Prescribed burning of Pinus oocarpa in Honduras II. Effects on nutrient cycling

The effects of prescribed burning on nutrient cycling were studied in a stand of oocarpa pine (Pinus oocarpa Schiede) in central Honduras. Small paired runoff plots were subjected to a low-intensity prescribed burn in one of two seasons and nutrient losses in runoff water and sediment were monitored together with nutrient inputs in bulk precipitation.Burning increased the loss of nutrients and net losses of calcium, potassium and magnesium were recorded during the first $\text{16}\text{1}\text{2}$ months after burning. Increased rates of sediment loss rather than increased losses in surface runoff were responsible for the greater part of the increase in total nutrient losses from the burned plots. Greater quantities of nutrients were lost from the plots burned just prior to the dry season than from those burned after the start of the wet season because of slow recovery of the ground vegetation during the dry season and exposure to the intense rains of May and early June. Recovery from burning was rapid and runoff and sediment loss from the burned plots approached the levels of controls 2 years after burning. As nutrient losses are a reflection of total runoff and sediment losses, it is probable that no net nutrient losses would have been detected during the 2nd year after burning had sampling been continued. The net nutrient losses recorded were small and this is attributed to the low initial fuel loadings, low fire intensities and the rapid rate of recovery of the ground vegetation. The significance of other potential sources of losses and gains of nutrients is discussed and it is concluded that low-intensity prescribed burning for hazard reduction, with burning cycles ranging from three to seven or more years, will reduce the rate of nutrient accumulation on the site rather than deplete the nutrient capital.     

Agroforestry for soil and water conservation in the western Himalayan Valley Region of India 1. Runoff, soil and nutrient losses

Erosion losses were investigated in a field experiment, from 90 × 15 m erosion plots at 4% slope in the western Himalayan valley region of India. The plots were planted with Leucaena leucocephala and Eucalyptus hybrid, either as block plantation or in alley farming with maize (Zea mays), Chrysopogon fulvus grass or turmeric (Curcuma longa). The treatments also included contour planting of sole maize, sole Chrysopogon fulvus grass and a clean weeded cultivated fallow. During the nine year study period, the average annual monsoon rainfall was about 1000 mm and it caused 347 mm runoff and 39 Mg ha^-1 soil loss per year from fallow plots. The runoff and soil loss were reduced by 27% and 45% by contour cultivation of maize. Contour tree-rows or leucaena hedges reduced the runoff and soil loss by 40% and 48%, respectively, over the maize plot, reducing soil loss to about 12.5 Mg ha^-1. This reduction in erosion was primarily due to the barrier effect of tree or hedgerows and micro-terraces formed through sediment deposition along the contour barriers. Such vegetative measures, that are productive while being protective, offer viable alternative for erosion control in areas with gentile slopes of the valley region. High density block plantations of eucalyptus and leucaena almost completely controlled erosional losses and can be recommended for steeper slopes that are vulnerable to heavy erosion. This revised version was published online in June 2006 with corrections to the Cover Date.     

Prescribed fire,snag population dynamics,and avian nest site selection

Snags are an important resource for a wide variety of organisms, including cavity-nesting birds. We documented snag attributes in a mixed-conifer forest dominated by ponderosa pine in the Sierra Nevada, California where fire is being applied during spring. A total of 328 snags were monitored before and after fire on plots burned once, burned twice, or left unburned to assess the effects of prescribed fire on snag populations. The greatest loss of snags (7.1 snags ha⁻¹ or 43%) followed the first introduction of fire after a long fire-free period. On plots burned a second time 21% of snags (3.6 snags ha⁻¹) were lost, whereas 8% (1.4 snags ha⁻¹) were lost on unburned control plots in the same time period. New snags replaced many of those lost reducing the net snag losses to 12% (2.0 ha⁻¹) for plots burned once, and 3% (0.5 ha⁻¹) for plots burned twice and unburned plots. We also examined snags used by cavity-nesting birds. Snags preferred for nesting were generally ponderosa pine (Pinus ponderosa), larger diameter, and moderately decayed as compared to available snags. For monitored snags that met the preferred criteria, there was a net loss (1.7 snag ha⁻¹ or 34%) after the first burn, while the loss of useable snags was less than 1 snag ha⁻¹ following the second burn (15%) or on unburned controls (8%). We recommend protection of preferred snags, in particular large ponderosa pines, especially during primary fire applications on fire-suppressed landscapes.     

南方红壤区不同管理模式下的马尾松林水沙特征与植被作用

基于标准坡面径流小区观测,比较不同雨量水平植被对地表径流量的贡献,同时利用水文功能连通性原理和灰色关联法对相关土壤和植被参数进行比较。结果表明:①小区减流减沙效果为“林下流”<封禁管理<马尾松纯林<人工乔灌草混交;②地上和地下的植被特征对坡面水文过程阻抗明显;③相关参数的影响程度为土壤含水量<植被盖度<植被高度<有机质含量。不同雨量下小区水沙过程不相一致,小区尺度上植被特征对坡面水土保持起重要作用。     

黄土丘陵区主要人工林水保及其土壤水分生态效益研究

通过研究表明，不同类型植被减少径流和泥沙的功效不同，东坡不同类型植被小区径流深由大到小的顺序为：荒坡＞沙棘（平茬）＞沙棘＋小叶杨＞农地；北坡为：油松＞油松＋沙棘＞沙棘林；东坡产沙量由大到小的顺序为：农地＞荒坡＞沙棘（平茬）＞沙棘＋小叶杨，北坡为：油松＞油松＋沙棘＞沙棘。不同类型植被在生长季的土壤水分平均值均很低，其中对土壤水分利用强度由大到小分别为柠条＞沙棘＞刺槐，100cm以下土层，不同类型林地土壤均存在利用性干层。     

Impact of vegetation restoration on hydrological processes in the middle reaches of the Yellow River, China

Sediment discharge from the Yellow River originates mainly from the drainage area between Hekouzhen and Longmen, i.e., the Helong area. Spatial-temporal variations of the vegetation cover in this area during the 1981–2007 period have been investigated using GIMMS and SPOT VGT NDVI data. We have also analyzed the interannual variations in vegetation cover and changes in annual runoff and sediment discharge, the consequences from precipitation change and the Grain for Green Project (GGP). The results show that vegetation cover of the Helong area has increased during the 1981–2007 period. The northwestern part the Helong area, where the flat sandy lands are covered by grass, has experienced the largest increase. The region where the vegetation cover has declined is largely found in the southern and southeastern Helong area, which is a gullied hilly area or forested. Although precipita-tion was relatively low during the 1999–2007 period, the vegetation cover showed a significant increase in the Helong area, due to the implementation of the GGP. During this period, the most significant improvement in the vegetation cover occurred mainly in the gullied hilly areas of the Loess Plateau, such as the drainage basins of the Kuyehe and Tuweihe rivers and the middle and lower reaches of the Wudinghe and Yanhe rivers. A comparison of the average annual maximum NDVI between the earlier (1998–2002) stage and the next five years (2003–2007) of the GGP indicates that the areas with increases of 10% and 20% in NDVI account for 72.5% and 36.4% of the total area, respectively. Interannual variation of annual runoff and sediment discharge shows a declining trend, especially since the 1980s, when the decrease became very obvious. Compared with the 1950–1969 period, the average runoff during the 1980–2007 period was reduced by 34.8×108 m3 and the sediment discharge by 6.4×108 t, accounting for 49.4% and 64.9% of that in the 1950–1969 period, respectively. There is a positive correlation between the annual maximum NDVI and annual runoff and sediment discharge. This correlation was reversed since the implementation of the GGP in 1999 and vegetation cover in the He-long area has increased, associated with the decrease in runoff and sediment discharge. Less precipitation has been an important fac-tor driving the decrease in runoff and sediment discharge during 1999–2007. However, restoration and improvement of the vegetation cover may also have played a significant role in accelerating the decrease in annual runoff and sediment discharge by enhancing evapotranspiration and alleviating soil erosion.     

Tree mortality after low-intensity prescribed fires in managed Pinus sylvestris stands in southern Finland

Abstract

Tree mortality, its causes, and the input of dead charred wood were studied in 11 managed 30–45-year-old Scots pine (Pinus sylvestris L.) stands 1 year after experimental low-intensity prescribed burnings in southern Finland. First, the relationship between fire-induced tree damage and several external variables, e.g. stand density, within-stand wind speed, open-air wind speed, the Finnish Forest Fire Index (FFI) and flame height, was studied. Secondly, the study examined which damage and morphological characteristics best predicted tree mortality. Tree mortality was very variable in the experimental plots, ranging from 0% to 48% on the basis of stem number and from 0% to 41% in terms of wood volume. The input of dead and charred wood decreased with stand age, being 19.4 m³ ha^?1 in 30–35-year-old stands, but only 1.7 m³ ha^?1 in 45-year-old stands. The input of dead wood was on average 10 m³ ha^?1, representing less than 5% of the mean volume before the prescribed fire. The external variables that best explained fire-induced damage were within-stand wind speed, flame height and FFI. Tree mortality was best predicted by charred stem ratio with bark thickness, and by charred stem ratio with tree diameter. The results indicate that prescribed burning that is conducted downwind increases tree mortality and changes subsequent stand structure with increasing within-stand wind speed.     

Effects of wood-ash on the tree growth,vegetation and substrate quality of a drained mire: a case study

The effects of wood-ash fertilisation on tree stands, soil characteristics and ground vegetation were studied on a drained pine mire in Finland (64°51′N, 26°04′E, 62 m a.s.l.). The original site type was a treeless, mesotrophic Sphagnum papillosum fen. The site was drained in 1933 and the wood-ash fertilisation experiment was started in 1947. The treatments were: (i) unfertilised, (ii) wood-ash 8 t ha⁻¹, and (iii) wood-ash 16 t ha⁻¹.Drainage and ash application had radical and long-lasting consequences on the biological activity on the site and the vegetation compartments studied. The understorey vegetation had been profoundly affected by the ash with almost complete transformation of the species and other life forms. Even 50 years after the ash treatment the changes in vegetation/site type and the tree stand were clearly visible. On the unfertilised plot, the biomass of ground vegetation consisted mostly of mosses and dwarf shrubs, but on the ash-treated plots it consisted mostly of herbs and grasses typical of upland forests.Ash treatment had promoted stem volume growth of Scots pine (Pinus sylvestris L.) substantially and for a long time. The total wood production on the ash plots during 1947–1994 was 13 and 17 times over that of the control plot. Unfertilised pine trees suffered from P and K deficiency throughout the study period. The concentrations of some plant nutrients (P, K) decreased during the past years on Ash8. No nutrient shortage afflicting the tree stand was observed on Ash16 during the study period.Ash application has also led to increased concentration of nutrients in the peat. A sizeable proportion of the mineral nutrients applied were still in the 0–20 cm peat layer. On the ash-treated plots the amount of soil nitrogen (0–20 cm) was 18 and 29 times and the amount of soil phosphorus 9 and 13 times over the amount bound by the tree stand and the ground vegetation (Ash8 and Ash16, respectively). The stock of potassium was generally small in the surface peat—only 60–90% of the amount of potassium bound in the tree stand and the ground vegetation.It was concluded that wood-ash had powerfully influenced the biological processes in surface peat. The decomposition of cellulose was significantly accelerated by both ash treatments. Ash fertilisation also increased the emissions of CO₂. The intensified decomposition rate in the litter, vegetation and peat explained to a large extent the accelerated growth of the Scots pine stands studied.     

Short-term impacts of harvesting and burning disturbances on physical and chemical characteristics of forest soils in western Newfoundland,Canada

The short-term impacts of prescribed burning were determined for the physical and chemical properties of a forest soil on a clear-cut in western Newfoundland, Canada. The experimental site was predominantly covered by black spruce trees (Picea mariana Mill.) that were harvested in 1996. In August 1998, prescribed burning removed most of the logging residues left on the site after harvesting, all above-ground surface vegetation, and parts of the upper humus layer. In October 1998, field samples were taken from four replicated burned plots and four replicated unburned plots. In each study plot, soil samples were taken from the organic layer (F+H) and from the top 10 cm of the mineral soil. In the burned treatments, mass of the humus layer (F+H) was reduced by 24% and, because of the accretion of basic ash materials, acidity of the humus layer was also reduced by up to 1 unit. In the organic layer and the mineral soil, total contents of Mg, Ca, and P, extractable Mg and Ca, available P, sum of NH₄Cl-extractable cations, and C/N ratios were increased by burning, while total C and N as well as total and extractable K remained unaffected by burning.     

三峡库区典型森林植被类型对坡面产流影响

在一定地形地貌基础上,植被与降水是决定坡面产流、产沙量的主要因素.2005～2006年对重庆缙云山4个天然林地径流场和农地径流场进行定位观测,结果表明:①森林植被在一定范围内对坡面径流量有削减作用,前期降雨对坡面径流有影响;②坡面径流峰值与暴雨峰值呈一一对应关系,马尾松和常绿阔叶林的变化幅度较小,楠竹林和常绿阔叶灌丛变化较大;③地下径流深与降雨量呈一一对应关系,地下径流深与降雨量关系反映出前期降雨对地下径流量的影响;④马尾松阔叶混交林、常绿阔叶林和常绿阔叶冠丛地下径流峰值的总体趋势是随降雨强度的增加而增加;楠竹林和农地随降雨强度的增加而呈下降趋势.     

Residue management of a planted fallow on an acid soil in Cameroon: crop yields and soil organic matter fractions

A fallow management trial was initiated in March 1990 in Yaounde, Cameroon, with the objective of identifying an efficient method of managing residue derived from a planted fallow of Cajanus cajan. Ten months after establishment, the shrubs were slashed and residues were treated in one of the following ways: burnt, incorporated in the soil, mulched on soil surface or removed. Following that, maize and groundnut were planted. Soil organic matter was fractionated after the residue treatments, and elementary partial budgeting was conducted. After three cycles of fallow and cropping seasons, maize yield was similar, about 3 t ha^-1, in all plots except in the residue-removed plots, where it was 1.4 t ha^-1. Yield trend of groundnut was also similar. The residue management method did not affect either the nature of fractions or the total content of soil organic matter. The highest net return, US$5945 ha^-1 year^-1, was obtained from the residue incorporated treatment, whereas the highest return to labour, US$11 per manday^-1 was associated with burning of residues. In areas such as the forest zone of Cameroon where labour is a major constraint and climatic conditions allow vigorous vegetation growth, burning appears to be the best method of residues management, at least in the short run. This revised version was published online in June 2006 with corrections to the Cover Date.     

Implications of fires on carbon budgets in Andean cloud montane forest: The importance of peat soils and tree resprouting

Fire in tropical montane cloud forests (TMCFs) is not as rare as once believed. Andean TMCFs sit immediately below highly flammable, high-altitude grasslands (Puna/Páramo) that suffer from recurrent anthropogenic fire. This treeline is a zone of climatic tension where substantial future warming is likely to force upward tree migrations, while increased fire presence and fire impacts are likely to force it downwards. TMCFs contain large carbon stocks in their peat soils and their loss through fire is a currently unaccounted for regional source of CO₂. This study, conducted in the southern Peruvian Andes (>2800 m), documents differences in live tree biomass, fine root biomass, fallen and standing dead wood, and soil organic carbon in 4 paired-sample plots (burned versus control) following the severe ground fires that occurred during the 2005 Andean drought. Peat soils contributed the most to biomass burning emissions, with lower values corresponding to an 89% mean stock difference compared to the controls (mean ± SE) (54.1 ± 22.3 vs. 5.8 ± 5.3 MgC ha⁻¹). Contrastingly, carbon stocks from live standing trees differed by a non-significant 37% lower value in the burned plots compared to the controls, largely compensated by vigorous resprouting (45.5 ± 17.4 vs. 69.2 ± 13.4 MgC ha⁻¹). Both standing dead trees and fallen dead wood were significantly higher in the burned plots with a three-fold difference from the controls: dead Trees 45.2 ± 9.4 vs. 16.4 ± 4.4 MgC ha⁻¹, and ca. a 2 fold difference for the fallen dead wood: 11.2 ± 5 vs. 6.7 ± 3.2 MgC ha⁻¹ for the burned plots versus their controls. A preliminary estimate of the regional contribution of biomass burning emissions from Andean TMCFs for the period 2000-2008, resulted in mean carbon emission rates of 1.3 TgC yr⁻¹ (max-min: 1.8-0.8 TgC yr⁻¹). This value is in the same order of magnitude than South American annual fire emissions (300 TgC yr⁻¹) suggesting the need for further research on Andean forest fires. On-going projects on the region are working on the promotion of landowner participation in TMCFs conservation through REDD+ mechanism. The heart of the proposed initiative is reforestation of degraded lands with green fire breaks enriched with economically valuable Andean plant species. The cultivation of these species may contribute to reduce deforestation pressure on the Amazonian cloud forest by providing an alternative income to local communities, at the same time that they prevent the spread of fire into Manu National Park and adjacent community-held forests, protecting forest and reducing CO₂ emissions.     

Characteristics of runoff and sediment generation of forest vegetation on a hill slope by use of artificial rainfall apparatus

We studied the impact of forest vegetation on soil erosion,surface runoff, and sediment generation by using field simulated rainfall apparatus. We measured runoff and sediment generation of five 4.5 × 2.1m runoff plots (a bare soil as a control; two Pinus tabulaeformis forestplots and two Platycladus orientalis forest with row spacing of 1 m × 1m and 1.5 m × 1.5 m, respectively) in Beijing Jiu Feng National ForestPark under three rainfall intensities (0.42, 0.83, 1.26 mm per minute).Forest vegetation significantly reduced soil erosion and sediment yield.Mean total runoff volume in the four tree stand plots was 93% of that inthe control plot, demonstrating the limited effectiveness of forest vegetation in runoff control. With increasing rainfall intensity, runoff reductionin forest plots declined from 28.32% to 2.1%. Similar trends in runoff coefficient and the relationship between runoff volume and rainfall duration was observed. Mean total sediment yield and mean sediment yield reduction rate under different treatments was 55.05% and 43.17% of those in the bare soil control plot, respectively. Rainfall intensity playedan important role in runoff and sediment generation processes, and had agreater impact on runoff than on soil erosion and sediment generation.When considering several factors in runoff and sediment transport processes, the P. tabulae form plot with row spacing at 1 × 1 m had a greater effect on soil and water conservation than did other forested plots.     

Immediate effects of fire-severity on soil invertebrates in cut and uncut pine forests

We studied how variations in fire severity and the degree of cutting before burning affected soil invertebrates in a Pinus sylvestris forest in central Sweden. A varied depth of burn in the mor layer was obtained by exclusion of rain and addition of fuel in small plots (1 m×2 m) in clear-cut, selectively cut and uncut part of the forest before large-scale prescribed burning took place. Soil samples were taken from the plots immediately before, the day after, and two months after the fire. The overall mortality of invertebrates depended on the proportion of organic soil consumed by the fire, and for individual taxa it ranged between 59 and 100%. Invertebrates that lived deeper in soil suffered lesser mortality than those in the vegetation and litter layers did. Greater mobility in soil (Staphylinidae) or a thick cuticle (Oribatediae, Elateridae) may have contributed to the higher survival observed in these taxa. The beetles Atomaria pulchra (Cryptophagidae), Corticaria rubripes (Lathridiidae), and other fire-favoured insects colonised the burned forest the very day the fire burned. These species preferred the hard-burned plots and the uncut stand for colonisation. Sixty days after the fire, the abundance of invertebrates was lower in the burned cut stands compared to the burned uncut stand. The species composition of beetles in the burned stands was then characterised by a few very abundant fire-favoured species.     

Effects of foliage and traffic intensity on runoff and sediment in skid trails after trafficking in a deciduous forest

After ground-based skidding operations, soil particles are rearranged closer together resulting in increased bulk density, reduced total porosity, and decreased infiltration capacity. The aim of the present study was to assess and quantify runoff and sediment in both leafed and leafless periods (foliage) over a 1-year duration following three levels of skidder traffic intensity (low, medium, high) performed in the Hyrcanian forests, Iran. By increasing traffic frequency, soil bulk density increased significantly, whereas total porosity decreased as compared to undisturbed (control) plots. After trafficking, runoff and sediment yield were significantly affected by foliage and traffic frequency. Regardless of foliage, mean total runoff and sediment increased with increasing traffic frequency performed on the skid trail. Mean runoff in the low, medium, and high traffic plots in the leafless period were 95.5, 54.2, and 21.7% higher than the values of runoff in the leafed period, respectively. Mean sediment yield in the low, medium, and high traffic plots in the leafless period was 7.1, 5.1, and 3.3 times higher than the values of sediment in the leafed period, respectively. Based on regression analysis, the runoff and sediment response to rainfall for the control plots and three traffic intensity classes were linear and statistically significant for both leafless and leafed periods. The total runoff in the high traffic intensity plot was 239.8 mm (62%) during the leafless period, and 38% of total runoff (148 mm) occurred during the leafed period. The sediment response to runoff over both leafless and leafed periods at all the traffic intensity classes and control plots were significantly linear. Restricting ground-based skidding operations to trail segments where the machine traffic is less than 4 (low)—9 (medium) traffic cycles can be effective to reduce runoff and sediment over compacted soil after machine-induced traffic.     

Effect of Vegetation and Precipitation upon Runoff and Sediment Production on Slope Lands of the Loess Area in China

INTRODUCTION The vegetation construction on the Loess Plateau is of significant importance in ameliorating the serious situation of soil and water conservation. The Loess Plateau is characterized by lots of gullies, and its surface soil is fragile and easily eroded, the factors that influence soil and water loss are: rainfall, vegetation, slope and others. Rainfall is the motive, power of causing soil and water loss; besides, vegetation and slope also greatly affect soil and water loss. …     

Impacts of low-intensity prescribed fire on microbial and chemical soil properties in a <Emphasis Type="Italic">Quercus frainetto</Emphasis> forest

Prescribed fire is a common economical and effective forestry practice, and therefore it is important to understand the effects of fire on soil properties for better soil management. We investigated the impacts of low-intensity prescribed fire on the microbial and chemical properties of the top soil in a Hungarian oak (Quercus frainetto Ten.) forest. The research focused on microbial soil parameters (microbial soil respiration (R_SM), soil microbial biomass carbon (Cmic) and metabolic quotient (qCO₂) and chemical topsoil properties (soil acidity (pH), electrical conductivity (EC), carbon (C), nitrogen (N), C/N ratio and exchangeable cations). Mean annual comparisons show significant differences in four parameters (C/N ratio, soil pH, Cmic and qCO₂) while monthly comparisons do not reveal any significant differences. Soil pH increased slightly in the burned plots and had a significantly positive correlation with exchangeable cations Mg, Ca, Mn and K. The mean annual C/N ratio was significantly higher in the burned plots (28.5:1) than in the control plots (27.0:1). The mean annual Cmic (0.6 mg g^?1) was significantly lower although qCO₂ (2.5 µg CO₂–C mg Cmic h^?1) was significantly higher, likely resulting from the microbial response to fire-induced environmental stress. Low-intensity prescribed fire caused very short-lived changes. The annual mean values of C/N ratio, pH, Cmic and qCO₂ showed significant differences.     

尖山河流域不同植被类型坡面产流产沙量研究

对云南省玉溪市小尖山流域径流小区进行定位观测试验,对珠江石灰岩溶地区的不同植被类型坡面产流产沙量进行分析,结果表明：①该小流域降雨量年内分配不均,雨季（6—9月）占全年降雨量的83．2％,当降雨量大于5mm或10min雨强大于9mm／h时都会使该流域坡耕地发生产流．②灌草丛和次生林比坡耕地减少产流54．29％和92．06％,比人工林减少产流52．78％和87．13％．4个径流小区的产流大小顺序为：坡耕地〉人工林〉退耕地-灌草丛〉次生林．③土壤侵蚀量随着径流量的增加而增加,灌草丛和天然次生林分别比坡耕地减少土壤侵蚀量95．96％和99．02％,比人工林减少土壤侵蚀量64．60％和91．40％,其大小顺序与产流量一致．坡耕地、人工林、灌草丛的侵蚀量主要是由几次单次侵蚀贡献的,而次生林则是平均贡献．因此次生林防止水土流失的效果最好,在建立人工林时,应该考虑合适的复层结构,乔灌结合才能达到水土保持的最佳效果．     

Gradient analysis of an eastern sand savanna's woody vegetation,and its long-term responses to restored fire processes

Little information is available comparing historic and modern sand savannas, and how remnants respond to restored fire. We compared short- and long-term effects of restored fire on the Tefft Savanna, a 197 ha eastern sand savanna in northwest Indiana that had undergone three decades of fire protection. U.S. Public Land Survey data from Tefft in 1833 indicate black and white oak barrens, and pin oak savanna, with trees averaging 50 stems/ha and 4 m²/ha basal area. We used ordination and a digital elevation model to assess topographic distribution of tree species in 1986. In 1986, we also compared initial effects of high- and low-intensity dormant season fire on woody vegetation among nine blocks containing black oak, white oak, and pin oak stands. Twenty years later, we compared the same blocks, all of which had been burned three times per decade with low-intensity fires. In 1986, black oak, white oak and pin oak occurred across a gradient of decreasing elevation and slope. At that time, unburned black oak and white oak stands averaged >400 stems/ha and about 10 m²/ha basal area, and their smaller size classes contained non-oak woody vegetation that apparently had invaded with fire exclusion. After initial burns, black oak and white oak stands receiving high-intensity fire averaged <200 stems/ha and had significantly lower oak canopy cover and basal area than unburned stands. Stands receiving low-intensity fire had intermediate oak canopy cover, with basal area similar to unburned stands. Pin oak stands were more fire-resistant, apparently because spring flooding often reduced fire effects. Density, cover and basal area of non-oak tree species were much lower than oaks, and were not reduced by initial burning. Repeated low-intensity burning over 20 years tended to maintain structure caused by initial fires. However, it reduced lower size class stem densities, promoted post-fire sprouting into the shrub layer, and allowed oak basal area to increase in larger size classes. Time since fire regulated shrub layer structure on a 4-year cycle. Density and cover of trees and shrubs returned to pre-burn conditions by the second and fourth growing seasons after fire, respectively, with non-oak tree species exceeding pre-burn cover and density by the fourth season. These results suggest that high-intensity fire is more important than repeated low-intensity burning in structuring and restoring eastern sand savanna, and that non-oak tree species, once established, may be resistant to low-intensity fire.     

Oleoresin flow and chemical composition of Corsican pine (Pinus nigra subsp. laricio) in response to prescribed burnings

Oleoresin is an important defensive made up of acidic and neutral diterpenes together with a variable fraction of monoterpenes and sesquiterpenes. It plays a major role in plant-herbivore defense. Fire, or prescribed burning, can be a stress factor leading to metabolic changes in pine needles. Prescribed burning is used to: (i) restore fire to the ecosystem, (ii) recreate natural disturbance dynamics and (iii) reduce fuel loadings and fire risks. To our knowledge, no study has been performed on the effects of fire on the oleoresin defense system in pine trees, apart from the measurement of resin flow. We examined the effect on oleoresin production (resin flow) and the chemical composition (volatile and diterpene components) in Corsican pine (Pinus nigra subsp. laricio (Poir.) Maire var. corsicana (Loud.) Hyl.). This species is an ecologically and economically important pine in France. The chemical composition of samples was studied using HS-SPME GC and GC/MS for the volatile fraction and ¹³C NMR for diterpenes. A considerable increase in resin production and, consequently, an increase predominantly in diterpenes together with volatile compounds were observed at stations which had sustained recent prescribed burnings. Resin flow in trees in low-intensity prescribed burning after 4 years were no different than trees in unburned plots. Resin flow can be considered as a response to thermal stress. The major components of the two fractions were α-pinene (66.9–75.6%) and levopimaric acid (19.8–23.0%), respectively. The chemical composition of the resin remained unchanged after prescribed burning, even though resin production increased. No insect attacks were observed after low-intensity prescribed burning of P. laricio.