Spatial and temporal variation in crop diversity in agroforestry homegardens of southern Ethiopia

A key assumption in many homegarden studies is that homegardens are ecologically and socio-economically sustainable due to their species diversity. The precise relation between diversity and sustainability is still heavily debated, however. A basic question is how diversity in homegardens can best be characterized in view of the various dimensions of species diversity and their variation in time and space. This paper assesses different types of species diversity in the homegardens of Sidama region of southern Ethiopia. In a survey of crop species in 144 homegardens a total of 78 cultivated crop species (excluding trees) belonging to 10 functional groups were recorded; there were on average 16 crop species and 8 functional groups per farm. Within homegardens, plots differ in species composition and crop diversity. Four types of homegarden systems are distinguished differing in both type and area-share of dominant species, relative orientation at subsistence or cash production and overall crop diversity. The gradual replacement of enset by maize and of coffee by more financially attractive cash crops khat and pineapple causes a decrease in overall crop diversity. Our data demonstrate that it is incorrect to consider homegardens as generic systems with a uniform distribution of species diversity: important within and between homegarden variation exists. Ecological and socio-economic sustainability is not just related to species diversity per se, but rather to more specific features such as presence of keystone species and diversity in functional species groups. Socio-economic sustainability in terms of adjustment to socio-economic change implies dynamics in species diversity.     

Diversity,structure and standing stock of wood in the homegardens of Kerala in peninsular India

A survey was conducted in 17 selectedthaluks (revenue sub-divisions) of Kerala state to elucidate the floristic structure, composition and the extent of similarities and diversities in the composition of homegardens of Kerala, with particular reference to small (below 0.4 ha), medium (0.5 to 2 ha) and large (>2.0 ha) holding size categories. Besides attempts wer also made to characterize the potential of homegardens to supply commercial timber and fuelwood. Two hundred and fifty two farmers were selected through a stratified random process. In addition to gathering general information on crop and livestock production enterprises, all scattered trees and shrubs (15 cm girth at breast height) on the homestead and border trees except palms and rubber were enumerated.There was tremendous variability both in the number of trees and shrubs present and species diversity of the selected homesteads in different provinces. The small, medium and large sized holdings also exhibited profound variability in the number of woody taxa and individuals present. In total, 127 woody species (Girth at Breast Height (GBH) 15 cm) were encountered. The mean number of woody taxa found in the homegardens ranged from 11 for Pathanapuram to 39 in Perinthalmanna. Floristic diversity was higher in the smaller homesteads. It decreased with increasing the size of holdings. Mean Simpson's diversity index for the homesteads ranged from 0.251 (Kochi) to 0.739 (Kottarakkara) suggesting that floristic diversity of homegardens was moderate to low compared to a value over 0.90 for the species-rich evergreen forests of the Western Ghats. The Sorensen's similarity indexes suggested a moderately high degree of similarity for the different tree species encountered in the homesteads of Kerala.No clear cut planting pattern was discernible in the homegardens of Kerala. The homegarden trees and shrubs were either scattered throughout the homestead or on farm boundaries. Farmers tend to prefer timber trees such as ailanthus (highest frequency) and teak besides fruit trees such as mango, jack, cashew and the like. Major homegarden species were represented in all diameter classes. The diameter structure, however, exhibited a slightly skewed (+) distribution pattern, having the highest frequency in the 20–30 cm classes ensuring adequate regeneration status and in that process making homegardens a sustainable as well as dynamic land use system.Standing stock of timber and firewood in the homestead of differentthaluks are presented. Average commercial standing stock of homesteads ranged from 6.6 to 50.8 m³ ha^–1 and fuel wood volume was of the order of 23 to 86 m³ ha^–1. Implicit in the high commercial timber volume and fuel wood volume is that a substantial proportion of the society's wood demands are met from the homesteads. Palms, however, constituted the dominant component of standing commercial timber and fuel wood volumes accounting as much as 63% and 72%, respectively, of the total wood in these categories.     

The Chagga homegardens: a multistoried agroforestry cropping system on Mt. Kilimanjaro (Northern Tanzania)

The homegardens are characterised by an intensive integration of numerous multipurpose trees and shrubs with food crops and animals, simultaneously on the same unit of land.The Chagga are skilled farmers with an intimate knowledge of the crops and their ecological requirements. They have a good idea of functions/uses of the plant species on their farms. The large species diversity provides both subsistence and cash crops. It enables the farmer to keep his management options open and provides insurance against drought, pest and economic risks.     

Dynamics of Homegarden Structure and Function in Kerala, India

Homegardens in Kerala have long been important multi-purpose agroforestry systems that combine ecological and socioeconomical sustainability. However, traditional homegardens are subject to different conversion processes linked to socioeconomic changes. These dynamics were studied in a survey of 30 homegardens. On the basis of a cluster analysis of tree/shrub species density and subsequent further grouping using homegarden size as additional characteristic, six homegarden types were differentiated. These were assessed regarding structural, functional, management and dynamics characteristics. Four development stages of homegardens were found along a gradient from traditional to modern homegardens. Fifty percent of the homegardens still displayed traditional features, whereas 33% incorporated modern practices. The process of modernisation includes a decrease of the tree/shrub diversity, a gradual concentration on a limited number of cash-crop species, an increase of ornamental plants, a gradual homogenization of homegarden structure and an increased use of external inputs. One homegarden was characterised as an adapted traditional homegarden combining multispecies composition and intensive management practices using internal inputs with commercial production. In comparison to modern homegardens, this homegarden type offers an example of an alternative development path in adapting homegardens to changing socioeconomic conditions. The study of structural and functional dynamics of homegardens offers the opportunity to understand the trends in socio-economic sustainability and how these relate to ecological sustainability.     

The Effect of the Surrounding Landscape and Socioeconomic characteristics on Woody Species Diversity in Homegardens of Shewarobit District,Northeast Ethiopia

Although homegardens are most often suggested as the refuges for biodiversity in human-dominated landscapes, how the surrounding landscape and socioeconomic characteristics affect this diversity in the tropics has received little research attention. Hence, this study has examined how these factors affect woody species diversity in homegardens of northeast Ethiopia. Three landscapes which are similar in agroecology were selected and in total 54 households were used for both a survey and a woody species inventory in respondents’ homegardens. The homegardens were stratified based on their locations in relation to crop fields and natural vegetation using satellite images from Google Earth. The variation in Shannon–Wiener diversity index among homegardens and the effect of the socioeconomic factors including household wealth status, homegarden area and the households’ types of uses of woody plants on species diversity were analyzed using one-way ANOVA. Altogether 53 woody plant species belonging to 35 botanical families were identified. Survey results indicated that the woody species diversity was higher in homegardens situated close to crop land as compared with the diversity in homegardens close to natural vegetation. Higher woody species diversity was recorded in homegardens which are larger in area and where the households’ types of uses of woody plants is higher. Moreover, the woody species diversity was found to be higher in homegardens of the high and medium income households when compared with that of poor households. Overall, results suggest that the concurrent ecological and socioeconomic studies are needed to design conservation strategy and policy for plant biodiversity in agricultural landscapes.     

Ecological rationalities of the traditional homegarden system in the Chao Phraya Basin, Thailand

Although the homegarden systems in the tropics are claimed to sustain basic community needs without environmental deterioration, the ecological rationalities behind the harmony between the humans, homegardens, and the environment are not well understood. Four study sites (Sukhothai, Srisatchanalai, Ayudhaya, and Nonthaburi) representing the four Thai eras in the Chao Phraya Basin were selected for studying these rationalities. The size and physical stature of the homegardens, their plant association and community features, physical environmental factors, nutrient and soil fertility parameters, and cultivation practices were studied. The major factor that determines species selection in homegardens is the utilization of the products, while the various practices within the homegardens are determined by such factors as the species, the system, and the environment. All homegardens had four vertical stratifications, with physical structures almost similar to that of dry dipterocarp forest, but with lower height for each layer, lower diversity of plants, and sparser crown layer. The analysis also shows a high possible utilization efficiency for space, light, water and nutrients in the soil in the homegardens. Shannon-Wiener's indices of species diversity of the homegardens were close to those of dipterocarp forest. The homegardens are in-situ reservoirs for biodiversity at genetic-, species-, and ecological levels. There was no complete harvesting from these homegardens. This practice ensured minimal nutrient export from the systems, while high amounts and diversity of litter biomass should contribute to high efficiency of nutrient cycling. Futhermore, phosphorus availability could be better in homegardens. The homegardens had more favorable microenvironment with lower soil and atmospheric temperature and higher relative humidity than outside. There has been no single incident of a pest outbreak at a threatening level. This revised version was published online in June 2006 with corrections to the Cover Date.     

Plant-soil interactions in multistrata agroforestry in the humid tropicsa

Multistrata agroforestry systems with tree crops comprise a variety of land use systems ranging from plantations of coffee, cacao or tea with shade trees to highly diversified homegardens and multi-storey tree gardens. Research on plant-soil interactions has concentrated on the former. Tree crop-based land use systems are more efficient in maintaining soil fertility than annual cropping systems. Certain tree crop plantations have remained productive for many decades, whereas homegardens have existed in the same place for centuries. However, cases of fertility decline under tree crops, including multistrata agroforestry systems, have also been reported, and research on the causal factors (both socioeconomic and biophysical) is needed. Plantation establishment is a critical phase, during which the tree crops require inputs but do not provide economic outputs. In larger plantations, tree crops are often established together with a leguminous cover crop, whereas in smallholder agriculture, the initial association with food crops and short-lived cash crops can have both socioeconomic and biological advantages. Fertilizers applied to, and financed by, such crops can help to `recapitalize' soil fertility and improve the development conditions of the young tree crops. Favorable effects on soil fertility and crop nutrition have been observed in associations of tree crops with N<sub>2</sub>-fixing legume trees, especially under N-deficient conditions. Depending on site conditions, the substitution of legume `service' trees with fast-growing timber trees may lead to problems of competition for nutrients and water, which may be alleviated through appropriate planting designs. The reduction of nutrient leaching and the recycling of subsoil nutrients are ways to increase the availability of nutrients in multistrata systems, and at the same time, reduce negative environmental impacts. These processes are optimized through fuller occupation of the soil volume by roots, allowing a limited amount of competition between associated species. The analysis of temporal and spatial patterns of water and nutrient availability within a system helps to optimize the use of soil resources, e.g., by showing where more plants can be added or fertilizer rates reduced. Important research topics in multistrata agroforestry include plantation establishment, plant arrangement and management for maximum complementarity of resource use in space and time, and the optimization of soil biological processes, such as soil organic matter build-up and the stabilization and improvement of soil structure by roots, fauna and microflora. This revised version was published online in June 2006 with corrections to the Cover Date.     

Interdisciplinary analysis of homegardens in Nicaragua: micro- zonation, plant use and socioeconomic importance

The choice of plant species, their arrangement and management varies between and within tropical homegardens in the same community. Relationships between agroecological and socioeconomic characteristics of 20 homegardens were examined at Masaya, Nicaragua. Variables analyzed were micro-zonation (area allocation to specific uses and management), plant use and diversity, occupation, labor investment, and product, benefit and income generation. Data was collected through surveys, participatory mapping, plant inventories, direct observation and interviews. Ten different micro-zones and nine plant uses were identified. Fruit trees, shaded coffee and ornamentals were the most important zones. Plant diversity was high, with a sample total of 324 species. Homegardens were an important occupation, with average labor investments of 32.6 h family^–1 week^–1. Families obtained at least 40 different plant products from homegardens, as well as the benefit of space for working and socializing. Six homegarden types were identified using a cluster analysis based on biophysical variables. Types reflected the relationship between income generation and the number and types of zones and plant species present. Labor inputs were high considering the small size of the homegardens (average size 3,240 m²), although no clear relationships between labor investment and plant and zone type or number were observed. Homegarden management strategies of plant selection and zonation were affected both by family choices and external forces. Although dependence on homegardens may vary according to specific conditions at a given time, they seem to be a consistent, flexible resource used to meet a diversity of needs. The methodological approach used in this paper may be appropriate for the study of other traditional agroecosystems since it includes both biophysical and socioeconomic variables, essential for understanding these complex systems.     

采伐作业对长白山阔叶红松林林下植被影响的研究(英文)

森林采伐是森林经营中的一个关键环节。采伐对树木天然更新的影响直接关系到森林的结构、组成及森林的可持续经营。本文以长白山阔叶红松林为研究对象,对择伐5 年后不同作业迹地(集材道、楞场和采伐后林窗)的幼苗更新及灌木、草本的多样性进行研究,并以没有进行采伐干扰的林地作为对照。共设计样地23 块,在各个小样地内调查幼树的种类、树木、高度;灌木和草本植物的种类、树木、高度和盖度,应用SPSS 软件进行一元方差分析。结果表明,择伐作业影响幼苗的更新,灌木和草本的多样性在作业迹地增加,尤其在林隙处的多样性最高。适当采伐能促进幼苗的更新,特别是阔叶树种的更新。集材道对幼苗密度的影响比对幼苗高度的影响大,幼苗密度在集材道上密度最高;采伐作业对针叶树种有严重的影响,在三种作业迹地上针叶树种的密度都比对照低。为了维持森林的结构和组成,在森林收获和楞场的建立时,必须采取科学措施保护针叶树种。此外,采伐林窗和楞场的面积都应该减小。为了确保采伐迹地建群树种的更新,经营中应该在保护生物多样性的前提下,适当控制过于旺盛的灌木和草本层的生长。本研究结果将为采伐后作业迹地的恢复和森林的可持续经营提供科学的依据。图1 表4 参14。     

Structure and floristics of Bangladesh homegardens

A vegetation survey of traditional homegardens in four regions (South western, North western, Eastern and Central northern) in Bangladesh was conducted with reference to marginal (> 0.002–0.08 ha), small (> 0.08–0.14 ha), medium (> 0.14–0.20 ha) and large (> 0.20 ha) farm categories. Eighty homegardens (five from each farm category in each of the four regions) were assessed in terms of perennial species. The floristic data were ordinated (DECORANA) with respect to homegarden categories based on size and region. Most species were planted in the border of the homegardens irrespective of farm size and region. Food and fruit producing species dominated near the living quarter and working areas and small plots of annual vegetables and crops separated this part of the garden from more distant parts favoured for timber species. Six vertical strata were recognised with higher plant density and species richness recorded in the lower three. In total ninety two perennial species were recorded for the set of 80 homegardens surveyed. From gardens in the South western region 67 species were recorded. Corresponding figures were 56 for the Central northern region, 54 for the Eastern region and 46 for the North western region. Within regions there were significant differences in species richness associated with farm size, and within each homegarden size category there were significant differences among regions. Diversity was highest among food and fruit producing species, followed by the timber species. The ordination showed a distinction between the North western and the other regions due to a combination of lower species richness in the North western region homegardens and several species exclusive to the region. Floristic differences led to less marked but nevertheless important differences among the other regions, also.     

Successional process of Picea crassifolia forest after logging disturbance in semiarid mountains: A case study in the Qilian Mountains,northwestern China

Selective logging is the most widely employed method of commercial timber production in Asia, and its impact on forest structure, composition, and regeneration dynamics is considerable. However, the successional processes in forest communities after logging in semiarid mountains are poorly understood. To provide more information on these processes, we used data from tree rings, direct and indirect age determinations, and field measurements of stand structure to reconstruct the historical disturbance regime, stand development patterns, and successional processes in a natural Picea crassifolia forest community in the Qilian Mountains of northwestern China. The results showed that the density of P. crassifolia forest increased significantly after logging. The densities of second growth forests 30 and 70 years after logging disturbance had increased to 2874% and 294% of primary forest's density, respectively. Logging disturbance did not alter tree species composition of logged stands. However, the diversity of understory species changed significantly among the successional phases. Logging disturbance decreased the spatial heterogeneity of second growth forest. The spatial distributions of recruitment were affected by the location of the remaining trees. There was less recruitment near the remaining trees than near forest that had been cut. In addition, logging disturbance also induced a growth release for the trees on the sites sampled. Our results imply that the succession and regeneration of P. crassifolia forest may be improved if the remaining trees could be retained relative uniform distribution pattern, thinning or selective logging could be performed to height density, exotic shrubs could be removed or the shrubs cover could be reduced during the earlier successional stages.     

Agroforestry systems in the highlands of the Tehuacán Valley,Mexico: indigenous cultures and biodiversity conservation

In this study we analysed: (1) the biodiversity conservation capacity of Agroforestry Systems (AFS) in temperate highlands of the Tehuacán–Cuicatlán Valley, Central Mexico, (2) human cultural motives and actions for conserving such diversity and (3) problems endangering that capacity. We evaluated the richness and diversity of perennial plant species maintained in AFS through vegetation sampling of 14 agricultural plots and compared their composition with that of natural forests (14 plots of 500 m² each). We examined the situations among communities of Náhuatl, Ixcatec and Cuicatec people, documenting through interviews the management practices of plant species and the whole system, reasons why people maintain vegetation cover within AFS, and factors influencing changes in decisions favouring agriculture intensification. In the AFS studied we recorded a total of 79 species of trees and shrubs, 86 % of them being native species and representing 43 % of all species of trees and shrubs recorded in the sampling of the natural forests the AFS derive from. People leave standing on average a total of 40 individual trees and shrubs per agricultural plot. Reasons for leave plant species standing were more frequently associated with their use as fruit trees, firewood, shade, beauty, respect to nature and other environmental benefits. Water availability for irrigation, land tenure, and dependence on agriculture and forest for peasant's subsistence were main decision factors influencing AFS variation in their composition. AFS in temperate zones are important reservoirs of biodiversity and biocultural heritage and should be keystones for conservation policies in the Tehuacán–Cuicatlán Valley.     

Woody plant diversity,structure and regeneration in the Ambo State Forest,South Gondar Zone,Northwest Ethiopia

For this study in the Ambo State Forest on woody plant diversity, structure and regeneration, 70 quadrats, each 25 m by 25 m, were selected using a systematic random sampling technique and intervals of 100 m along a transect line. For assessing seedlings and saplings,two 2 × 10 m sub quadrats were set upon opposite sides of each main quadrat. Data on species diversity, abundance,structure, basal area, density, frequency and regeneration status were collected and analyzed using standard procedures and programs. Of 58 woody plant species identified,69 % were trees, 16 % were shrubs, 12 % were tree/shrubs and 4 % were climbers. Fabaceae was the most speciesrich family comprising 17 species. The Shannon–Weiner diversity index was 2.73, and evenness was 0.67. The population structure in the cumulative diameter class frequency distribution revealed an interrupted and inverted J-shape with a very high decrease in higher diameter class.Acacia lahai(49 %) was the most important woody species with the highest importance value index. To maintain balanced structure, enhanced regeneration and protecting the forest from selective cutting are recommended.     

Exploring Agrobiodiversity on Farm: A Case from Middle–Hills of Nepal

A survey was conducted to examine agrobiodiversity status of farms in the Pokhare Khola watershed of Dhading district in the Middle-Hills of Nepal. A total of 53 farmland tree and one crop species from 22 families were documented. The most commonly found species were fruit and fodder species (on over 60 % farms) including banana (Musa paradisiaca), khasrato (Ficus hispida) and khanayo (Ficus semicordata). Tree density was highest (226/ha) on marginal farms (farm area ≤0.25 ha) and lowest (165/ha) on small farms (farm area 0.26–0.5 ha). For the study area as a whole, the Shannon-Wiener species diversity index was 3.26 and the species evenness index 1.89. Large farms (farm area >1 ha) had the greatest tree species diversity (4.47 ± 0.52) and marginal farms the lowest (2.18 ± 0.37), indicating the positive relationship between farm size and species diversity. A total of six types of cereals and 18 types of vegetable crops were grown in the study area. The major livestock component of each household was chickens (average 8/household) and goats (6/household). The mean value of livestock in the large farm category was estimated as $2235, significantly higher than that of the other three categories. A significant relationship was found between agrobiodiversity and livelihoods, irrespective of annual production, and the critical role of the farmed landscape in agrobiodiversity conservation was apparent.     

Diversity and structural composition of species in dipterocarp forests: a study from Fasiakhali Wildlife Sanctuary,Bangladesh

Fasiakhali Wildlife Sanctuary is a protected area composed of tropical remnant rainforest that harbor substantial number of large,old Garjan(Dipterocarpus spp.)trees.The present study assessed composition,structure and diversity of the species in this protected area.A total of 32 trees species were recorded with DBH ≥ 11 cm belonging to 24 genera and 19 families.The forest is low in plant diversity as represented by Shannon–Wiener diversity and Simpson Dominance indices.Dipterocarpus turbinatus was the most dominant species with maximum relative density,frequency,dominance,and importance value index.Syzygium firmum and Tectona grandis followed in terms of dominance.The structural composition indicated higher number of individuals in the medium growth classes(41 to 511 cm DBH and 16–20 m height ranges),whereas D.turbinatus was the only species that dominated most of the growth classes.Poor stem density in lower growth classes indicated meager recruitment of regeneration which may be due to lower annual precipitation,increased grazing and encroachments.This study will help to understand the patterns of tree species composition and diversity in the remnant dipterocarp forests of Bangladesh.It will also contribute to identifying threatened plants to undertake D.turbinatus based conservation and sustainable management of the Fasiakhali Wildlife Sanctuary.     

Phenology and pollination of Manilkara zapota in forest and homegardens

Knowledge on reproductive phenology and pollination biology are basic elements that should be considered in the management and exploitation of plant species that offer non-timber products. The tropical tree Manilkara zapota is a species from which non-timber products have been obtained for centuries by Mayan communities in Mexico. Nevertheless, there are no quantitative reports on its reproductive biology and the factors that limit fruit production. The present study describes the reproductive phenology, breeding system and pollination of this species in two contrasting environments: medium-height, subdeciduous forest, and homegardens (“solares”) in a Mayan community in the state of Yucatan. Significant differences were found between environments both in the temporal distribution of flower and mature fruit production, as well as in the proportion of mature fruits. Homegarden trees showed the greatest fruit production, although flower production did not differ between environments. Mature fruits were of better quality (i.e., greater fresh weight) in homegardens. Hand pollination experiments showed that M. zapota is self-compatible, and that there is pollinator limitation for fruit production in trees that grow in homegardens. We propose that water and soil nutrients are the main factors limiting M. zapota fruit production in forests, while in homegardens the main factor appears to be pollinator availability.     

Integration of indigenous wild woody perennial edible fruit bearing species in the agricultural landscapes of Amhara region,Ethiopia

The study was undertaken in three districts of Amhara region to assess the extent and purpose of integration of wild indigenous woody perennial fruit bearing species in the agricultural landscapes, appraise their species composition and diversity and identify factors holding back farm integration. Data were gathered through interviews administered to 90 randomly chosen household heads, and fruit trees census in each and every plot of the informants. Species diversity was found to be low and vary by farm, land use type and site. Altogether 17 species were recorded in the agricultural settings. Species retention in the different land use types appears to be governed by species relative importance and compatibility with annual crops, where farm edges recorded a higher ethno-ecological importance score. Nevertheless, as it stands now fruit bearing species are retained primarily for non-fruit utilities. Free availability, land shortage, slope and altitude as well as disgrace feelings are among the major factors holding back farm integration and diversity of indigenous species for fruit production. Should they are additionally exploited for their fruits proper, there is a critical need to raise recognition and awareness at the rural grassroots level and instill knowledge about the food values, and assist in appropriate tree management techniques and inter-cropping regimes as well as accessing markets.     

A 6-year longitudinal study on agrobiodiversity change in homegardens in Tabasco,México

Societal processes of rural change and globalization may change homegardens and their contribution to the conservation of agrobiodiversity, particularly of species occurring naturally in regional vegetation. The best way to determine if this occurs is through longitudinal studies. We conducted such a study, inventorying tree species in a sample of 38 homegardens in 2009, 2012 and 2015. The homegardens were located in the subregions of mountain slopes, fluvial plains and coastal plains in the tropical lowlands of Tabasco, Mexico. We analysed changes in species richness by geographic origin, species richness and species composition in each inventory. We identified 169 tree species in the three inventories, of which 74.6% were native or neotropical and 25.4% introduced. Of the 140 species recorded in 2009, 88% remained in 2015, whereas 12% had been replaced and nine additional species had arrived. Mean species richness increased between 2009 and 2015 (P = 0.03) and between 2012 and 2015 (P = 0.001). Increases resulted from increased mean neotropical (P = 0.01) and introduced (P = 0.01) species richness, and constant native species richness. Differences in species composition between the three subregions in 2009 persisted in 2012 and 2015 (P < 0.001 in all years). These results show how the highly dynamic character of homegardens combines with the renewal and persistence of their agrobiodiversity, and underpins the continued relevance of homegarden for agrobiodiversity conservation and livelihoods in tropical lowlands amidst rural change and globalization.     

Indigenous agroforestry of Pohnpei

Plant species and cultivars of the indigenous agroforestry system of Pohnpei were surveyed in transects through 54 randomly-selected farms. The agroforestry system was characterized by extensive cultivation of yams (Dioscorea), aroids (Alocasia), and Piper methysticum under a permanent overstory of breadfruit, coconut, and forest remnant trees and a middle canopy of Hibiscus tiliaceus, Musa spp., and Morinda citrifolia. In the 10 ha of survey plots, 161 species were found, of which 102 were trees, shrubs, and crops and 59 were uncultivated herbaceous plants. Numbers of tree, shrub, and crop species per farm ranged from 16 to 37 with an average of 26. Twenty-eight breadfruit and 38 yam cultivars were found in the survey plots, showing that cultivar diversity is an important component of the biological diversity maintained and utilized in Pohnpeian agroforests.