Effects of age-related increases in sapwood area,leaf area,and xylem conductivity on height-related hydraulic costs in two contrasting coniferous species

Introduction  

Knowledge of vertical variation in hydraulic parameters would improve our understanding of individual trunk functioning and likely have important implications for modeling water movement to the leaves. Specifically, understanding how foliage area (A _l), sapwood area (A _s), and hydraulic specific conductivity (k _s) vary with canopy position to affect leaf-specific conductivity (LSC) and whole-tree leaf-specific hydraulic conductance (K _l) may explain some of the contrasting patterns of A _l/A _s reported in the literature.     

冀北山地森林土壤水分物理性质和入渗规律研究

为了进一步了解冀北山地森林土壤水分物理性质和入渗规律,以华北落叶松桦木混交林、山杨桦木混交林和油松蒙古栎混交林为研究对象,对其土壤水分物理性质和土壤入渗性能进行测定,结果表明:随土层深度的增加土壤容重逐渐增大,土壤持水量和总孔隙度逐渐减小;油松蒙古栎混交林的土壤平均容重最大(1.11g/cm3),华北落叶松桦木混交林最小(0.92g/cm3);土壤最大持水量由大到小依次为山杨桦木混交林(63.55%)、华北落叶松桦木混交林(53.10%)、油松蒙古栎混交林(39.50%),田间持水量由大到小依次为山杨桦木混交林(49.95%)、落叶松桦木混交林(41.01%)、油松蒙古栎混交林(31.39%);土壤总孔隙度由大到小依次为华北山杨桦木混交林(51.61%)、落叶松桦木混交林(46.14%)、油松蒙古栎混交林(43.18%);华北落叶松桦木混交林稳渗速率由大到小依次为坡下、坡中、坡上,山杨桦木混交林和油松蒙古林混交林稳渗速率由大到小依次为坡中、坡上、坡下。     

Restoration thinning and influence of tree size and leaf area to sapwood area ratio on water relations of Pinus ponderosa

Ponderosa pine (Pinus ponderosa Dougl. ex P. Laws) forest stand density has increased significantly over the last century (Covington et al. 1997). To understand the effect of increased intraspecific competition, tree size (height and diameter at breast height (DBH)) and leaf area to sapwood area ratio (A(L):A(S)) on water relations, we compared hydraulic conductance from soil to leaf (kl) and transpiration per unit leaf area (Q(L)) of ponderosa pine trees in an unthinned plot to trees in a thinned plot in the first and second years after thinning in a dense Arizona forest. We calculated kl and Q(L) based on whole- tree sap flux measured with heat dissipation sensors. Thinning increased tree predawn water potential within two weeks of treatment. Effects of thinning on kl and Q(L) depended on DBH, A(L):A(S) and drought severity. During severe drought in the first growing season after thinning, kl and Q(L) of trees with low A(L):A(S) (160-250 mm DBH; 9-11 m height) were lower in the thinned plot than the unthinned plot, suggesting a reduction in stomatal conductance (g(s)) or reduced sapwood specific conductivity (K(S)), or both, in response to thinning. In contrast kl and Q(L) were similar in the thinned plot and unthinned plot for trees with high A(L):A(S) (260-360 mm DBH; 13-16 m height). During non-drought periods, kl and Q(L) were greater in the thinned plot than in the unthinned plot for all but the largest trees. Contrary to previous studies of ponderosa pine, A(L):A(S) was positively correlated with tree height and DBH. Furthermore, kl and Q(L) showed a weak negative correlation with tree height and a strong negative correlation with A(S) and thus A(L):A(S) in both the thinned and unthinned plots, suggesting that trees with high A(L):A(S) had lower g(s). Our results highlight the important influence of stand competitive environment on tree-size-related variation in A(L):A(S) and the roles of A(L):A(S) and drought on whole-tree water relations in response to thinning.     

A model of heat transfer in sapwood and implications for sap flux density measurements using thermal dissipation probes

A variety of thermal approaches are used to estimate sap flux density in stems of woody plants. Models have proved valuable tools for interpreting the behavior of heat pulse, heat balance and heat field deformation techniques, but have seldom been used to describe heat transfer dynamics for the heat dissipation method. Therefore, to better understand the behavior of heat dissipation probes, a model was developed that takes into account the thermal properties of wood, the physical dimensions and thermal characteristics of the probes, and the conductive and convective heat transfer that occurs due to water flow in the sapwood. Probes were simulated as aluminum tubes 20 mm in length and 2 mm in diameter, whereas sapwood, heartwood and bark each had a density and water fraction that determined their thermal properties. Base simulations assumed a constant sap flux density with sapwood depth and no wounding or physical disruption of xylem beyond the 2 mm diameter hole drilled for probe installation. Simulations across a range of sap flux densities showed that the dimensionless quantity k [defined as (ΔT(m) -ΔT)/ΔT, where ΔT(m) is the temperature differential (ΔT) between the heated and unheated probe under zero-flow conditions] was dependent on the thermal conductivity of the sapwood. The relationship between sap flux density and k was also sensitive to radial gradients in sap flux density and to xylem disruption near the probe. Monte Carlo analysis in which 1000 simulations were conducted while simultaneously varying thermal conductivity and wound diameter revealed that sap flux density and k showed considerable departure from the original calibration equation used with this technique. The departure was greatest for variation in sap flux density typical of ring-porous species. Depending on the specific combination of thermal conductivity and wound diameter, use of the original calibration equation resulted in an 81% under- to 48% overestimation of sap flux density at modest flux rates. Future studies should verify these simulations and assess their utility in estimating sap flux density for this widely used technique.     

A model of moisture movement in wood based on water potential and the determination of the effective water conductivity

Summary A model of isothermal moisture movement in wood during drying using the gradient in water potential as the driving force is proposed. The moisture transport coefficient used in this model is the effective water conductivity. It is a function of moisture content, temperature, and direction of flow. The boundary desorption curve of the effective water conductivity function is established in the radial and tangential directions of aspen sapwood from nearly saturated to dry conditions at 20, 35, and 50 °C using the instantaneous profile method. The results show that the effective water conductivity increases exponentially with moisture content and temperature. The effect of temperature cannot be solely explained by the variation of the viscosity of water. The variation of the moisture content-water potential relationship with temperature would explain a large part of this effect. The effective water conductivity was generally higher in the radial direction than in the tangential direction in a ratio varying from 1/1 to 25/1 depending on moisture content and temperature. The flux-gradient relationship obtained at given moisture contents were found to be linear, confirming the validity of the model for the experimental conditions considered in the present work.The authors wish to thank Peter Garrahan of Forintek Canada Corp., Ottawa for his comments and suggestions. This research was undertaken while the senior author was a grant-holder from the Fonds pour la Formation de Chercheurs et l'Aide à la Recherche, Gouvernement du Québec     

Effect of wide spacing on tree growth, branch and sapwood properties of young Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] in south-western Germany

The influence of stand density on Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] is conceptually understood, but for wide spacings not well quantified, particularly in Europe. This study used 41 trees from 7 different locations in south-western Germany to compare important tree- and branch-level attributes across three different densities, namely 100, 200, and 1,200 stems ha⁻¹. In general, there were only a few tree and branch attributes that were significantly different between the 100 and 200 ha⁻¹ densities. Crown projection area and diameter of the thickest branches were the most important differences between the 100 and 200 ha⁻¹ densities. The most obvious and significant differences in this study were between 100 and 1,200 ha⁻¹ densities, where nearly every examined tree and branch attributes were statistically significant. However, relative sapwood area, the number of branches, branch angle, and the occurrence of spike knots were insensitive to stand density. Although the two lowest stand densities in this study represent rather extremely wide spacings, these results still have important implications for the development of effective thinning regimes for Douglas-fir in south-western Germany. Important management recommendations from this study include thinned stands should be maintained to at least 200 stems ha⁻¹ to maintain high log quality and stand stability. Furthermore, even at stand densities exceeding more than 1,200 trees ha⁻¹ planted trees, artificial pruning may even be necessary to produce high quality logs.     

Effects of water vapour heating on structure and properties of cork

Summary The effects of heating cork in water vapour at 100°C and 300°C are studied and compared with those produced by heating in air at the same temperatures. Dimensional and mass changes were measured and radial compression curves were obtained following various treatments. The 300°C treatments originate a straightening of the originally corrugated cell walls, while the cell wall material undergoes thermal degradation. Larger swellings and larger mass changes are observed in the treatments in water vapour as compared to those in air. The 100°C treatments do not induce degradation and produce reversible changes in technological properties which are simply due to differences in the water content.     

The effect of bark decortication for hiwada production on xylem and phloem formation in Chamaecyparis obtusa

Of all plant materials used to cover the roofs of traditional Japanese buildings, Japanese cypress (Chamaecyparis obtusa) bark, hiwada, has the longest service life and has been used from ancient times. However, wood and bark properties after hiwada harvest have not been evaluated in detail. We studied whether decortication for hiwada production in winter affected xylem and phloem formation. Decorticated trees still preserved all inner bark and part of the outer bark, and both decorticated and control trees had similar annual ring structures at all stem heights in the xylem and phloem. In both xylem and inner bark, no significant difference in ring width at any stem height was found between annual rings before and after decortication. Thus, this study revealed that the decortication of bark for hiwada production does not affect the formation of xylem and the inner and outer bark if decortication is carried out by highly skilled workers in winter.     

基于Google Earth的大围山水土保持信息系统研究

为加强大围山国家森林公园水土保持的研究与管理,在水土保持地理信息子系统、监测信息子系统、专家系统中引入了Google Earth,设计和构建了基于Google Earth的大围山水土保持信息系统。通过对该系统框架及应用方法和前景的阐述,并比较了该系统与其它相关信息系统的优势和不足,实现对大围山水土流失数据信息的高效存储和管理。研究表明:Google Earth在水土保持信息系统中具有重要的推广应用价值。     

A potential role for xylem-phloem interactions in the hydraulic architecture of trees: effects of phloem girdling on xylem hydraulic conductance

We investigated phloem-xylem interactions in Acer rubrum L. and Acer saccharum Marsh. Our experimental method allowed us to determine xylem conductance of an intact branch by measuring the flow rate of water supplied at two delivery pressures to the cut end of a small side branch. We found that removal of bark tissue (phloem girdling) upstream of the point at which deionized water was delivered to the branch resulted in a decrease (24% for A. rubrum and 15% for A. saccharum) in branch xylem hydraulic conductance. Declines in hydraulic conductance with girdling were accompanied by a decrease in the osmotic concentration of xylem sap. The decrease in xylem sap concentration following phloem girdling suggests that ion redistribution from the phloem was responsible for the observed decline in hydraulic conductance. When the same measurements were made on branches perfused with KCl solution (approximately 140 mOsm kg(-1)), phloem girdling had no effect on xylem hydraulic conductance. These results suggest a functional link between phloem and xylem hydraulic systems that is mediated by changes in the ionic content of the cell sap.     

Determination of reaction rate parameters for the acid copper chromate fixation reactions on oak (Quercus castaneifolia C. A. Mey) sapwood

Acid copper chromate fixation reactions on oak (Quercus castaneifolia C. A. Mey) sapwood were characterized by at least two distinguished reaction periods which differ in their kinetic behavior, particularly reaction rate. In this research, chemical rate equation parameters were studied in detail as a function of chromium fixation time. Theoretically, the kinetic data were distinguished between the adsorption and reduction reactions and consequently, the rate parameters were determined for each stage distinctly. The obtained results showed that at the investigated thermal range (25–33°C), almost for every 3°C increase in temperature the reaction duration has decreased 20.4?min for the first reaction period and 78?min for the second reaction period. Besides that, in all sample replications, up to 58.3% of hydrogen ions function at the reduction reactions of the initial period, demonstrating that acidity (H⁺ ions) has a major effect on reaction rate constant. Accordingly, adsorption reactions of the hexavalent chromium at the starting point follow a pseudo-first-order kinetic reaction. Different to the adsorption reaction the Cr^VI reduction reaction is exothermic with a measured exothermal energy of 48.2?kJ/mol.     

A noninvasive optical system for the measurement of xylem and phloem sap flow in woody plants of small stem size

Over the past 70 years, heat has been widely used as a tracer for estimating the flow of water in woody and herbaceous plants. However, most commercially available techniques for monitoring whole plant water use are invasive and the measurements are potentially flawed because of wounding of the xylem tissue. The study of photosynthate transport in the phloem remains in its infancy, and little information about phloem transport rates is available owing to the fragility of the vascular tissue. The aim of our study was to develop a compact, stand-alone non-invasive system allowing for direct detection of phloem and xylem sap movement. The proposed method uses a heat pulse as a tracer for sap flow. Heat is applied to the surface of the stem with a near-infrared laser source, and heat propagation is monitored externally by means of an infrared camera. Heat pulse velocities are determined from the thermometric data and related to the more useful quantity, mass flow rate. Simulation experiments on the xylem tissue of severed silver birch (Betula pendula Roth.) branch segments were performed to assess the feasibility of the proposed approach, highlight the characteristics of the technique and outline calibration strategies. Good agreement between imposed and measured flow rates was achieved leading to experimentation with live silver birch and oak (Quercus robur L.) saplings. It was demonstrated that water flow through xylem vessels can be monitored non-invasively on an intact stem with satisfactory accuracy despite simultaneous sugar transport in the phloem. In addition, it was demonstrated that the technique allows for unequivocal detection of phloem flow velocities.     

A case-study of water transport in co-occurring ring- versus diffuse-porous trees: contrasts in water-status, conducting capacity, cavitation and vessel refilling

Recent work has suggested that the large earlywood vessels of ring-porous trees can be extraordinarily vulnerable to cavitation making it necessary that these trees maintain a consistent and favorable water status. We compared cavitation resistance, vessel refilling, transport capacity and water status in a study of ring-porous Quercus gambelii Nutt. (oak) and diffuse-porous Acer grandidentatum Nutt. (maple). These species co-dominate summer-dry foothills in the western Rocky Mountains of the USA. Native embolism measurements, dye perfusions and balance pressure exudation patterns indicated that the large earlywood vessels of 2-3-year-old oak stems cavitated extensively on a daily basis as predicted from laboratory vulnerability curves, resulting in a more than 80% reduction in hydraulic conductivity. Maple branches showed virtually no cavitation. Oak vessels refilled on a daily basis, despite negative xylem pressure in the transpiration stream, indicating active pressurization of embo-lized vessels. Conductivity and whole-tree water use in oak were between about one-half and two-thirds that in maple on a stem-area basis; but were similar or greater on a leaf-area basis. Oak maintained steady and modest negative xylem pressure potentials during the growing season despite little rainfall, indicating isohydric water status and reliance on deep soil water. Maple was markedly anisohydric and developed more negative pressure potentials during drought, suggesting use of shallower soil water. Although ring porosity may have evolved as a mechanism for coping with winter freezing, this study suggests that it also has major consequences for xylem function during the growing season.     

A general framework for the quantification and valuation of ecosystem services of tree-based intercropping systems

This study provides the first complete framework for the valuation of ecosystem services of agroforestry and uses a tree-based intercropping (TBI) system in southern Québec, Canada, as a case study. Ten ecosystem services were estimated, all of which were of interest and directly applicable to most agricultural systems worldwide: nutrient mineralization, water quality, soil quality, pollination, biological control, air quality, windbreak, timber provisioning, agriculture provisioning, and climate regulation. A mix of mathematical models for the quantification and economic valuation of various ecosystem services were used. The results revealed a total annual margin of $2,645 ha^?1 y^?1 (averaged over 40 years). The economic value of combined non-market services was $1,634 ha^?1 y^?1, which was higher than the value of marketable products (i.e. timber and agricultural products). An analysis of the present value suggested that agricultural products ranked highest among the ecosystem services taken singularly, followed by water quality, air quality, climate regulation, and soil quality maintenance. Total economic value of all ecosystem services for the rotation period was $54,782 ha^?1, only one third of which was contributed by agricultural products. Although the total value of the ecosystem services provided by TBI was high, farmers only benefited from agricultural and timber products. Thus, government incentives are needed to interest farmers in adopting practices that benefit society as a whole.     

Effects of water soaking and outdoor exposure on nail joint properties of particleboard

Phenol-formaldehyde resin-bonded particleboard (PF board) and isocyanate resin-bonded particleboard (MDI board) were soaked in water at 40, 70 and 100 °C, and the relationships between soaking conditions and nail joint properties were analyzed. The soaking time to reach the lower limit of nail-head pull-through (NHPT) of the PF board was 2 h at 100 °C, while it took 168 h at 70 °C. The soaking time to reach the lower limit of lateral nail resistance (LNR) of the PF board was 24 h at 100 °C, but it did not take 168 h at 70 °C to reach it. The lower limits of NHPT and LNR for the MDI board were higher than those for the PF board. For the PF board, there was a high correlation between modulus of rupture, internal bond strength and nail joint properties. Based on the results of water soaking and outdoor exposure, it was shown that thickness change has a significant effect on NHPT and LNR, and that the reduction in NHPT and LNR results from the collapse of bonding points owing to swelling of the board.     

A model of tree-crop competition for windbreak systems in the Sahel: description and evaluation

A model was developed to simulate the effects of competition for soil water and radiation between windbreaks and pearl millet crops in the Sahel. These effects on millet (Pennisetum glaucum (L.) R. Br.) growth were simultaneously simulated for each millet row parallel to the windbreak with small time steps for soil water processes, radiation availability and crop assimilation. The crop routine of the model was based on an existing semi-deterministic model. The soil-water flow was simulated in two dimensions to account for horizontal gradients. Competition for water was expressed by distributing the available soil water between trees and millet in proportion to its uptake rates in a non-competitive situation. Competition for light was incorporated as light reduction through a two-dimensional (windbreak) barrier with time-increasing height and density. Tree parameters were introduced as fixed values or as time-dependent forcing functions. Crop, windbreak trees (Bauhinia rufescens Lam.), and soil data inputs were either field-determined or obtained from literature. Reasonable agreement between simulated and measured soil water content and dry matter production was obtained under the conditions in Niger. Global radiation intensities and soil water contents were simulated satisfactorily as a function of time and the distance from the windbreak. Hence, the model is appropriate to analyse competition for light and water between windbreaks and crops.     

柿树春季带木质部单芽嵌接育苗技术试验

试验采用君迁子做砧木,通过在春季带木质部单芽嵌接法快速繁育技术的试验,为新品种柿树的快速扩繁提供科学依据,试验表明春季在君迁子和柿树枝芽同时萌动时,采用带木质部单芽嵌接法快速繁育柿树,成活率达95%以上,嫁接育苗成效显著。     

落叶松直径动态变化规律及其机理的研究

作者依据标准地的资料来源,采用数据分析的方法,对落叶松直径动态变化规律进行研究,并得到一定的结论,以供参考。     

使用林地可行性报告编制的技术问题及对策探讨

以国家林业局2002年颁发的《使用林地可行性报告编写规范》为技术依据,系统阐述了使用林地可行性报告编制的实质要求、可行性评价的主要技术路线及几个关键技术问题,提出了规范可行性报告编制管理工作的对策措施。