雷州半岛尾叶桉人工林耗水量研究

利用热脉冲技术对雷州半岛2种主要土壤类型下的4年生尾叶桉人工林林分耗水量进行了为期1 a的观测,并对林分耗水与林分生长特性、气象因子及4 m深土壤有效水关系进行研究.研究结果表明:尾叶桉人工林边材面积与胸径具有显著的相关性,利用边材面积进行单株向林分尺度耗水的转换可行;河头、纪家尾叶桉人工林日平均茎流密度分别为2 772、1 839 L·m-2·d-1;日平均耗水量分别为1.49、1.53 mm;河头与纪家尾叶桉人工林林分的年耗水量分别为542、559 mm,分别占年降水量的35%、36%;开阔的林冠及较小的叶面积指数、旱季时土壤有效水的减少及较低的饱和蒸汽压差等在不同程度上都限制了桉树的耗水量.     

雷州半岛尾叶桉人工林树液茎流特征的研究

应用热脉冲法对雷州半岛4年生尾叶桉单株树干液流时空的动态变化及与各环境因子的关系进行观测研究.研究结果表明(1)形成层以内木质部不同深度的茎流密度不同,其中靠近形成层部分液流速度较快,但各层都具有相同的日变化趋势,中午12点至1点达到最大;夜间因根压作用影响各层仍有微弱上升液流;各深度日平均茎流密度11.6 L·m-2·d-1,最大为15.3 L·m-2·d-1(晴天),最小仅为5.4 L·m-2·d-1(雨天);(2) 不同直径尾叶桉的日茎流密度具有相似日变化趋势,胸径与高度相近其茎流密度相同,大树较小树快,这主要与树木根系吸收土壤水分的能力有关;(3)热脉冲法与整树容器法对2年生幼树耗水量的同步测定结果相一致,误差仅为3.4%;(4)树干茎流量与饱和蒸气压差和太阳辐射相关性特别显著.     

雷州半岛尾巨桉生长固碳及蒸腾耗水特征

以2年生尾巨桉为研究对象,对其年内生长固碳和蒸腾耗水特征进行了研究。结果表明：2年生尾巨桉年内平均生长固碳速率为0.022 kg·d-1,1 a生长固碳量为7.9 kg;2年生尾巨桉单株年内平均蒸腾耗水3.67 L·d-1,其林分(株行距2 m ×3 m)1 a蒸腾耗水量为221.8 mm。2年生尾巨桉年内生长固碳水分利用效率平均为7.04 g·kg-1,变化范围为2.21~16.27 g·kg-1,旱季明显高于雨季。     

华北落叶松树干液流的个体差异和林分蒸腾估计的尺度上推

2005年6-10月在宁夏六盘山南侧的西峡林场,选择比较均匀的坡面(坡度45°),布设了20m×20m的华北落叶松固定标准地,应用热扩散茎流计连续测定13株树木的树干液流.结果表明:不同径级树木的树干日液流量存在较大差异,在6-7月,其值变化在11.17～24.46kg·d-1,变异系数CV为0.298(n=5);在8-10月,其值变化在5.01～22.25 kg·d-1,变异系数CV为0.454(n=13).方差分析表明,胸径和液流密度是2个显著影响树干日液流量变异的因子,前者主要通过决定树干边材面积来控制树干液流量大小,它可以解释变异方差的56.9%;树干液流密度可以解释变异方差的34.7%.相关性分析表明,树干液流密度与与林木个体的生长指标(树高、胸径、冠幅和边材面积)无显著相关关系,但与林木的空间指标--树冠重叠度呈显著线性负相关(r=-0.668),即树干液流密度随树冠重叠度增加而降低,说明树干液流密度主要受其林木所处的空间位置及周围树木遮荫影响而发生变化.最后,利用树干液流密度与树冠重叠度之间的关系,提出基于林木空间差异估计华北落叶松林分蒸腾量的方法,并与常用的基于边材面积的尺度转换方法进行对比.结果表明,2种方法估计的林分日蒸腾量的数值变化趋势基本相同,但基于林木空间差异的方法估计的华北落叶松林分平均日蒸腾量为1.15mm·d-1,而基于边材面积的方法的估计值为1.32mm·d-1,前者低于后者13.13%,说明不考虑林木空间特征可能会导致林分日蒸腾量估计值偏大.     

基于热消散技术的毛竹林蒸腾耗水估算

【目的】试验不同长度热消散探针(TDP)测量毛竹液流的可行性,分析年龄对立竹液流的影响,并据此对立竹液流进行尺度扩展,估算桂北毛竹林的蒸腾耗水,为区域毛竹林的生态水文效应研究和指导关键生态功能区植被结构调整提供依据。【方法】显微镜观察毛竹输水结构在竹壁上的径向分布。基于热消散方法,用5 mm和10 mm长度的TDP探针对1～2年生立竹和3龄以上立竹的基部液流进行连续测量,并同步测定环境因子。【结果】维管束在毛竹竹壁上不均匀分布,竹壁外侧维管束小而密,导管分化不完全,竹壁内侧维管束大而疏,导管分化完全,直径较大。10 mm TDP探针测得的液流密度显著高于5 mm探针,其平均液流密度是5 mm探针的4.03倍。在生长旺季的7月,基于10 mm TDP探针测量的1～2年生立竹正午液流密度显著高于3龄以上立竹,而在早上和傍晚二者基本相同。1～2年生立竹液流的平均日通量在测量生长季内均高于3龄以上立竹,二者的平均日液流通量分别为51.15和33.80 g·cm^-2 d^-1。以立竹年龄和基径作为液流尺度扩展依据估测的桂北毛竹林日蒸腾耗水量在观测生长季内为0.01～0.72 mm·d^-1,平均日蒸腾耗水量为0.31 mm·d^-1。【结论】10 mm长度的TDP探针较5 mm探针更适宜用于毛竹液流的测量。1～2年生立竹比3龄以上立竹具有更高的液流密度和日液流通量,因此年龄是毛竹液流由立竹到林分尺度扩展时除立竹直径外另一个必须要考虑的因素。     

不同桉树人工林土壤呼吸旱雨季变化特征

以雷州半岛气候背景条件下具有良好培育前景的5种桉树人工林(湿加松林为对照)为研究对象,测定并分析各个林分土壤呼吸速率在旱季和雨季的差异,以揭示其与土壤温、湿度的关系.结果表明:土壤呼吸速率在旱季表现为先减后增变化趋势,在雨季表现为先增后减或逐渐减小变化趋势,旱、雨季变化特征明显.6个林分旱、雨季土壤呼吸速率均值分别为1.63~3.32μmol·m-2·s-1和2.55~4.36μmol·m-2·s-1.旱季土壤温、湿度共同促进土壤呼吸作用,雨季土壤温度促进土壤呼吸,土壤湿度抑制土壤呼吸作用.     

不同时间尺度下兴安落叶松树干液流密度与环境因子的关系

基于连续1年的兴安落叶松树干液流密度和环境因子(光照、空气温度、空气湿度、土壤温度和土壤湿度)的测定结果,探讨不同时间尺度下树干液流密度与环境因子的关系差异.在月时间尺度上,土壤温度和土壤湿度显著影响树干液流密度变化,土壤温度单位增加引起树干液流上升0.084～0.123 L·cm-2 month-1;在天时间尺度上,显著影响因子有土壤温度、光照和空气温度,其中土壤温度为最主要的影响因子,单位增加会导致树干液流上升1.9 ～2.7 mL·cm-2 d-1;在小时时间尺度上,主要影响因子在不同季节不同,但最主要因子多是直接影响地上叶片生理指标如光照和空气湿度,二者单位上升平均分别引起树干液流上升1.239 mL·cm-2 min-1和下降0.0566 mL·cm-2 min-1.随尺度由大到小,对树干液流影响最大的因子有从地下直接与根系水分吸收相关的土壤环境因子向地上直接影响叶片蒸腾的环境因子(光照和空气湿度)转变的趋势.同时,随着尺度增大,与树干液流显著相关的环境因子数明显下降,且相关系数R2显著提高,长期监测树木耗水可以采用监测环境因子反推的方法,而在小尺度上相同方法可能导致很大误差,最好采用直接测定法.     

西藏拉萨河谷区藏川杨和北京杨树干液流特征及其对环境因子的响应（英文）

【目的】研究西藏拉萨半干旱河谷地带藏川杨和北京杨树干液流的干湿季变化特征及环境影响因素,以期为该地区造林树种选择和合理配置提供科学依据。【方法】运用Granier热扩散探针技术,于2015年旱季和雨季在西藏拉萨半干旱河谷对藏川杨和北京杨的树干液流进行连续观测,并同步测定太阳辐射(Q)、空气温度(T_a)、土壤含水量(SWC)、饱和蒸气压亏缺(VPD)等环境因子。【结果】1)拉萨河谷区主要气象因子在旱季和雨季存在较大差异,雨季的Q、T_a、SWC、VPD等均高于旱季,主要气象因子的平均值为雨季的60%左右。2)藏川杨和北京杨的树干液流日变化表现出昼高夜低的规律,二者在雨季的日变化为多峰曲线,旱季为单峰曲线。旱季和雨季藏川杨液流速率均显著大于北京杨,北京杨液流密度在旱季和雨季相比藏川杨的液流密度分别下降71%和58%;相比雨季的液流密度,2种杨树的液流密度在旱季分别下降65%和76%。3)在旱季和雨季,主要环境因子Q和VPD与2种杨树树干液流间具有显著相关性,且Q和VPD的日变化与2种杨树的液流速率日变化之间存在明显的时滞效应。Q和VPD对北京杨树干液流的影响小于对藏川杨树干液流的影响。【结论】在拉萨河谷区旱季和雨季较大气候差异的环境下,藏川杨和北京杨树干液流速率均具有较为明显的昼夜节律。但2树种树干液流在峰值、日平均值、启动时间以及受环境因子的影响方面均存在明显差异,特别是在干旱季节尤为显著,藏川杨作为乡土树种显示出更强的适应性。     

雷州半岛尾巨桉人工林树干液流对台风天气的响应

[目的]为正确认识台风对桉树人工林蒸腾耗水的影响。[方法]采用热扩散茎流计对尾巨桉树干液流在台风前后进行连续监测,分析其对台风天气的响应,并同步测定林分气象条件,分析台风过程中树干液流与气象因子之间的相关性。[结果]表明:台风当天,树干液流密度与台风登陆前后(单峰)不同,呈不规律多峰型曲线,树干日均液流密度(2.89 mL·h~(-1)·cm~(-2))较台风前减少53%,夜间液流波动较大;日间最大峰值(7.76 mL·h~(-1)·cm~(-2))出现在下午,峰值较台风前降低55%,树干液流日通量极显著小于台风登陆前后;台风登陆前树干液流密度峰值(17.4 mL·h~(-1)·cm~(-2))及平均液流密度(6.15 mL·h~(-1)·cm~(-2))分别是台风登陆过后的1.24倍和1.14倍,台风后树干液流日通量较台风前均有所下降;台风当天,影响液流的主要气象因子为风速、大气温度、空气湿度、水汽压亏缺和降雨量,液流与风速的相关系数较台风前增大52%,与降雨量的相关系数是台风前的2.6倍,与其他因子的相关系数较台风前均下降了50%左右,而台风登陆前后之间的主要影响因子差异不大。[结论]台风当天的日均液流密度、白天的平均液流密度和日通量均极显著小于台风前后,而夜间液流显著大于台风前后;台风前后的日均液流密度差异不显著,夜间液流台风前显著大于台风后;台风过程中风速成为影响液流的关键因子,与太阳辐射相关性不显著,其他影响因子与台风前后相同,但相关系数除风速和降雨量外均显著下降。     

大青山油松人工林树干液流动态及其蒸腾耗水规律研究

应用TDP(Thermal Dissipation Probe)技术对大青山古路板林场的30a生油松人工林树干液流以及不同林分密度下的树木蒸腾耗水规律进行了研究。结果表明:1)在生长季内,树干径向断面形成层以下不同部位输水能力差异较大,最大流速位于形成层下3cm。2)油松树干液流的日进程呈现明显的昼夜变化规律。在11:00左右达到峰值,其值可达0.286 6~0.306 0cm3/s。3)树干液流量(Y)与树木的胸径(X)之间的关系可用Y=0.0053EXP(0.4823X)的指数函数模型表达。4)在相同立地条件下,随着阴坡林分密度从2 147株/hm2增加到4 463株/hm2,单株蒸腾耗水量从0.543 4cm3/s降低到0.319 7cm3/s,而林分蒸腾耗水量变化幅度较小,平均蒸腾耗水量为0.3710±0.0489(5)mm/h。结果指出,大青山30a生油松人工林经营密度应控制在3 307株/hm2左右。     

Sap flux – a real time assessment of health status in Norway spruce

Top dieback of Norway spruce (Picea abies), triggered by drought in 2004–2006, has been observed in Southeast Norway and trees died within four years after appearance of the first symptoms. The aim of our study was to use sap flux measurements as a diagnostic method for assessment of tree vitality. We used the heat field deformation method to monitor the sap flux density (SFD) in four pairs of healthy and declining trees in situ. To provide retrospective information on hydraulic performance of the trees we took samples for wood anatomical analysis. After felling the trees we used the modified differential translucence method (MDT) as a proxy for the SFD measurements. Healthy trees had three times higher SFD values as declining trees. In some healthy trees we detected decreasing SFD with time. The MDT method agreed with the SFD measurements. In conclusion, we detected sap flux dysfunction in declining trees and showed that the SFD reduction may occur during a short period, prior to occurrence of any visual symptoms. We suggest incorporating the SFD measurements into the repertoire of diagnostic tools in forest pathology.     

Estimating water use of a black locust plantation by the thermal dissipation probe method in the semiarid region of Loess Plateau, China

Black locust (Robinia pseudoacacia) is a major reforestation species in the semiarid region in the Loess Plateau of China. There has been increasing concern about the sustainability of the plantations because of their possible high water-use. This study was, accordingly, undertaken to quantify the stand-scale water use of a middle-aged black locust plantation in the region. The thermal dissipation probe method was applied to 27 trees to measure sap flux densities in an experimental plot during the growing season of 2008. The monoculture stand has a basal area of 23.3 m² ha^?1 and a maximum plant area index (PAI) of 2.89. Sapwood areas were estimated by use of a regressive relationship with the diameter at breast height (DBH) for scaling up of stand transpiration. The results showed that DBH could be a good predictor of sapwood area of individual trees. The diurnal cycles of average sap flux densities differed among DBH classes. Daily transpiration can be predicted from mean daily daytime vapor pressure deficit (VPD_m) using a fitted exponential saturation model. Model variables were different among seasons, probably owing to different soil water conditions and leaf phenology. By using the derived model for each month, stand canopy transpiration over the growing season was estimated to be 73.8 mm, with an average daily value of 0.41 mm day^?1 and a maximum of 0.89 mm day^?1. The relatively small estimates of stand transpiration might be attributed to low PAI and sap wood area of the middle-aged stand.     

生长季内官厅库区小叶杨单株树干液流通量密度的研究

利用热脉冲液流仪和微型自动气象站,在北京市延庆县上辛庄水土保持站观测研究了雨季不同天气条件下官厅库区的小叶杨树干边材液流通量密度变化规律及其对环境因子(气象条件、土壤水分)的响应.结果表明,边材液流通量密度曲线在不同天气条件下均具有明显的昼夜变化规律:液流通量密度通常于清晨5∶30-6∶30时开始上升;12∶00-13∶00时达到峰值;16∶00时开始持续下降,20∶00时下降速度变慢,至次日日出之前达到低谷.日液流通量密度(SFA)与日太阳辐射量(ESR)、日均空气温度(Ta)和日均空气相对湿度(RH)均存在极显著正相关关系,与日均0～40 cm土层蓄水量(Sw40)存在极显著负相关关系,经回归分析建立了日液流通量密度与各环境因子的线性关系:SFA=69.7 5.63ESR 3.12Ta 0.239RH-1.13Sw40(R=0.752).     

Variations in water use by a mature mangrove of Avicennia germinans, French Guiana

? In the tropical intertidal zones, little is known on water uptake by mangroves. Transpiration rates are generally measured at leaf level, but few studies exist on water use at tree or stand levels.

? The objective of this study was to measure sap flow in trees of different sizes to appreciate the range of variation in water use that may exist in a site dominated by 80% mature Avicennia germinans.

? The results showed that from the dry to the wet season the mean water use increased from 3.2 to 5.3 dm³ d^?1 in small trees (DBH ～ 13 cm), from 11.5 to 30.8 dm³ d^?1 in medium trees (～24 cm) and from 40.8 to 64.1 dm³ d^?1 in large ones (～45 cm).

? Sapwood remained active up to a depth of 8 cm with radial variations within the stem. Weak correlations were obtained with VPD and net radiation.

? This study confirmed that transpiration was larger under low levels of salinity. Water use at stand level (～1900 living stems ha^?1) was estimated to be in the range of 5.8 to 11.8 m³ ha?1 d^?1 according to the season.

     

Manipulating phenology and water relations in <Emphasis Type="Italic">Senna spectabilis</Emphasis> in a water limited environment in Kenya

The effect of shoot pruning on leaf phenology, stem wood anatomy and sap flow was investigated on Senna spectabilis (DC.) Irwin and Barneby in Machakos, Kenya. Unpruned trees (single stem) were compared to hedges (two to four stems), pruned 4 times a year during two rainy seasons (April–June, 1997 and November, 1997–January, 1998) separated by a dry season (July–October 1997). Trees attained peak leaf area of 55 m² plant⁻¹ during the rainy seasons, and shed all their leaves naturally during the dry season. Maximum hedge leaf area was 4 m² plant⁻¹ between pruning events and 5.2 m² plant⁻¹ during the dry season. Pruning induced multiple stems and narrow xylem vessels with low hydraulic conductivity. Average cross sectional area of conducting wood per plant was at least 1.8 times greater in trees than in hedges. Xylem lumen diameter at 5 mm depth below the cambium was significantly (P < 0.001) larger in trees (53.6 ± 6.21 μm) than that in hedges (36.2 ± 8.21 μm). Maximum sap flow occurred in the wet season for trees (4800 g d⁻¹ plant⁻¹) and in the dry season for hedges (1400 g d⁻¹ plant⁻¹). Wet season pruning suppressed crown expansion and modified the natural phenology of senna, reducing transpiration rate and therefore soil water depletion, causing crowns to grow. This enhanced the ecological combining ability of senna managed as hedges with annual crops.     

应用热扩散式探针法对南方红壤丘陵区湿地松人工林耗水量的研究

运用Granier热扩散式探针法对湿地松(Pinus elliottii)树干液流密度进行长期连续测定,并对其周围多个气象要素和土壤要素进行同步测定。湿地松生长旺季不同月份晴天液流密度日变化规律基本一致,但液流启动、到达峰值的时间以及开始升高、开始下降的时间间隔存在差异。样木解析结果表明,处于低龄期的湿地松,整个木质部都可看作边材,边材面积和胸径可用二次函数很好地拟合。方差分析结果表明:不同胸径湿地松树干液流密度不存在显著性差异,与胸径无紧密关系。回归分析结果表明:湿地松日累计液流量与边材面积呈显著的线性相关关系,相关关系的判定系数达0.99。另外,单位边材面积日累计液流量与入选各气象因素的相关性显著,相关程度大小顺序为光合有效辐射>冠层相对湿度>冠层温度。结合边材面积-胸径回归方程和样木胸径调查资料,分别采用2种方法推算整个标准地的耗水量。经比较,2种方法所得结果差别不大,各自适用于一定的范围。     

Cacao trees under different shade tree shelter: effects on water use

We asked how shade tree admixture affects cacao water use in agroforests. In Central Sulawesi, Indonesia, cacao and shade tree sap flux was monitored in a monoculture, in a stand with admixed Gliricidia trees and in a mixture with a multi-species tree assemblage, with both mixtures having similar canopy openness. A Jarvis type sap flux model suggested a distinct difference in sap flux response to changes in vapor pressure deficit and radiation among cacao trees in the individual cultivation systems. We argue that differences originate from stomatal control of transpiration in the monoculture and altered radiation conditions and a different degree of uncoupling of the VPD from the bulk atmosphere inside shaded stands. Probably due to high sap flux variability among trees, these differences however did not result in significantly altered average daily cacao water use rates which were 16 L day^?1 in the multi-species assemblage and 22 L day^?1 in the other plots. In shaded stands, water use of single cacao trees increased with decreasing canopy gap fraction in the overstory since shading enhanced vegetative growth of cacao fostering transpiration per unit ground area. Estimated transpiration rates of the cacao tree layer were further controlled by stem density and amounted to 1.2 mm day^?1 in the monoculture, 2.2 mm day^?1 for cacao in the cacao/Gliricidia stand, and 1.1 mm day^?1 in the cacao/multi-species stand. The additional transpiration by the shade trees is estimated at 0.5 mm day^?1 for the Gliricidia and 1 mm day^?1 for the mixed-species cultivation system.     

柠条主根液流与叶面积关系初探

运用热扩散技术对3株柠条主根部液流速率进行了监测,对试验期间典型晴天日(30d)液流速率的分析表明:柠条根部液流传输受大气蒸发力、冠层输出、边材面积的共同影响;1、2、3号样株边材面积分别为6.61、7.17、5.06 cm2,日平均液流总量分别为457.92、1014.66、292.12 g·d-1,日平均液流总量随边材面积的增加而增大;各样株间日平均液流总量的差异显著,而单位叶面积的液流量无显著差异;2号柠条样株的液流速率日变化动态与潜在蒸发散的相关性最强,叶面积/边材面积为0.62m2·cm-2,可以作为评定柠条单株是否具有与当地水文条件相适应的根冠比例关系的重要指标.试验期间样株根区60～140 cm内土壤平均含水量分别占田间持水量的51.04%、63.26%和87.0%,土壤水分状况较好.     

Temporal variations in transpiration of <Emphasis Type="Italic">Vitellaria paradoxa</Emphasis> in West African agroforestry parklands

Lack of data on water use of key species of drylands constitutes an obstacle to understanding their role in hydrological processes in this environment. To elucidate seasonal variation in water consumption by Vitellaria paradoxa, the dominant species of parklands of the semi-arid areas of West Africa, we’ve measured its transpiration using heat ratio method (HRM) and seven potential explanatory variables. Sap flux was found to be significantly different among years with 0.64, 0.59 and 0.67 L h^?1 dm^?2 in 2008, 2009 and 2010, respectively. Sap flux was significantly higher in the dry (0.73 L h^?1 dm^?2) than in the wet season (0.53 L h^?1 dm^?2). Nighttime sap flux during dry season (0.48 L h^?1 dm^?2) was significantly higher than that of the wet season (0.20 L h^?1 dm^?2) and it contributes on average to 26% of daily sap flow with a maximum reaching 49%. The mean transpiration rate per tree was 151 L day^?1 and all measured variables except rainfall and soil water content were significantly correlated with sap flux. These correlations were stronger (higher R value) during the rainy than in the dry season. Vapor Pressure Deficit (VPD) explained the highest proportion of sap flux variation and their curve was of parabolic type (R² = 0.54) indicating that V. paradoxa can probably down-regulate its canopy conductance beyond a certain threshold of VPD, which is about 3 kPa in the present study. Future studies should investigate such hypothesis as well as the impacts of the variation of V. paradoxa transpiration due to climatic variables on hydrological cycles.