Spatial and structural uniformity of lowland agricultural grassland in England: a context for low biodiversity

The contention that agricultural grassland within the UK is spatially and structurally uniform across wide differences in livestock farming intensity is examined. Total nitrogen (N) input was used as a surrogate for management intensity. Farms with average annual N inputs of >200 kg ha^?1 were categorized as highly intensive, those with N inputs of 50–200 kg N ha^?1 as moderately intensive and those with inputs of <50 kg N ha^?1 were categorized as extensive. Four farms within each management category were selected in two discrete regions: one in south‐west England, typifying a landscape dominated by agricultural grassland, the other in south‐east England with a more mixed‐farming landscape. Specific N input and management data, and sward botanical composition and structure, were obtained on four fields per farm. The south‐west region had a higher proportion of farms with dairy cattle than in the south‐east England region. The average N input and stocking rates were higher within the intensive‐management category farms in south‐west England than in south‐east England. Old permanent grasslands contained variable amounts of sown species, such as Lolium perenne, and were dominated by a few (generally <5) non‐sown grass species. Grassland that received >75 kg N ha ^?1 year^?1 contained <7 plant species and <3 forb species m^?2. Grassland communities with >12 plant species and >5 forb species m^?2 were only found in study fields that received <50 kg N ha^?1 year^?1. Grasslands with >10 forb species m^?2 were found only in sites receiving <15 kg N ha^?1. Of importance at a landscape scale was ubiquity of species‐poor and similar grassland plant communities across the management intensity range within and between livestock farms. Grassland in the extensive management category was similar in structure in terms of sward height and/or sward density to more intensively managed grasslands. This study revealed, albeit in two lowland sample regions in the UK, the ubiquity of grass‐dominated, species‐poor and structurally uniform grasslands, irrespective of apparently broad differences in farming intensity. The consequences of this spatial uniformity on grassland biodiversity are likely to be profound.     

1986 survey of upland grassland in England and Wales: changes in age structure and botanical composition since 1970–72 in relation to grassland management and physical features

A sample of 10000 ha of grassland in seven upland districts originally surveyed in 1970–72 was re-surveyed in 1986. Forty percent of the area was above 240 m elevation, 38% had gradients above 81/2°, 29% had poor or bad drainage and 54% was classified as having difficult or severe limitations to management. Twelve percent had been reseeded within the previous 5 years and 39% within 20 years; 6% was classified as arable grassland (in rotation with crops). Thirty-four percent was usually mown, two-thirds of it for silage. In most years 72% of grassland received fertilizer N (average application 123 kg N ha⁻¹) and 35% received organic manure; nearly 20% received neither. Older swards tended to receive lower rates of fertilizer N than reseeds, hay fields and grazed fields less than silage fields, and fields on beef/sheep farms less than those on dairy farms. The proportion of sown species in swards averaged 35%, of which Lolium perenne accounted for 23%. A good contribution of Trifolium repens was recorded in 15% of the grassland. Herb species were recorded in 30% of the grassland, mostly in situations with low fertilizer N under hay or grazing. Agrostis spp., Poa spp., Holcus lanatus and Festuca rubra were the main unsown grasses. The average contribution of L. perenne had increased since 1970–72; it was also present in two-thirds of the area from which it was absent in 1970–72, whilst the contribution of T. repens had decreased. The incidence of Cirsium arvense and Ranunculus spp. had also decreased, whilst that of Pteridium aquilinum had increased.     

Hay production from lowland semi-natural grasslands: a review of implications for ruminant livestock systems

Data on total, seasonal and between-year variability in the productivity and nutritional quality of cut forages obtained from a range of lowland semi-natural grasslands are reviewed. Dry-matter production, as hay, from unfertilized species-rich semi-natural grasslands ranged from less than 20% to about 80% of the production that might be expected from agriculturally improved and intensively managed grasslands. Unfertilized species-rich grasslands had lower growth rates, reflecting low nutrient availability in spring and early summer. Low soil phosphorus availability appears to be a key factor allowing high species-richness to be maintained. The in vitro digestibilities of herbage cut for hay from semi-natural grasslands were 20%, or more, below values for conserved forages from intensively managed grassland. When losses incurred during the hay-making process were taken into account, then the nitrogen content of hays from many semi-natural grasslands appear to be sub-optimal for productive livestock. Metabolizable energy values of herbage cut for hay were 10–40% below values for intensively managed conserved grass. Phosphorus content of forages from most species-rich grasslands was either below the metabolic requirement of livestock or inadequate to sustain high individual animal performance. Potential problems associated with supplementation of the diet with phosphorus, and with high calcium-to-phosphorus ratios, are discussed. Although potassium content of most of the herbage cut for hay was adequate for productive ruminant livestock, exceptions arise with hay cut after July. The magnesium content of cut herbage from some semi-natural grasslands was below an adequate level for productive livestock. Most cut herbage appeared to contain adequate amounts of sodium. Inadequacy of the mineral content of hays from semi-natural grasslands for ruminant livestock will be exacerbated by nutrient losses incurred during the hay-making process and particularly so when hay- making is prolonged by wet weather. Few feeding studies using hays from semi-natural grasslands of high nature conservation interest have been conducted. Lack of such data makes it difficult to plan any integrated use of these hays in ruminant feed budgets for livestock systems.     

The productivity of grassland farms in seven climatic zones of England and Wales

Records of grassland productivity were kept for 2 years on 152 dairy and 179 beef farms. Results were collated for six lowland zones, delineated primarily on the basis of average rainfall, and an upland zone. Contemporary and long-term average meteorological records were also collected.
In most of the lowland zones stocking rates and use of fertilizer N were similar, but utilized metabolizable energy (UME) output from grass varied; it was 25% higher in the wet, cool zone of north-west England and east Wales than in the dry zone of eastern England. The ranking of zones for utilized output corresponded closely with the ranking for summer rainfall.
On upland farms stocking rate was 25% lower and UME output 15% lower than on lowland farms, but this was achieved from little more than half the N input.
The differences between zones were similar to those shown in other published farm data. They also showed a similar trend to that demonstrated in grass cutting experiments. This suggests that farmers were, on average, able to exploit the extra grass grown in wetter climates. The incidence of difficult topography and impeded drainage was similar in dry and wet zones, but did have a more serious effect in the wet zones. The much higher level of concentrate feeding in the driest zone may have reduced the utilization of grass.     

The long-term effects of a range of pasture treatments applied to three semi-natural hill grassland communities.

Five pasture treatments were applied to three semi-natural hill grassland communities. The pasture treatments were: (1) controlled grazing, (2) controlled grazing + lime, (3) controlled grazing + lime + phosphate, (4) controlled grazing + lime + phosphate + oversown white clover and (5) controlled grazing + lime + phosphate + oversown white clover + oversown perennial ryegrass. The communities were dominated by Agrostis/Festuca (site 1), Molinia (site 2) and Nardus (site 3). The Nardus at site 3 was substantially reduced by herbicide before treatments were applied. All treatments were grazed simultaneously by mature wether sheep on three occasions each year to a residual herbage mass of 560 kg DM ha^?1. There were two grazing periods, each of 4 weeks duration, beginning in mid-May and mid-July respectively and a further grazing period of 3 weeks duration beginning in mid-October. The number of sheep grazing days and liveweight gain of sheep were recorded during each grazing period for 13 years at each site. Organic matter intake (OMI) and digestibility (OMD) of ingested herbage were measured in years 2, 5 and 13. Mean daily OMI per head ranged from 1290 g for treatment 1 to over 1400 g for treatments 4 and 5. OMI values were higher for the Agrostis/Festuca site (mean OMI 1450 g) than for the Molinia and Nardus sites (1310 g and 1370 g respectively) largely owing to differences in the values for treatments 1, 2 and 3. OMI values for each treatment decreased by around 300 g between May and October. Mean OMD values from treatments 1, 2 and 3 were higher for the Agrostis/Festuca site (0·66) than for the Molinia and Nardus sites (0·63 and 0·64 respectively). The OMD values for treatments 1, 2 and 3 at the Molinia and Nardus sites declined by between 0·05 and 0·02 between years 2 and 13 and also showed the greatest decrease between May and October (0·08). OMD values for treatments 4 and 5 (0·67 and 0·69 respectively) were higher than for treatments 1, 2 and 3 at all sites and levels were maintained over 13 years. Individual liveweight gains of sheep increased from treatment 1 (?18 g d^?1) to treatment 5 (82 g d^?1), but they decreased over time on all treatments. The Agrostis/Festuca site gave 17% more annual sheep grazing days than the Molinia site and 33% more than the Nardus site. The mean number of annual sheep grazing days for each treatment ranged from 2250 for treatment 1 to 3640 for treatment 5. Annual sheep grazing days increased over 13 years by between 35 and 45% for treatment 1 and 55 and 70% for treatment 5. The results are described in relation to the changes in pasture composition and herbage accumulation.     

Observations on the incidence of plant parasitic nematodes in grassland in England and Wales

Soil samples taken from one- to four-year-old swards of white clover and perennial ryegrass at 34 sites, and from older clover/grass swards at 56 sites in England and Wales, were examined for plant parasitic nematodes. Stolons of white clover from 46 of these sites were examined for nematodes. Clover cyst nematode ( Heterodera trifolii ) and white clover stem nematode ( Ditylenchus dipsaci ) were found in 62% and 41% of sites respectively. The grass cyst nematode ( Punctodera punctata ) was present in 56% of 71 sites examined, and other grass cyst nematodes ( H. bifenestra and species of the H. avenae group) occurred in 13% and 17% of sites. Two root-knot nematodes were found, Meloidogyne naasi and M. hapla, in 15% and 4% of the samples examined. Migratory nematodes were extracted from 63 samples, and spiral, pin and lesion nematodes were found in more than 90%. Stunt nematodes were less frequent (65%) and criconematids (19%) were found relatively infrequently. The significance of these observations to grassland productivity is discussed.     

The long-term effects of a range of pasture treatments applied to three semi-natural hill grassland communities. 1. Pasture production and botanical composition

Five pasture treatments: (1) controlled grazing, (2) controlled grazing + lime, (3) controlled grazing + lime + phosphate, (4) controlled grazing + lime + phosphate + oversown white clover and (5) controlled grazing + lime + phosphate + oversown white clover + oversown perennial ryegrass were applied to three semi-natural hill grassland communities. The communities were those dominated by Agrostis/Festuca, Molinia and Nardus. The proportion of Nardus at the Nardus-dominant site was substantially reduced by herbicide before treatments were applied. All treatments were grazed simultaneously by mature wether sheep on three occasions each year. There were two grazing periods each of 4 weeks duration between mid-May and mid-August with a further grazing period of 3 weeks starting mid-October. During each grazing period stock numbers were set so that a residual herbage mass of 560 kg DM ha ^?1 remained at the end of the grazing period. Measurements of net herbage accumulation (NHA) were made annually over a period of 13 years at each site. The green:dead ratio of grasses, species composition of the pasture and the pH of the soil were measured at intervals during the experiment. Estimates of mean annual NHA ranged from 3860 kg DM ha^?1 for treatment 1 (controlled grazing) to 5170 kg DM ha^?1 for treatment 5 (oversown white clover and perennial ryegrass). The application of lime and phosphate increased annual NHA by 300–350 kg DM ha^?1 with a further increase of around 400 kg DM ha^?1 when white clover was sown. Increases in NHA between year 1 and year 13 ranged from 30% for treatment 1 to around 55% for treatment 5. Although there was no difference in the mean NHA between sites, the herbage from the Agrostis/Festuca site contained a higher proportion of green grass and white clover than that from the other sites. The highest levels of green grass and white clover were found on the oversown treatments at each site. The grazing pressure exerted produced relatively little change in the botanical composition at the Agrostis/Festuca site. At the Molinia-dominant site the Molinia was largely replaced by Nardus during the first 6 years and Nardus also increased in cover at the Nardus site. Application of lime and phosphate generally increased the proportion of Poa pratensis, Festuca rubra and Agrotis tenuis but did not halt the spread of Nardus at the Molinia and Nardus sites. White clover and perennial ryegrass were successfully introduced by oversowing and proportions remained high throughout the 13 years. The cover by bryophyte spp. increased at all sites with the greatest increases occurring in each case on the less comprehensive pasture treatments. These results suggest that on Agrostis/Festuca-dominant pastures herbage biomass production can be increased with relatively low-cost pasture treatments while maintaining Species diversity. However, Nardus and Molinia dominant pastures are likely to require more comprehensive pasture treatments involving sown grasses and white clover to provide herbage of acceptable quality and avoid an increase in Nardus and bryophytes in the sward. With a regime of episodic summer grazing and the addition of fertilizers oversown pastures can be maintained over long periods of time.     

Botanical composition of permanent grassland in England and Wales in relation to soil, environment and management factors

Grassland in England and Wales accounts for nearly 60% of the enclosed agricultural land and approximately 50% is at least 20 years old. Botanical composition, especially amongst older grassland, is extremely variable.
A survey of 502 grassland farms was conducted in the 1970s. Factors affecting botanical composition of individual fields have been investigated using correlation and regression analyses and swards of similar botanical composition grouped using cluster analysis. Lolium perenne, Agrostis spp. and Holcus lanatus were numerically the most important species, contributing, on average, 35%. 21% and 10%, respectively. Most older swards contained Lolium perenne and Agrostis spp. in varying proportions depending on drainage, soil fertility and management. Swards containing a significant proportion of Trifolium repens were associated with adequate drainage, good soil nutrient status, low inputs of fertilizer nitrogen and hard grazing. Of the other common species Holcus lanatus was associated with poor drainage, low soil fertility, hay cutting, short-season grazing and low fertilizer N. Festuca rubra was mainly in older swards, at higher elevations and with low fertilizer N but heavy stocking. Swards containing Rumex spp., Poa trivialis and other sown grasses were associated with mowing together with reasonably high soil fertility.     

South-west England grassland survey 1983. 2. Trends in cropping pattern, reseeding and sward composition 1970–83

Seventy-eight farms in south-west England, originally visited in 1970–72, were revisited in 1983 in order to identify changes in land use and re-seeding over the c. 12-year period; changes in sward composition were assessed on fifty-nine of these farms. The results were also compared with Ministry of Agriculture, Fisheries and Food (MAFF) Census results and MAFF Seeds Branch data.
The proportion of tillage had declined from 29 to 24%. This was in agreement with census results for the south-western counties, but contrasted with those for England and Wales as a whole, which showed an increase from 38.8 to 41.7% tillage. It is indicative of grass becoming more predominant in south-west England as east England has become more dominated by tillage cropping.
There had also been a decline in the amount of short-term grassland. In 1970–72 17% of the grassland was intended to last from 1 to 4 years; in 1983 the proportion was only 10%. That these intentions were being carried out was confirmed by the fact that in 1983 there was more grassland in each of the age categories 5·8,9·20 and over 20 years. The trend towards older grassland confirms that shown by regional and national census data, and the trend away from short-term swards is in line with deliveries of herbage seed which indicate reduced use of Italian ryegrass (Lolium multiflorum).
Botanical composition had changed appreciably, with the proportion of Lolium perenne having increased in swards of all ages, e.g. from 35 to 51% in swards 5–8 years old and from 17 to 31% in those aged over 20 years. Other sown species, predominantly Dactylis glomerata and Phleum pratense , had declined markedly in swards over 4 years old. The proportions of Agrostis spp., Festuca spp. and Poa trivialis had fallen, but Holcus lanatus had increased slightly. There had been a marked reduction in infestations of perennial broad-leaved weeds, particularly Ranunculus , but also Rumex and Cirsium spp.     

Potassium cycling and losses in grassland systems: a review

Cycling of potassium in grassland systems has received relatively little attention in research and practice in recent years. Balanced nutrient systems require consideration of nutrients other than nitrogen (N). Potassium (K) is needed in large amounts and is closely related to N nutrition. In intensive dairy farming, surpluses of K arise from the input of concentrates and fertilizer and are returned to the grassland and may lead to increasing K content in the soil. Organic farming, on the other hand, is characterized by limitations in input of nutrient sources and quantities. Leaching of K from grassland is usually low, but high levels of available soil K, high K input from fertilizer or at urine patches lead to increasing losses. High K inputs have a negative influence on Mg and Ca uptake by plants and can cause accelerated leaching of these cations. High levels of K have been associated with inducing nutrition‐related dairy cow health problems such as milk fever (hypocalcaemia) and grass tetany (hypomagnesaemia). This review gives an overview of the cycling of potassium and related cations in grassland systems especially with regard to leaching losses and identifies limitations to knowledge.     

The conservation management of upland hay meadows in Britain: a review

Upland hay meadows conforming to MG3 in the National Vegetation Classification of the UK are a rare habitat in Britain and are largely confined to upland valleys in northern England. Agricultural intensification, particularly ploughing and reseeding and a shift from hay‐making to silage production over the last 50 years, has resulted in large losses of species‐rich upland hay meadows. Remaining species‐rich meadows have been the focus of much nature conservation effort resulting in many of the species‐rich sites being protected by statutory designations or through voluntary agri‐environment scheme agreements. Research and monitoring has tended to confirm that species richness is maximized by management involving spring and autumn grazing, a mid‐July hay cut, no inorganic fertilizer and possibly low levels of farmyard manure. Deviations from this regime result in a loss of species richness. Restoration of semi‐improved grassland to swards resembling species‐rich MG3 also requires a similar regime but is also dependent on the introduction of seed of appropriate species. The role of Rhinanthus minor as a tool for manipulating meadow biodiversity during restoration management is discussed. Suggestions for future research are outlined.     

Extensification of grassland use in the Welsh uplands: sheep performance in years 1–6

An experiment was established in 1991 on a 25-year-old perennial ryegrass/bent ( Lolium perenne L. /Agrostis capillaris L.)-dominated pasture in Wales to study the effects of reducing nutrient inputs to previously fertilized upland pasture. The effects of the removal of applications of (1) N (denoted by CaPK) (2) N, P and K (Ca) and (3) N, P, K, and Ca (Nil) were compared with a treatment which received applications of all four nutrients (CaPKN) over a 6-year period (1991–96) in a randomized block design replicated three times. The experiment was managed under a continuous variable stocking regime (ewes and lambs until weaning and ewes thereafter) maintaining a sward surface height of 4·0 cm throughout the grazing season. Although individual liveweight gain of the lambs was unaffected by the treatments, there was a significant reduction ( P  < 0·05) in total lamb liveweight gain, ewe stocking rate and length of grazing season as a result of the withdrawal of nutrients. Over the 6 years total lamb liveweight gain was reduced by 17%, 32% and 45% and ewe stocking rate by 21%, 36% and 49% on treatments CaPK, Ca and Nil, respectively, compared with treatment CaPKN. The effect of withdrawing nutrient inputs on ewe stocking rate was progressive and by 1996 the Nil input treatment displayed a 63% reduction compared with the CaPKN treatment and this was also coupled with a 21-day reduction in length of the grazing season. During the post-weaning period, ewes from the Nil input treatment recorded a liveweight loss in 1995 and only a modest liveweight gain in 1996. This coupled with significantly lower body condition scores ( P  < 0·01) of these ewes in the autumn indicated that the Nil input treatment could lead to reductions in reproductive performance.     

The deposition of Caesium 137 on grassland at a site in SW England following the Chernobyl accident

Following the Chernobyl accident in April 1986 ¹³⁷Cs was detected in samples of grass collected at AGRI, North Wyke, Devon, UK. Samples of grass from permanent and reseeded swards under uniform management were taken on three occasions. The ¹³⁷Cs content of the samples was assessed as both Bq kg^-1 of sample and Bq m^-2, Samples taken from the same site in 1985 were used as a control group; these contained no measurable ¹³⁷Cs. The samples taken on 8 May 1986 had a ¹³⁷Cs content of approximately 30 Bq kg^-1; by 31 July 1986 the content had dropped to 1% of this value. The permanent pasture initially showed a higher level of ¹³⁷Cs than the reseeded sward. Analysis of soil cores from the sampling area showed a much higher level of ¹³⁷Cs than that found in the herbage. This was attributed to the atmospheric testing of nuclear devices in the early 1960s: the short-lived ¹³⁷Cs isotope was found in the 1986 herbage samples but not in the soil cores.     

The conservation management of mesotrophic (meadow) grassland in northern England. 1. Effects of grazing, cutting date and fertilizer on the vegetation of a traditionally managed sward

The results are reported from an experiment on the effects of cutting date (14 June, 21 July and I September), fertilizer application (none or 80 kg ha^?1 N plus 40 kg ha^?1 P and K) and grazing treatments (none, autumn or autumn plus spring) on the vegetation of an upland mesotrophic grassland in Upper Teesdale. northern England, UK. Effects on plant species number and cover are reported for 4 years (1989–93) of treatment. Effects on ‘species -attributes’ are given for the fourth year. The cessation of grazing combined with the use of fertilizer progressively reduced species number by about 25%. Under traditional management (no fertilizer, cutting date on 21 July, autumn and spring grazing) the species number and cover remained relatively static over the 4 years. Comparison between treatments in the fourth year showed a reduction in species number under the fertilizer application, cutting date on 1 September and no-grazing treatments. Fertilizer use together with cutting date on 1 September particularly lowered species number and cover. Analysis of variance was used to assess the effect of treatment on species that occurred frequently in the sward. A cutting date of 1 September favoured Agrostis capillaris. Alopecurus pratensis, Poa trivialis, Phleum pratense and Trisetum flavescens, The absence of grazing favoured Dactylis glomerata and Holcus lanatus. The use of fertilizer particularly favoured A. pratensis and H. lanatus. Ordination methods were used to assess the effect of treatment on the less frequent species. These were primarily associated with the treatment combination that matched ‘traditional’ management. Deviations from this ‘traditional’ regime acted separately, rather than in combination, and favoured different grass species. Traditional management was associated with ruderal, stress-tolerant ruderal and competitive ruderal strategists and with longer seed germination times, heavier seeds, some of which needed scarifying or chilling to break dormancy, and transient seed banks that germinated in the autumn. The original sward was an Anthoxanthum odora-turn-Geranium sylvatirum grassland, Briza media subcommunity (MG3b). After 4 years, Festuca ovina-Agrostis capillaris-Galium saxatile grassland, Holcus lanatus-Trifolium repens subcom-munity (U4b) and Lolium perenne-Alopecurus pratensis-Festuca pratensis grassland (MG7c) were found in many of the fertilized and late-cutting treatments.     

The composition of first-cut grass for ensilage in England and Wales from 1988 to 1991

A total of 2225 samples of herbage, typical of that taken for first-cut silage on commercial farms in England and Wales, was taken between 1988 and 1991. Samples were analysed for dry matter (DM), water soluble carbohydrate (WSC), crude protein (CP) and metabolizable energy calculated from modified acid detergent fibre. The results were used to investigate the effect of weather conditions upon herbage quality for ensilage. In general, in areas of high rainfall, i.e. western areas, herbage DM and WSC were significantly lower than in eastern areas, whilst the reverse situation occurred with CP. Italian ryegrass (IRG) had a significantly higher WSC and lower CP than other grasses, whilst samples from permanent pasture (PP) had significantly lower DM and WSC, and significantly higher CP than other grasses. It was calculated that to produce a WSC concentration in the fresh grass of 37 g kg⁻¹, to produce well-fermented silage without additive use, IRG grown in the east would require a DM of 160 g kg⁻¹, In contrast PP grown in the west would require a DM content of 250 gkg⁻¹.     

Implications of climate change for grassland in Europe: impacts, adaptations and mitigation options: a review

Climate change associated with greenhouse gas (GHG) emissions may have important implications for Europe's grasslands. Projected scenarios indicate that increased temperatures and CO₂ concentrations have the potential to increase herbage growth and to favour legumes more than grasses, but changes in seasonal precipitation would reduce these benefits particularly in areas with low summer rainfall. Further implications for grasslands may arise from increased frequency of droughts, storms and other extreme events. Potential farm-scale adaptive responses to climate change are identified. Grassland agriculture also contributes to GHG emissions, particularly methane and nitrous oxide, and management of grassland affects net carbon balances and carbon sequestration. Management options are identified for mitigating grassland's contribution to GHG emissions which need to be developed in a holistic way that also considers other pressures.     

The effect of stocking rate and fertilizer usage on income variability for dairy farms in England and Wales

A mathematical model of grass production and utilization on a dairy farm is described. Using the model an assessment has been made of the extent to which financial uncertainty arising from year-to-year variability in grass yields, coupled with a preference among farmers for minimizing risks, may explain the relatively low stocking rates and observed nitrogen usages on many dairy farms in England and Wales. The degree of risk has been equated with the probability of profits in a particular year being less than those required to cover the consumption needs and short-term borrowing requirements of the farmer. The results of the analysis indicate that a strategy of minimizing risks may lead to a significantly lower stocking rate than one of maximizing profits. Thus, considerations of risk may lead to stocking rates which are suboptimal from the viewpoint of economic and biological efficiency. However, there is no evidence to suggest that the comparatively low average usage of nitrogen on dairy farms in the UK is determined by risk considerations. On the contrary, increasing nitrogen usage lowers the apparent financial risk at a given stocking rate.     

Effects of species aggregation, habitat and season on the accuracy of double-sampling to measure herbage mass in a lowland grassland ecosystem

Double‐sampling, as a cost‐effective and sufficiently accurate method, is an acknowledged method for estimating herbage mass at the level of the plant species up to the total plant community. It was investigated whether adjusting visual estimations of double‐sampling is applicable at different levels of species aggregation when different seasons and habitats are involved. The method was used in six different habitats of the same ecosystem, a grassland nature reserve in western Belgium, based on eight sampling periods. A general linear model was used to analyse the data at four levels of species aggregation. The correlation between estimated and measured values of herbage mass was significant for the majority of graminoid species at the levels of the sub‐life‐form and total plant community but not at the level of the life‐form. Interaction of habitat type and measured herbage mass, and interaction of date and measured herbage mass, were significant for a minority of species. A significant interaction of both date and habitat with measured herbage mass was found in some cases at the level of the sub‐life‐form and total plant community. Results suggest that the double‐sampling method is reliable for most graminoids and at the sub‐life‐form level, but should be applied more cautiously for forbs species, and when different sampling periods and habitat types are involved at the level of the life‐form. To avoid producing several calibration lines at the level of the species and to overcome deficiency of observations at this level, lumping morphologically similar species from the same vegetation stratum in sub‐life‐form groups is recommended for adjusting estimated herbage mass.     

The conservation management of mesotrophic (meadow) grassland in Northern England. 2. Effects of grazing, cutting date, fertilizer and seed application on the vegetation of an agriculturally improved sward

The plant species number and composition, iind yield of herbage biomass of an agriculturally improved hay meadow were assessed after 4 years under various combinations of grazing, fertilizer applicution. cutting date and seed addition treatments in a replicated split-plot design. Grazing treatments consisted of either autumn grazing with cattle and sheep, spring grazing with sheep or both regimes. Fertilizer application treatments consisted of either 25 kg ha^?1 N plus 12–5 kg ha^?1 P and K or no fertilizer. Cutting date treatments consisted of cuts on either 14 June, 2i July or 1 September. Seed addition treatments consisted of either no addition or sowing with a range of meadow species in the autumn. Data analysis was by correspondence analysis and analysis of variance. Species number decreased with fenilizer use and when the cutting date was 1 September. A range of species was affected by the main treatments and there were some first-order interactions, mainly between cutting date and fertilizer application. Rhinanthus minor was particularly favoured by the seed addition treatment. Species attnbutes in the regenerative and established phase were related to treatments and their effect on species composition. The National Vegetation Classification communities were associated with particular treatment regimes. The 21 July cutting date favoured ‘improved’ over ‘unimproved-traditional’ swards, with spring grazing favouring ‘unimproved-traditional’ swards. Lowest yields of herbage biomass were associated with autumn and spring grazing, the 14 June cutting date and no fenilizer treatments. The fenil-izer, 1 September cutting date and auiumn grazing treatments gave the highest yields. The implications of these results are discussed in terms of the conservation management required to return agriculturally improved mesotrophic grassland to a species composition similarto that of traditionally managed grassland.