Tiller traits of spring cereals under tiller-depressing long day conditions

Long days partially depress tiller growth of spring cereals. In this study we characterised and quantified growing conditions and cultivar-induced variation in tiller traits and contribution of tillers to grain yield. Experiments were done at two locations in southern Finland, incorporating two nitrogen fertiliser application rates (80 and 120 kg N ha⁻¹) and 12 two-row barley (Hordeum vulgare L.), 10 six-row barley, ten oat (Avena sativa L.) and 11 wheat (Triticum aestivum L.) cultivars. Spring cereal species differed significantly in their tillering: two-row barley was superior compared with other spring cereals, with the highest number and growth capacity of tillers and head-bearing tillers. Grain yield produced by tillers was, however, always modest compared with that of main shoots, ranging from 13% and 15% in oat and wheat to 20% in six-row barley and 64% in two-row barley. Cultivar and growing condition-induced differences were marked in tiller traits, but tillers were never able to out-yield the main shoot. This is likely to be emphasised by standard use of high seeding rates. Tillers are dominated by the main shoot under long day conditions at high latitudes, and tiller yield potential remains clearly underutilised even under conditions favouring growth.     

Cereal yield trends in northern European conditions: Changes in yield potential and its realisation

In recent decades there has been steady reduction in the rate of increase in cereal yields worldwide. This paper aims to (1) characterise the changes in yield trends of spring (barley, oat and wheat) and winter cereals (rye and wheat) grown in Finland and (2) distinguish between plant breeding achievements and other potential contributors to changes in national yield trends and differences among crops. In this work we used long-term datasets from FAO (FAOSTAT, 2007) (1960–2005) and results from multi-location MTT Official Variety Trials (1970–2005). A mixed model technique was used to divide the yield trends in variety trials into two components: genetic improvements and environmental changes. All trends were characterised using 5 years moving averages. The study period was divided into three agronomically relevant sub-periods: (1) in 1960–1980 agriculture was mechanised and improved basic agricultural practices largely introduced, (2) in 1981–1994 intensified crop management practices were increasingly applied and (3) in 1995–2005 Finland had joined the European Union, which resulted in altered agricultural policies and markets. Plant breeding has successfully increased genetic yield potential of all cereal crops without any indication of reduced rates of improvement. Since 1995 the Finnish national yield trends declined for all crops except wheat, for which the trend levelled-off. The main reasons for this are: cereal production has become less intensive because of (1) application of an environmental programme aimed at increasing the sustainability of agriculture by reducing the environmental load it represents and (2) markedly decreased economic incentives to produce intensively as producer prices for cereals decreased and input prices remained unchanged. National yield trends did not decline because of lack of genetic improvement in yield potential. It is likely that future cereals yield trends will again respond to increasing demand for food, feed, biofuel production and global climate change.     

Grain number dominates grain weight in temperate cereal yield determination: Evidence based on 30 years of multi-location trials

Multi-location data collected over 30 years in Finland on major cereal crops (spring barley, oat and wheat, and winter wheat and rye) provide an opportunity to characterise variation, interrelation and relative importance of the grain yield determining components, grain number per square meter (GNO) and single grain weight (SGW). In addition to evaluating differences among cereal crops in yield determination, changes among different age groups (AG) were compared. Field experiments were carried out in 1970–2001 at 25 locations in Finland. Grain yield was recorded (at 15% moisture) and SGW (mg) and GNO (number m⁻²) determined. Frequencies of different GNO and SGW combinations were analysed for all crops and component means in different yield groups as well as among four AGs. In most cases GNO dominated SGW, representing the major yield component determining grain yield. GNO was highly responsive to favourable growth conditions. The change in GNO was especially high when recurrent, relatively low yield groups were compared, while at higher yields the role of SGW became increasingly important, but did not exceed that of GNO. The degree of GNO domination varied according to crop. Increased grain yield in spring oat and winter rye was associated with higher numbers of set grain, while spring barley and winter wheat responded to yield favouring conditions also through higher SGW. Despite GNO dominated SGW due to its responsiveness to growing conditions, its role in yield improvement was not obvious when the entries (n ≥ 30) were classified into four AGs. Improvements in grain yield derived from AG were associated with crop-specific GNO and/or SGW changes.     

Association of growth dynamics, yield components and seed quality in long-term trials covering rapeseed cultivation history at high latitudes

Brassica oilseed yield trend has declined in Finland by over 20% during the last 15 years. Improved genetic yield potential of turnip rape (B. rapa L.) and oilseed rape (B. napus L.) can be better realised with increased understanding of yield determining processes under northernmost growing conditions. This study aimed at (1) determining the genetic improvements in seed yield, yield components, quality traits and duration of the main growth phases and (2) comparing all these traits in turnip rape and oilseed rape. This work is based on the dataset from long-term Official Variety Trials (1976–2006), covering the entire historically relevant period of rapeseed cultivation in Finland. The results indicated that number of seeds per square metre dominated production of high yields, while single seed weight was not correlated with yield. Over the years, seed yields were produced with very different combinations of seed numbers and weights, differing markedly between the two crops. While high seed numbers were required for production of superior seed yields, single seed weights were not particularly high. Environmental variation markedly affected seed yield, seed number per square metre and duration of flowering compared with its effect on single seed weight. Duration of flowering was, however, negatively associated with seed number. Even though seed weight was largely determined by genotype, no marked plant breeding improvements were identified, contrary to those in number of seeds per square metre, seed yield, oil content and oil yield. For oilseed rape the increase in seed yield (41%) was far higher than for turnip rape (19%) without being associated with a longer seed-filling phase and later ripening, but rather the contrary.