A non-linear hierarchical mixed model to describe tree height growth

A non-linear hierarchical mixed model approach is used to describe height growth of Norway spruce from longitudinal measurements. The parameter variation in the model was divided into unknown random effects, fixed effects and covariate-dependent effects in order to model tree height growth. The values for fixed effect parameters and the variance–covariance matrix of random effects were estimated. Covariates could only explain up to 10% of parameter variability. Height curves were calibrated by means of BLUPs for the unknown random effects using prior height measurements and evaluated using a separate dataset. The resulting curves had a small error variance and plausible shapes.     

长白山北坡林线分布趋势的研究(英文)

为了揭示高山树线随着气候变化而变化的现象，对长白山的高山树线变化趋势进行了研究。根据长白山北坡地区的气象数据，重点调查了（对树线有最大限制作用）一月平均气温和年积温。并在岳桦林与高山苔原之间的交叉带内测出岳桦的地径及样方内岳桦株数。根据地径与年龄有线性关系，地径可以用来代替年龄。结果表明岳桦林年龄随着海拔的上升而降低。近些年长白山北坡的气温在渐渐上升，因此导致了高山树线的上升。     

Light intensity and hydrogel soil amendment differentially affect growth and photosynthesis of successional tree species

Global climate changes have increased temperatures,radiation indexes,and consequently,irregularities in rainfall in mainly tropical countries,considerably hindering plant establishment in recovering degraded areas.The objective of this study was to evaluate the growth and physiological characteristics of one species of each successional group:pioneer,secondary,and climax when subjected to different light intensities and hydrogel as a soil conditioner during rainy and dry periods.The experiment w...     

The effect of different atmospheric ozone partial pressures on photosynthesis and growth of nine fruit and nut tree species

Nursery stock of peach (Prunus persica L. Batsch, cv. O'Henry), nectarine (P. persica L. Batsch, cv. Fantasia), plum (P. salicina Lindel., cv. Casselman), apricot (P. armeniaca L., cv. Tilton), almond (P. dulcis Mill., cv. Nonpareil), prune (P. domestica L., cv. Improved French), cherry (P. avium L., cv. Bing), oriental pear (Pyrus pyrifolia Rehd., cv. 20th Century), and apple (Malus pumula Mill., cv. Granny Smith) were planted in open-top chambers on April 1, 1988 at the University of California's Kearney Agricultural Center located in the San Joaquin Valley (30 degrees 40' N 119 degrees 40' W). Trees were exposed to three atmospheric ozone partial pressures (charcoal-filtered air (C), ambient air (A), or ambient air + ozone (T)) from August 1 to November 17, 1988. The mean 12-h (0800 to 2000 h) ozone partial pressures measured in open-top chambers during the experimental period were 0.030, 0.051, and 0.117 microPa Pa(-1) in the C, A and T treatments, respectively. Leaf net CO(2) assimilation rate decreased linearly with increasing 12-h mean ozone partial pressure for the almond, plum, apricot, prune, pear, and apple cultivars. Stomatal conductances of apricot, apple, almond, and plum decreased linearly with increasing ozone partial pressure. Cross-sectional area relative growth rates of almond, plum, apricot, and pear decreased linearly with increasing ozone partial pressure. Net CO(2) assimilation rate, stomatal conductance, and trunk growth of cherry, peach and nectarine were unaffected by the ozone treatments. Reduced leaf gas exchange probably contributed to ozone-induced growth reduction of the susceptible species and cultivars. Several of the commercial fruit tree species and cultivars studied were relatively tolerant to the ozone treatments.     

Effects of understorey vegetation on tree establishment and growth in a silvopastoral system in northern Greece

The interaction between understorey plants and trees in a young silvopastoral system was studied in a sub-humid Mediterranean environment with cold winters in northern Greece. The experimental design was a split-split plot with three replications and included: three understorey treatments (grass, legume and control), two tree species (Acer pseudoplatanus and Pinus sylvestris) and two spacings (2.5×2.5 m and 3.5×3.5 m). The competition between herbaceous plants and trees resulted in significant differentiation in tree growth while their spacing did not produce any significant differences. The influence of trees on herbage yield either by plant species or spacing was not significant. During the early stages of establishment, a significant positive correlation was observed between sycamore growth and soil moisture, leaf weight, leaf area and leaf number as well as nitrogen and potassium concentration in leaves. Six years after planting the height increase of the sycamore trees was largest in the control treatment (415%), lowest in the grass treatment (134%) and intermediate (192%) in the legume treatment. Much higher was the increase obtained for the diameter, 161%, 207% and 536% respectively for the grass, legume and control treatments. The Scots pine trees grew faster than sycamore over the course of the experiment. Height increased by 397%, 351% and 400% and diameter by 518%, 443% and 683% respectively for the grass, legume and control treatments. This revised version was published online in June 2006 with corrections to the Cover Date.     

Soil N mineralization profiles of co-existing woody vegetation islands at the alpine tree line

Tree lines form a transition ecotone from forest to tundra both at high elevation and high latitude and occur in a number of different forms. Nitrogen (N) deficiency is considered to be a factor involved in tree line formation, and also N dynamics are considered to differ between the trees and the ericaceous vegetation of the tundra. In the Austrian Alps at the tree line, N availability and N mineralization in soils of different vegetation types (Picea abies, Pinus mugo and Rhododendron ferrugineum) as well as total phenols were determined. Soil from under P. abies was taken from two different tree line forms, an island type and a diffuse type, as well as from P. abies growing at a lower elevation. N mineralization was measured in situ using a covered PVC tube incubation method and in a laboratory incubation under controlled conditions. Ion exchange resin capsules were installed at the interface of humus and mineral soil for estimating N in the soil solution. Net N mineralization showed a similar pattern for the vegetation types for both the in situ and laboratory incubation. The soil humus layer had the highest levels of N mineralization compared to the other soil layers. N mineralization rates were similar in P. abies and P. mugo at the tree line regardless of tree line form. Rates of N mineralization were lower under R. ferrugineum than the tree species, but this lower rate was not related to the occurrence of high levels of total phenols in the soil. Nitrogen deficiency was not evident in the island-type tree line, but was evident in the diffuse tree line type.     

Early growth of six native Australian tree species in windbreaks and their effect on potato growth in tropical northern Australia

A study to examine the early growth of a number of tree species planted in windbreaks and to quantify the benefit of two 30-month-old windbreaks (one with two rows of trees and one with three rows of trees) to the production of potato (Solanum tuberosum L.) was conducted on a farm land in tropical north-eastern Australia. Three Eucalyptus (E. microcorys F. Muell, E. tessellaris F. Muell and E. torelliana F. Muell), two Callistemon (C. salignus Smith and C. viminallis Smith) and two Melaleuca species (M. armillaris Smith and M. linariifolia Smith) were used to form the windbreaks. Tree performance in the windbreaks was evaluated based on the measurements of height, height to the lowest branch, crown length along the row, and number of branches per plant. Potato vegetative growth including plant height, crown size, and leaf length, width (measured at the widest part of the leaf), and number per plant, was measured at ages 2, 4, 6 and 8 weeks.E. microcorys and E. torelliana, showed rapid height and branch growth while retaining low branches and are considered highly suitable for windbreaks. The porosity of the three row and two row windbreaks were 37.2% and 60% respectively at age 30 months. The windbreaks increased potato plant growth in height and leaf number with vegetative plants being tallest at 3 h from the windbreaks. However, the windbreaks had limited effects on leaf length and width. It was found that potato plants grown close to windbreaks yielded more potatoes than those grown at the furthest positions, with the highest production at 3 h. Overall, the windbreaks studied increased potato yield by up to 7.7%.     

燕山北部山地干旱阳坡不同造林树种生长状况的比较

为了了解燕山山地干旱阳坡人工造林树种生长状况的差异,对人工栽植的油松、樟子松及蒙古栎的生长状况及其幼林的空间分布格局进行了研究,结果表明:栽植5a后,油松、樟子松和蒙古栎的存活率分别达到了93.79%、87.98%和66.69%,其地径分别为4.68cm、5.34cm、1.52cm,树高分别为1.58m、1.75m、0.80m,油松、樟子松、蒙古栎的单株生物量分别为1 857.36g、2 018.30g、234.35g,形成的人工幼林的密度在867.00~2 183.00株/hm~2之间;立地条件较好的样地幼林的空间分布格局为均匀型,立地条件较差的样地空间分布格局为聚集型。油松、樟子松、蒙古栎均为燕山北部山地干旱阳坡的适宜造林树种,栽植5a后已基本形成稳定的乔木层,人工造林加速了干旱阳坡由绣线菊灌丛向乔木林的更新与恢复。     

Effects of calcium on seed germination, seedling growth and photosynthesis of six forest tree species under simulated acid rain

We selected six tree species, Pinus massoniana Lamb., Cryptomeria fortunei Hooibr. ex Otto et Dietr., Cunninghamia lanceolata (Lamb.) Hook., Liquidambar formosana Hance, Pinus armandii Franch. and Castanopsis chinensis Hance, which are widely distributed as dominant species in the forest of southern China where acid deposition is becoming more and more serious in recent years. We investigated the effects and potential interactions between simulated acid rain (SiAR) and three calcium (Ca) levels on seed germination, radicle length, seedling growth, chlorophyll content, photosynthesis and Ca content in leaves of these six species. We found that the six species showed different responses to SiAR and different Ca levels. Pinus armandii and C. chinensis were very tolerant to SiAR, whereas the others were more sensitive. The results of significant SiAR?×?Ca interactions on different physiological parameters of the six species demonstrate that additional Ca had a dramatic rescue effect on the seed germination and seedling growth for the sensitive species under SiAR. Altogether, we conclude that the negative effects of SiAR on seed germination, seedling growth and photosynthesis of the four sensitive species could be ameliorated by Ca addition. In contrast, the physiological processes of the two tolerant species were much less affected by both SiAR and Ca treatments. This conclusion implies that the degree of forest decline caused by long-term acid deposition may be attributed not only to the sensitivity of tree species to acid deposition, but also to the Ca level in the soil.     

Effects of growth rates, tree morphology and site conditions on longevity of Norway spruce in the northern Swiss Alps

Longevity of trees is known to be associated with growth rates, but also with tree morphology and spatial influences. However, very little quantitative information is available on the effects of these biotic and abiotic influences on maximum ages of trees. The objectives of this study were to investigate the trade-off between longevity and growth rates of Norway spruce (Picea abies) and to quantify the effects of tree morphology and abiotic site conditions on longevity of this species. Data were collected along different topographical and climatic gradients in a 20?×?25?km study area in the northern part of the Swiss Alps (Glarus). The ages of the more than 100 sampled dead Norway spruces ranged between 50 and 367?years. Longevity of these trees was negatively related to tree growth, i.e. slow-growing trees tended to grow older than fast-growing trees. Tree height was positively associated with longevity for both upper and lower storey trees. Longevity of lower storey trees was increased with large crown diameter, but decreased with long crown length. Upper storey trees growing at higher altitude tended to get older than at lower altitude. We conclude that the combined effects of growth rates, variability in site conditions and different traits of tree morphology determine tree longevity of Norway spruce in the Swiss Alps. Because longevity is tightly linked to mortality rates of tree populations, our study may improve our understanding of long-term processes of forest dynamics under current and future climate.     

Bridging process-based and empirical approaches to modeling tree growth

The gulf between process-based and empirical approaches to modeling tree growth may be bridged, in part, by the use of a common model. To this end, we have formulated a process-based model of tree growth that can be fitted and applied in an empirical mode. The growth model is grounded in pipe model theory and an optimal control model of crown development. Together, the pipe model and the optimal control model provide a framework for expressing the components of tree biomass in terms of three standard inventory variables: tree height, crown height and stem cross-sectional area. Growth rates of the inventory variables and the components of biomass are formulated from a carbon balance. Fundamentally, the parameters of the model comprise physiological rates and morphological ratios. In principle, the values of these parameters may be estimated by lower-level process models. Alternatively, the physiological and morphological parameters combine, under reasonable assumptions, into a set of aggregate parameters, whose values can be estimated from inventory data with a statistical fitting procedure.     

西南石漠化地区两种经济林木光合日动态特征

为给我国石漠化地区经济林的栽培和生产提供科学依据,以‘香玲’核桃和‘贵农5号’刺梨为材料,利用光合测定仪对西南石漠化治理区的核桃和刺梨的光合日变化特征进行了测定和分析。结果表明:‘香玲’核桃的日均净光合速率5.57μmol/(m2·s)、蒸腾速率3.70mmol/(m2·s)、水分利用效率1.94μmol/mmol;‘贵农5号’刺梨的日均净光合速率7.65μmol/(m2·s)、蒸腾速率4.48mmol/(m2·s)、水分利用效率1.84μmol/mmol。核桃表现出低光合效率、低蒸腾效率;刺梨表现出高光合效率、高蒸腾效率的特点。石漠化地区由于特殊地质环境,在栽植经济林时要充分考虑树种的净光合速率、蒸腾速率和水分利用效率,较高的净光合速率和较低的蒸腾速率可以充分利用喀斯特稀缺的水分资源,同时产生更多的物质累积,提高潜在的生态经济效益。利用核桃与刺梨的光合效率特性差异,进行乔(核桃)-灌(刺梨)结合,可以增加地表覆盖度,提高光能利用率,减少土壤侵蚀。     

Use of a single-tree simulation model to predict effects of ozone and drought on growth of a white fir tree

A physiologically based, single-tree simulation model, TREGRO, was parameterized with existing phenological, allometric, and growth data and used to predict effects of ozone and drought on growth of a 53-year-old white fir (Abies concolor (Gord. & Glend.) Lindl. ex Hildebr.) tree following a 3-year model simulation. Multiple experimental simulations were conducted to assess the individual and interactive effects of ozone (O(3)) exposure and drought on growth of white fir. The effects of O(3) were imposed as reductions in carbon (C) assimilation of 0, 2.5, 5, 10, and 20%. Drought was imposed as 0, 10, 25, and 50% reductions in total annual precipitation. The results of the simulations were compared with the effects of O(3) on white fir seedlings grown in the presence and absence of ozone in open-top chambers and with a field survey of white fir trees subjected to a gradient of O(3). In the O(3) simulations, an O(3)-induced reduction in C assimilation of 2.5% reduced total tree biomass and branch total nonstructural carbohydrate (TNC) content by < 7%. Although quantifiable in simulation experiments, such small reductions would probably not be detectable in the field. Results from both an open-top chamber experiment and a field survey indicated that reductions in C assimilation of white fir growing in elevated O(3) were much greater than 2.5%, but were not statistically different from control values. A simulated O(3) reduction in C assimilation of >/= 10% reduced total tree biomass by 7% and branch TNC by 55%. Results from the field survey indicated that branch elongation was reduced in response to increased O(3) concentration, corroborating the simulated response of reduced C allocation to the branches of white fir. Although simulated reductions in total annual precipitation of >/= 25% reduced final tree biomass, the simulated reductions also reduced O(3) uptake and therefore reduced the O(3) response of white fir. However, a combination of low amounts of O(3) (2.5% reduction in C assimilation) and drought (25% reduction in annual precipitation) synergistically reduced C gain of white fir more than either stress individually. Our simulations predict that moderate drought (no more than a 25% reduction in total annual precipitation) may not ameliorate the response of white fir to O(3) and that moderate amounts of atmospheric O(3) and drought could be more detrimental to white fir than either stress singly.     

Patterns of photosynthesis and starch allocation in seedlings of four bottomland hardwood tree species subjected to flooding

Effects of short-term (32 days) flooding on photosynthesis, stomatal conductance, relative growth rate and tissue starch concentrations of flood-intolerant Quercus alba L. (white oak), bottomland Quercus nigra L. (water oak), bottomland Fraxinus pennsylvanica Marshall. (green ash) and flood-tolerant Nyssa aquatica L. (water tupelo) seedlings were studied under controlled conditions. Net photosynthetic rates of flooded N. aquatica seedlings were reduced by 25% throughout the 32-day flooding period. Net photosynthetic rates of flooded Q. alba seedlings fell rapidly to 25% of those of the control seedlings by Day 4 of the flooding treatment and to 5% by Day 16. In F. pennsylvanica and Q. nigra, net photosynthetic rates were reduced to 50% of control values by Day 8 but remained at approximately 30 and 23%, respectively, of control values by Day 32. Leaves of flooded Q. alba seedlings accumulated approximately twice as much starch as leaves of non-flooded control plants, whereas root starch concentrations decreased to 67% of those of control plants by the end of the 32-day flooding treatment. In contrast, flooding caused only a small increase in leaf starch concentrations of N. aquatica plants, but it increased root starch concentrations to 119% of those of the control plants by the end of the experiment. The co-occurring bottomland species, Fraxinus pennsylvanica and Q. nigra, differed from each other in their patterns of stomatal conductance and root starch concentrations. We conclude that the maintenance of low leaf starch concentrations, and high pre-flood root tissue starch concentrations are important characteristics allowing flood-tolerant species to survive in flooded soils.     

Variation in tree diameter growth in response to the weather conditions and tree size in deciduous broad-leaved trees

The diameter growth of trees may vary depending on the tree size, species and other tree-related factors. However, such variation is challenging to consider while estimating the growth response to weather conditions. This study investigated the effects of the weather conditions and variations in tree factors on diameter growth in a mature forest. In particular, we focused on diameter growth in June, the month of active diameter growth. We recorded the monthly diameter growth of 6 tree species for 7 years and constructed a generalized linear mixed model in which the mean growth was dependant on tree size, weather conditions, and the interaction between tree size and weather.     

Application of Grey System Theory to tree growth prediction

IntroductionDahurian larch plantations were planted affer the1950's in Heilongjiang Province. Some of them areless than 40 years old. There lack permanent plots oflong-term. Sometimes, statistical models for predicting tree growth are not always desirable. The GreySystem is a new branch of mathematics founded byProf. Deng Julong in 1982 (Deng 1985). In contrast tostatistic models, which need a lot of measured data,the Grey System only need a f6w data for setting upmodels. In recent years…     

Dynamic response of tree growth to changing environmental pollution

Long-term investigations of Scots pine (Pinus sylvestris L.) growth were carried out in the vicinity of one of the biggest air pollution sources in Lithuania—mineral fertilizers plant “Achema.” It is detected that initial stages of the dynamic tree response to the external stress factors in the polluted environment with an increased quantity of nitrogen compounds have started with a stimulation stage, followed by a gradual transition to a depression of growth. The recovery of the damaged stands took place along with the reduced environmental pollution, and the overdraft of the “normal” annual increment was a characteristic feature of all the investigated stands. This phase is still continuing for the most damaged stands. A very high individual variability of the tree growth response to the environmental impact is a characteristic feature of the damaged Scots pine stands. The homeostatic mechanisms of the survived trees stipulated reaching approximately the same or even higher growth rate as prior to the depression period, and the tree growth rate before the depression period can serve as the most powerful predictor of tree growth recovery capacity under the reduced environmental pollution. Crown defoliation is the next most important predictor of individual tree recovery capacity. Lower stand density and lower competition cause higher recovery capacity of damaged trees. The conclusion is made that a reduction in the environmental pollution on the local and regional scale and especially a decrease in emissions and deposition of sulfur and nitrogen compounds caused the recovery of damaged forests. These trends should be taken into account while analyzing and modeling forest dynamics. Interaction of environmental pollution and climatic factors is very important for the response of tree growth to the environmental stressors.     

土壤pH对蓝莓生长及光合作用的影响

以南高丛蓝莓薄雾(Misty)为试验材料,用H2SO4、Al2(SO4)3和NaOH溶液浇淋(处理)盆栽蓝莓的土壤,以调节土壤pH值,测定蓝莓叶片的叶绿素含量、光合作用及生长指标。结果表明:随着土壤pH值降低,蓝莓叶片叶绿素含量,最大净光合速率(Pmax)、表观量子效率(AQY)、光饱和点(LSP)和光补偿点(LCP)均升高。其中以pH 4.5的处理蓝莓叶绿素含量最高,植株生长势最好;pH 8.5的处理光合强度与生长势均最低。因此,南高丛薄雾蓝莓适合栽培的酸性土壤p H值范围为4.5~5.5,而碱性土壤则抑制其生长。     

Tree growth as indicator of tree vitality and of tree reaction to environmental stress: a review

The intensive monitoring plots (Level II) of ICP Forests serve to examine the effects of air pollution and other stress factors on forest condition, including tree vitality. However, tree vitality cannot be measured directly. Indicators, such as tree growth or crown transparency, may instead be used. Tree growth processes can be ranked by order of importance in foliage growth, root growth, bud growth, storage tissue growth, stem growth, growth of defence compounds and reproductive growth. Under stress photosynthesis is reduced and carbon allocation is altered. Stem growth may be reduced early on as it is not directly vital to the tree. Actual growth must be compared against a reference growth, such as the growth of trees without the presumed stress, the growth of presumed healthy trees, the growth in a presumed stress-free period or the expected growth derived from models. Several examples from intensive monitoring plots in Switzerland illustrate how tree-growth reactions to environmental stresses may serve as vitality indicator. Crown transparency and growth can complement each other. For example, defoliation by insects becomes first visible in crown transparency while stem growth reaction occurs with delay. On the other hand, extreme summer drought as observed in large parts of Europe in 2003 affects stem growth almost immediately, while foliage reduction becomes only visible months later. Residuals of tree growth models may also serve as indicators of changed environmental conditions. Certain stresses, such as drought or insect defoliation cause immediate reactions and are not detectable in five-year growth intervals. Therefore, annual or inter-annual stem growth should be assessed in long-term monitoring plots. An erratum to this article can be found at