August) increased white clover presence, these effects being intensified with increasing white clover leaf size (very large-leaved clover: unrested, 20.6; April to late May rest, 8.3; July to mid-August rest, 41.1 and small-leaved clover: unrested, 96.3; April to late May rest, 84.8; July to mid-August rest, 97 - s.e.d. 9.2; P <0001). Tetraploid ryegrass/white clover associations had consistently and significantly more white clover than diploid ryegrass/white clover associations of similar ryegrass maturity group (tetraploid, 53.4; diploid 44.8 - s.e.d. 2.12; P <0.001) and associations with early maturing ryegrass contained more white clover than those with late maturing ryegrasses, the effect of maturity group being greater than that of ploidy. Overall, white clover presence increased with increasing openness of grass growth habit.     

The spatial distribution of excreta under intensive cattle grazing

The spatial distribution of excreta from intensive cattle grazing was recorded and was described by a negative binomial function. This enabled the proportion of a pasture surface affected by excreta to be calculated for various grazing periods. Where the effect of excreta on the sward was short-lived, as for urine N, or where the total area covered by excreta remained small, as for dung, then little overlapping of effective excreta patches could be expected. Appreciable overlapping of effective patches could, however, occur for the long-lived urine K effect and this should be taken into account when estimating the proportion of a pasture surface affected.     

The effect of fasting on short-term cattle grazing behaviour

The effects of fasting on feeding behaviour, intake rates, and diet quality were studied with steers grazing vegetative Lolium multiflorum swards. Fasting interrupted the normal alternation of grazing and rumination and led to longer grazing times (0·65 vs. 0·45 of observed time), shorter rumination times (0·15 vs. 0·30 of observed time), and higher total dry matter (DM) intakes (6200 g DM vs. 3750 g DM d⁻¹)-Fasting did not affect prehended bite size which averaged 047 g DM bite⁻¹. Declining bite size within a day was explicable by changes in herbage mass. Biting rate (38.9 bites min⁻¹) and instantaneous intake rate (2675 g DM min⁻¹) of fasted animals were considerably higher than non-fasting animals in the mornings, but neutral detergent fibre (NDF) concentration of the diet (0 742 of organic matter) was generally not depressed by fasting. Consequently, non-fasted animals appeared to be feeding suboptimally since their nutrient intake rates were considerably lower than that which they are capable of exhibiting. Mastication rates per unit NDF intake were considerably lower in the morning for fasted animals (0·78 chews g⁻¹ NDF intake) which suggested that their intake rates may have led to larger ingested particle size. Therefore, on Lolium multifiorum swards, the steers attained a higher intake rate by masticating the forage less while maintaining diet quality.     

Short-term fasts and the ingestive behaviour of grazing cattle

Twelve tethered Angus cows (Bos taurus) weighing 471 ± 18 kg were used in a balanced changeover design to estimate the direct (D), residual (R), and permanent (P) effects of 1,2, and 3 h fasts following 2 h grazing sessions on the ingestive behaviour of cattle grazing swards of endophyte- (Acremonium coenophialum Morgan-Jones and Gams)-free tall fescue (Festuca arundinacea Schreb. cv Kentucky 31) (Exp I) and lucerne (Medicago sativa L. cv Arc) (Exp II). Herbage dry-matter (DM) mass (> 5 cm) and individual herbage DM allowances were 1-16 t ha-1 and 50 kg per hour in Exp I and 1-861 ha-1 and 78 kg per hour in Exp II, respectively. In Exp I, P effects of fasting followed trends of R effects since the D effects were small. Mean DM intake rates (P) were 2-2., 1-8, and 1-3 kg h-1 for cows given 1, 2, and 3 h fasts, respectively. Mean intakes (DM) per bite (P) were 082, 071, and 045 g bite-1 for cows assigned 1, 2, and 3 h fasts, respectively. Rates of biting were not affected by fasting and averaged 45 bites min"'. In Exp II, P effects followed trends in D effects since R effects were small. Mean DM intake rates (P) were 20 , 22, and 3-0 kg for cows subjected to 1,2, and 3 h of fast, respectively. Rates of biting (P) were 24, 27, and 32 bites min-1 for cows after 1, 2, and 3 h fasts, respectively. DM intakes per bite (P) were not affected by fasts and averaged 1-46 g bite-1. The behavioural responses of cattle to interval between meals were attributed to differences in flow characteristics of ingesta of lucerne and tall fescue in the gastrointestinal tract.     

The impact of cattle dung pats on earthworm distribution in grazed pastures

Background

Grazed grassland management regimes can have various effects on soil fauna. For example, effects on earthworms can be negative through compaction induced by grazing animals, or positive mediated by increases in sward productivity and cattle dung pats providing a food source. Knowledge gaps exist in relation to the behaviour of different earthworm species i.e. their movement towards and aggregation under dung pats, the legacy effects of pats and the spatial area of recruitment. The present study addressed these knowledge gaps in field experiments, over 2 years, using natural and simulated dung pats on two permanent, intensively grazed pastures in Ireland.

Results

Dung pats strongly affected spatial earthworm distribution, with up to four times more earthworms aggregating beneath pats, than in the control locations away from pats. In these earthworm communities comprising 11 species, temporally different aggregation and dispersal patterns were observed, including absence of individual species from control locations, but no clear successional responses. Epigeic species in general, but also certain species of the anecic and endogeic groups were aggregating under dung. Sampling after complete dung pat disappearance (27 weeks after application) suggested an absence of a dung pat legacy effect on earthworm communities. Based on species distributions, the maximum size of the recruitment area from which earthworms moved to pats was estimated to be 3.8 m² per dung pat. Since actual grazing over 6 weeks would result in the deposition of about 300 dung pats per ha, it is estimated that a surface area of 1140 m² or about 11% of the total grazing area can be influenced by dung pats in a given grazing period.

Conclusions

This study showed that the presence of dung pats in pastures creates temporary hot spots in spatial earthworm species distribution, which changes over time. The findings highlight the importance of considering dung pats, temporally and spatially, when sampling earthworms in grazed pastures. Published comparisons of grazed and cut grasslands probably reached incorrect conclusions by ignoring or deliberately avoiding dung pats. Furthermore, the observed intense aggregation of earthworms beneath dung pats suggests that earthworm functions need to be assessed separately at these hot spots.

     

A comparison of methods of recovering oesophageal extrusa from grazing sheep

Extrusa samples were collected from two groups of oesophageally fistulated sheep grazing together. One group had foam plastic oesophageal plugs fitted below the fistula to assist in the complete collection of extrusa; the second group grazed without plugs. The extrusa were collected in four periods chosen to cover as wide a range of herbage conditions as possible varying from herbage with a high proportion of green leaves to predominantly dry material with a high proportion of dead leaves and stems. The plant components of the extrusa were determined and there was no evidence that stems or any other plant component preferentially by-pass the fistula in grazing sheep. It was concluded that it is not necessary to insert foam plastic plugs in the oesophagus of grazing sheep to obtain representative samples of ingested herbage.     

Performance of white clover varieties combined in blends and alone when grown with perennial ryegrass under sheep and cattle grazing

Abstract Two experiments were carried out to evaluate the performance of blends of three white clover (Trifolium repens L.) varieties in comparison with the component varieties and three other varieties sown individually in a mixture with perennial ryegrass (Lolium perenne L.). The plots were grazed rotationally in Experiment 1 by cattle and sheep and in Experiment 2 by sheep alone. In both experiments, the blend was composed of three medium‐leaved varieties (AberDai, AberVantage and AberHerald), but with different relative contributions of the three varieties in the two experiments. Dry matter (DM) yields of white clover and perennial ryegrass were assessed in replicate plots for two years (1999 and 2000) after the establishment year. In Experiment 1, there was no significant difference between the DM yields of white clover or perennial ryegrass in either year. The decline in DM yield of white clover between years that was observed for some varieties was not found for the blend. In Experiment 2, significant differences were found in DM yields of white clover in both years. In 1999, AberDai had the highest DM yield. In 2000, AberHerald and AberVantage had the highest DM yields, and AberDai showed a decline in DM yield that was mirrored by the mean for all the white clover varieties. In both experiments, the blend did not show significantly higher DM yield than one or more of its components; indeed, in Experiment 2, it was significantly lower yielding than AberDai in 1999. However, where one component of the blend declined in DM yield between years, this was compensated for by an increase in the yield of another component. These preliminary findings suggest that the yield stability of blends may give them a potential role in agricultural practice.     

Herbage intake and N excretion by sheep grazing monocultures or a mixture of grass and white clover

In 1988 and 1989, swards of grass (G0), while clover (C0) and grass/white clover (GC0) receiving no N fertilizer, and a grass sward supplied with 420 kg N ha^?1 (G420), were grazed by non-lactating sheep to maintain a sward surface height of 6 cm. Herbage organic matter (OM) intakes averaged between 1200 and 1700 g OM ewe^?1 d^?1. For treatments G0, C0, GC0 and G420 respectively, the ewes' live weight gain was 102, 112, 100 and 110 g d^?1 and changes in body condition scores were +0·28, +0·52, +0·36 and +0·44 units season^?1. However, the effect of treatment was not significant for either variable. There were similar levels of output of faecal N ewe^?1 but significantly more urinary N ewe^?1 was excreted on treatments C0 and G420, where the concentrations of N in herbage laminae were also higher. For example, in 1989, total daily N excreted was 39·7, 64·4, 44·0 and 63·3 g N ewe^?1 for G0, C0, GC0 and G420 respectively. Taking into account the mean daily stocking rates, which were 19·4, 26·6, 27·2 and 36·5 ewe ha^?1, the total faeces and urine returns over the season were 161, 358, 249 and 484 kg N ha^?1 for each treatment respectively. The herbage OM intakes ewes^?1 d^?1 measured in September and October were similar for C0 and G420, and so the intake of herbage OM ha^?1 d^?1 was related to stocking rate, i. e. the estimated herbage intake ha^?1 over the growing season for the white clover monoculture was 73% of that for N-fertilized grass. Excretal nitrogen returns to the pasture from grazed mono-cultures of clover were high, and similar to those from a grass sward receiving 420 kg fertilizer N ha^?1. Consequently potential losses of N to the environment are high under these management systems.     

The nutritional evaluation of Italian ryegrass cultivars fed as silage to sheep and cattle

Three feeding trials with big bale silage were carried out to determine whether previously established intake differences between artificially dried Italian ryegrass cultivars could be repeated under differing conditions of food preparation and presentation. The silage was prepared in 2 consecutive years from the same field crop and fed to groups of cattle in both years. The second year silage was also fed to sheep indoors. All animals were fed ad libitum and the voluntary intakes recorded for groups of cattle and individual sheep. Chewing activity was also recorded on all groups of sheep.
The chemical composition of the two cultivars as fresh grass was similar in 1984 but there were small differences in 1983 with Tarquin showing slightly higher organic matter digestibility (DOMD) and lower dry matter and water-soluble carbohydrate compared to RvP. The only consistent differences in silage composition between cultivars was a slightly higher level of modified acid detergent (MAD) fibre in RvP compared to Tarquin. Despite these small differences the voluntary intake of cattle fed Tarquin was 19% higher than RvP in the first year and 35% in the second, while sheep showed a similar difference of 40% with the second year silage.
Recording of jaw activity with sheep showed that animals spent 40% more time eating and 27% more time ruminating RvP than Tarquin per unit of food eaten. This suggested that the intake differences, which were independent of digestibility, were related to the physical resistance of the feed to breakdown by chewing. These differences depend on the structural characteristics of the plants and were associated with differences in leaf/stem ratios and MAD-fibre content. These are less likely to be affected by the method of conservation or feed presentation and, although the degree of difference between the cultivars changed with the different procedures, the inherently higher intake of Tarquin persisted.     

Acoustic measurement of intake and grazing behaviour of cattle

Acoustic analysis of grazing behaviour was found to allow accurate identification of chewing and biting, and estimation of intake by cattle. Four steers grazed six types of turves, three were short (14, 16 and 20 cm tall), leafy turves from mowed Setaria lutescens with high application of N fertilizer, and three were tall (36, 49 and 62 cm tall), mature turves from uncut areas with low application of N fertilizer. Each animal grazed ten bites from each turf type while behaviour was videotaped and grazing sounds were recorded with a wireless microphone taped to the animal's forehead. The intake was measured by the difference in pre- and post-grazing turf weight, corrected for water loss. Chews and bites differed in acoustic characteristics, with bites producing more output in the 3–8 kHz frequency band. Discriminant analysis of the acoustic characteristics correctly classified chews and bites with 94% accuracy. Intake was predicted with a high degree of accuracy by total energy flux density (fJ m^–2) in chewing sounds (EFDC), EFDC per chew and average intensity of chewing sound in the 1–2 and 2–3 kHz bands ( R ²=0·90, CV=3·6%). Although bite mass ranged from 1·37 to 4·50 g, EFDC per g of intake (CV=0·015) was not affected by treatments or animals. Chewing sounds carried more information about intake than biting sounds and chews per bite. Analysis of grazing sounds has the potential to overcome many of the problems associated with the measurement of grazing intake.     

Relationships between the yield of perennial ryegrass and of small-leaved white clover under cutting or continuous grazing by sheep

Seven varieties or advanced breeding lines of white clover ( Trifolium repens L.), all of small leaf size, were grown separately in mixtures with perennial ryegrass ( Lolium perenne L.) in an experiment encompassing three harvest years. Harvestable dry-matter (DM) yield measurements were taken of these mixtures and of perennial ryegrass monocultures under two management regimes: cutting and continuous sheep grazing. Considerable differences were observed in the harvestable DM yields of white clover, perennial ryegrass and total yields of the mixtures between plots containing different white clover varieties. White clover yields were generally higher under cutting, and perennial ryegrass yields were higher under grazing. The difference between perennial ryegrass yield in monoculture and in mixture was variable. In the second harvest year, a significant interaction effect was seen between management and white clover variety for white clover yield but not for perennial ryegrass yield. The relationship between clover yield and grass yield differed between the two management regimes. Under cutting, a negative correlation was observed, indicative of competitive effects. However, under grazing, no such correlation was seen. Possible mechanisms underlying these outcomes are discussed.