人为干扰对典型草原生态系统土壤养分状况的影响

从土壤N，P，K及土壤有机质等方面研究了人为干扰（包括放牧、开垦和禁牧）对典型草原生态系统土壤养分含量的影响。研究结果表明：开垦和放牧会导致土壤，主要是表层土壤养分含量的下降，而开垦对土壤养分的影响更为明显；禁牧会提高土壤各种养分的含量，而且随着禁牧时间的增加，土壤养分含量有逐渐增加的趋势。对深层土壤养分含量没有明显影响。同时，草原生态系统土壤全氮、速效氮及有机质含量与土层深度呈明显的负直线相关关系；速效K、速效P含量则与土壤深度的关系符合二次幂函数关系；全磷含量随深度的增加有逐渐升高的趋势；土壤全钾含量随土壤深度的增加呈微弱的降低趋势，但变化不明显。本研究表明，禁牧可以提高典型草原土壤养分元素的含量，有利于遏制草原土壤的退化。     

放牧对荒漠草原植物生物量及土壤养分的影响

以宁夏荒漠草原为研究对象,探讨放牧对荒漠草原植物多样性、 生物量及土壤养分特征的影响。结果表明, 放牧对荒漠草原植物群落多样性、 均匀度和丰富度影响显著。植物群落多样性和均匀度随着放牧强度的增加均呈先增加后降低的趋势,在轻度放牧达到最大值。同围封禁牧相比,重度、 中度和轻度放牧草地的植物地上和地下部生物量显著降低,分别降低了43.8%、 42.0%、 15.4% 和 27.7%、16.2%、11.9%。土壤有机碳随着放牧强度的增加而降低,而土壤全氮含量随着放牧强度的增加呈先增加后降低的趋势。围封禁牧草地土壤有机碳比重度放牧增加了18.1%,而土壤全磷、 速效磷和全钾含量分别降低了 21.1%、 51.9% 和 11.0%。土壤有机碳含量对植物群落地上和地下部生物量的影响大于土壤全氮、 全磷、 全钾、 速效磷和速效钾。放牧干扰下荒漠草原土壤环境及其养分含量,能在一定程度上反映植物群落多样性和生物量的变化。     

放牧强度对荒漠草原植被、土壤及其侵蚀特征的影响

[目的] 研究不同放牧强度对荒漠草原植被特征、土壤物理性能及其土壤侵蚀的影响,为荒漠草原水土流失治理和退化及生态系统修复提供科学依据。[方法] 以内蒙古希拉穆仁荒漠草原为研究区,采用原位监测和模拟试验相结合的方法,对重度、中度、轻度和无放牧4种不同放牧强度样地的植被群落、土壤物理性质、土壤侵蚀过程开展了研究。[结果] ①随着放牧强度的增加,群落平均盖度、高度和地上生物量呈降低趋势,物种丰富度指数、多样性指数及均匀度指数整体呈上升趋势。②土壤砂粒含量随放牧强度的增加呈上升趋势,表层土壤(0—10 cm)容重及入渗性能更易受放牧影响。③相同降雨强度下,随放牧强度的增加,土壤的侵蚀产沙量随之增加。土壤初渗速率、Simpson优势度指数和土壤砂粒含量是影响不同放牧强度下土壤侵蚀特征的关键因素。[结论] 放牧显著影响荒漠草原植被群落特征及土壤水文物理性能,因此,建议该区放牧强度控制在中度(2羊单位/hm²)及以下,对于受重度放牧破坏的草原区,围栏封育的同时还可适当进行人工补播或利用微生物改良土壤来提升土壤抗蚀性。     

肃北高寒草原不同放牧强度土壤养分变化特征

研究了肃北高寒草原不同放牧强度下不同土层土壤养分及 5 种微量元素有效态含量变化特征。结果表明：①高寒草原土壤物理性质的变化对土壤养分及微量元素具有重要的调控作用;②随着放牧强度的提高,0 ~ 10、10 ~ 20 cm 土层土壤体积质量均呈不同程度的增加,土壤孔隙度和土壤含水量则呈显著的递减趋势;③轻度放牧草地土壤有机质、土壤全 N 含量高于中度放牧和重度放牧草地;20 ~ 30 cm 土层有机质随放牧强度的增大呈明显下降趋势,即随放牧强度的增大深层土壤肥力呈退化趋势;肃北高寒草原的速效养分以多 N 少 P 富 K 为特点,土壤速效 N、P、K 含量在总体上随放牧强度的增加呈下降趋势;④肃北高寒草原 5 种微量元素的高低顺序依次是：Na＞Fe＞Mn＞Cu＞Zn,不同放牧强度下各微量元素的变化一致,顺序依次是：轻度放牧＞对照＞中度放牧＞重度放牧;⑤放牧强度对 10 ~ 20 cm 的土层影响最大,随放牧强度的增大,地表植物营养吸收层土壤营养成分、微量元素呈降低,导致地表植被生长能力降低,最终导致地表土壤沙化,最后使草地大面积退化。     

牧区土壤碳、氮的放牧反应

本研究把怀俄明州切恩附近高地４０年没有放牧的草场，采用三种管理制度，经过强度放牧（放牧６７天／ｈｍ２） １１年后，与连续轻度放牧（放牧２２天／ｈｍ２）和禁止放牧进行了比较。强度放牧时，年产草量的利用率不到５０％，与土地管理部门提出的水平相当。通过采集９１ｃｍ深的土样，来评估土壤有机碳、氮的变化，与禁牧区相比，放牧区地表下３０ｃｍ土壤的碳、氮含量比较高，而在３０ｃｍ以下土层，各种放牧条件下土壤的碳、氮含量相差不大。 ３／４的植物根部生物量存在于地表下３０ｃｍ的土层，该土层内的碳、氮活力最强。过去的１１年里，在这块混合大草原上采用各种放牧制度和放牧强度对土壤的有机碳、氮没有产生有害影响。该研究表明，用这些参数进行评估，合理放牧会全面提高土壤质量。     

放牧强度对草原砂质栗钙土某些物理性质影响的研究

作者测定了不同放牧强度下的草原砂质栗钙土机械组成、团聚体、水稳性团聚体、土壤结构系数及土壤微团聚体,并据此对该地区砂质栗钙土草原的合理放牧强度范围及安全放牧强度上限进行了探讨。作者建议以1.33~2.67只羊/hm2的放牧强度为该地区砂质栗钙土草原的合理放牧强度范围,但无论如何不应超过4.00只羊/hm2,超过此值,土壤中的一系列物理性状将显著恶化,草原土壤将出现明显的退化现象。     

黄土高原典型草原区草地土壤种子库的动态分析

采用野外调查取样和室内试验相结合的方法,研究了黄土高原典型草原区草地土壤种子库的物种组成、密度、时空格局和物种多样性等特征。结果表明,（1）土壤种子库中共统计到37个物种,隶属于16科,其中禾本科、菊科、唇形科、豆科和毛茛科的植物占优势。（2）从返青期到枯黄期,种子库中豆科、禾本科、堇菜科所占比例呈下降趋势,唇形科和菊科所占比例明显提高,种子库的密度也显著提高。（3）种子库中种子数量在垂直分布上呈现出由表层土壤向深层逐层减少的变化规律;在水平分布上的变化规律为：封禁〉放牧;阴坡〉阳坡;下坡〉中坡〉上坡。（4）封禁地土壤种子库中的Margalef丰富度和Shannon--Wienner指数都高于放牧地,Pielou均匀性指数低于放牧地。从返青期到枯黄期,封禁地和放牧地土壤种子库的Margalef丰富度、ShannonWienner指数、Pielou均匀性指数都呈下降趋势。     

放牧对植物群落特征和土壤微生物及酶活性的影响

[目的]探讨放牧对羊草群落和贝加尔针茅群落的影响,为草原土壤的保护和草原自然保护区建设提供科学依据。[方法]以呼伦贝尔草甸草原为样地,采用野外调查和实验室分析相结合的方法。[结果]在放牧干扰下,以羊草为建群种的群落所受的影响比贝加尔针茅群落小,植物群落盖度、高度和Simpson指数明显高于贝加尔针茅群,物种多样性更丰富,生长情况相对更好。羊草草原土壤全氮、全磷和有机质含量高于贝加尔针茅草原土壤。各放牧区土壤微生物数量均表现为:细菌放线菌真菌;垂直分布为0—10cm10—20cm;土壤脲酶、过氧化氢酶、转化酶活性、全N,全P和有机质等0—10cm土层的均高于10—20cm。相关分析表明,植物群落α多样性与土壤微生物数量、土壤酶活性、全N,全P和有机质等呈显著正相关。[结论]适度放牧有助于提高草甸草原羊草群落和贝加尔针茅群落α多样性、土壤微生物数量和土壤酶活性。两个群落相比较,羊草群落更适合放牧。     

希拉穆仁草原不同放牧强度对植被群落特征和土壤团聚体的影响

[目的] 为探究草原不同放牧强度下植被群落的改变,解析群落特征变化对土壤团聚体的影响。[方法] 以希拉穆仁荒漠草原控制放牧试验区为研究对象,采用野外调查、室内分析相结合的方法,分析轻度放牧(LG)、中度放牧(MG)、重度放牧(HG)和禁牧(CK)下植被群落特征和土壤团聚体特征及二者的关系,揭示植物群落变化对土壤团聚体的影响。[结果] (1)植物群落物种数随放牧强度的增加而增加,LG的群落Shannon-Wiener多样性指数(H'')和Simpson优势度指数(D)显著低于其他放牧强度(p＜0.05)。地上生物量随放牧强度的增加而降低,HG的地上生物量显著低于其他放牧强度(p＜0.05)。不同放牧强度下地下生物量差异显著,且不同土层的地下生物量均以LG最高。(2)平均重量直径(MWD)和几何平均直径(GMD)的变化趋势与大团聚体含量一致,在0—5, 5—10 cm土层表现为随放牧强度的增加先增加后降低再增加(LG最高、MG最低)。(3)群落Shannon-Wiener多样性指数(H'')、Pielous均匀度指数(J'')、地上生物量、容重是影响大团聚体含量、MWD和GMD的显著性因子(p＜0.05)。[结论] 放牧强度对土壤团聚体稳定性具有负反馈调节作用,主要通过影响植被群落多样性指数(H''和J'')、地上生物量、土壤容重的变化而引起土壤团聚体稳定性的变化。该结果为希拉穆仁草原放牧强度选择及生态恢复工作提供理论支撑和科学依据。     

旅游干扰对贺兰山典型草原生物多样性及土壤性质的影响

[目的]探讨旅游干扰对贺兰山典型草原生物多样性及土壤性质的影响,进一步理解旅游干扰的生态影响效应与机制,为风景区资源与环境的保护管理提供参考。[方法]采用样方法对贺兰山景区主要游览线路两侧植物群落及其土壤进行了调查,分析了不同强度旅游干扰(低度旅游干扰,中度旅游干扰,重度旅游干扰)对植物群落及其土壤性质的影响。[结果](1)旅游干扰显著降低了植被覆盖度、枯落物含量、丰富度指数、均匀度指数和多样性指数(p0.05);(2)旅游干扰对土壤理化性质具有较大影响,土壤容重随着干扰强度的增加而增加,含水量和电导率随着干扰强度的增加而降低;(3)在旅游干扰条件下,土壤微生物量碳和氮变化幅度较大,对旅游干扰表现最为敏感;(4)随着旅游干扰强度的增加,土壤有机碳、全氮、碱解氮和微生物量碳、氮均明显降低,全钾和有效钾随干扰强度的增加而增加;(5)旅游干扰区域土壤养分指标相关性基本与对照区域相一致,土壤全量养分均与其对应的有效养分呈极显著正相关(p0.01)。[结论]中度旅游干扰并没有影响植被属性;旅游干扰对土壤性质有一定影响,而此影响并没有改变土壤养分因子之间的相互作用效应。     

Integrated diurnal soil respiration model during growing season of a typical temperate steppe: Effects of temperature, soil water content and biomass production

Soil respiration was measured with the enclosed chamber method in an ungrazed Leymus chinensis steppe during the growing seasons of 2001 and 2002. Soil respiration rate (R_S) was significantly influenced by air temperature (T) at the diurnal scale, and could be described by Van't Hoff's equation (R_S = R₁₀ exp(β(T − 10))). At the seasonal scale, the normalized soil respiration rate at 10 °C (R₁₀) was mainly controlled by soil water content (R² = 0.717, P < 0.001), while the sensitivity of soil respiration to temperature (Q₁₀) was partially affected by absolute growth rate (R² = 0.482, P = 0.004). Thus, soil respiration could be described as R_S = (20.015W − 84.085) (0.103AGR + 1.786)^(T−10)/10 during the growing seasons, integrating soil water content (W) and absolute growth rate (AGR) into the temperature-dependent soil respiration equation. It was validated by the observed soil respiration rates in this study (R² = 0.890, P < 0.001) and observations from near-field experiment (R² = 0.687, P = 0.011). It implied that accurately evaluating annual soil respiration should include the effects of plant biomass production and other abiotic factors besides air temperature.     

Vegetation characteristics and soil properties in grazing exclusion areas of the Inner Mongolia desert steppe

In arid and semi-arid desert steppe areas, grazing exclusion with fencing is widely regarded as an effective strategy for restoring degraded vegetation and enhancing the quality of degraded soil. In this study, we hypothesized that grazing exclusion caused by fencing enhances both vegetation and soil properties, and that the longer an area is fenced, the more considerable the improvement. We conducted an observational study wherein random sampling was utilized to select 9 plots fenced for ten or more years, 25 plots fenced for four to nine years, 25 plots fenced for one to three years and 29 free-grazing plots within an area of approximately 63,000 km² of Inner Mongolia desert steppe. A one-way ANOVA revealed no significant differences in the characteristics of grassland vegetation or soil properties between grasslands fenced for one to three years and free-grazing grassland. After 4 years of fencing, noticeable increases in above-ground biomass, litter content, Simpson index, soil organic carbon, and available nitrogen were observed. Significant positive differences in vegetation coverage, height, species richness, soil available phosphorus, and available potassium were associated with plots with a minimum of 10 years of fencing. The soil layer with the greatest difference in the fenced-in areas for soil organic carbon was at 0–25 cm. For available nitrogen and available phosphorus, fencing produced the most significant differences in the 0–20 cm soil layer, while for available potassium, fencing produced the most significant differences in the 0–30 cm soil layer. However, the fencing did not indicate any statistically significant differences in terms of clay, silt, and sand content in any soil layer. The data support our hypothesis that grazing exclusion improves both vegetation and soil properties, and that longer periods of grazing exclusion result in greater degrees of improvement. This research offers technical guidance for the reasonable choice of fencing time across a vast area of the Inner Mongolian desert steppe.     

Partitioning soil respiration in a temperate desert steppe in Inner Mongolia using exponential regression method

Accurately partitioning soil respiration into autotrophic and heterotrophic components is important for understanding how ecosystem carbon budgets will respond to climate change. Usually, heterotrophic respiration can be estimated by a linear relationship between soil respiration and root biomass. In this study, however, we found that an exponential relationship was more appropriate than a linear relationship for relating soil respiration to root biomass in a temperate desert steppe in Inner Mongolia, China.     

Effects of livestock grazing on pollination on a steppe in eastern Mongolia

Widespread degradation of Mongolian grasslands by overgrazing is of global concern. The objective of this study was to reveal the effects of grazing on pollination as an example of interaction biodiversity in Mongolian grasslands. We established three plots according to grazing intensity on the eastern steppe of Mongolia. In each plot, we recorded the numbers of insect-pollinated plants and observed the foraging behavior of pollinators in June and August. The richness of insect-pollinated species was high and these species were most abundant in lightly grazed plots, and formed complex relations with diverse pollinators. But, frequency of flower visitation and pollination index were greater in heavily grazed plots. All pollination properties were poorest in intermediately grazed plots. These results suggest that the forb-biased foraging of sheep and goats reduces the floral diversity of insect-pollinated species, and consequently reduces pollinators in the intermediately grazed plots. In the heavily grazed plots, only limited ruderal species could survive under heavy cattle grazing, and such simple vegetation formed unbalanced but strong bonds with pollinators. Removal simulation showed that the mutual network was more fragile with respect to the extinction of certain species.     

放牧强度对中国内蒙古草原土壤水分状况与通量的影响

In the past few decades,the increase in grazing intensity has led to soil degradation and desertification in Inner Mongolia grassland,China,due to population growth and shift in the socio-economic system.Two sites with different grazing intensities,continuous grazing site(CG) with 1.2 sheep ha 1 year 1 and heavy grazing site(HG) with 2.0 sheep ha 1 year 1,were investigated at the Inner Mongolia Grassland Ecosystem Research Station(43 37 50 N,116 42 18 E) situated in the northern China to i) characterize the temporal distribution of soil water content along soil profile;and ii) quantify the water fluxes as affected by grazing intensity.Soil water content was monitored by time domain reflectometry(TDR) probes.Soil water retention curves were determined by pressure membrane extractor,furthermore processed by RETC(RETention Curve) software.Soil matric potential,plant available water and water flux were calculated using these data.Both sites showed an identical seasonal soil water dynamics within four defined hydraulic periods:1) wetting transition coincided with a dramatic water increase due to snow and frozen soil thawing from March to April;2) wet summer,rainfall in accordance with plant growth from May to September;3) drying transition,a decrease of soil water from October to November due to rainfall limit;and 4) dry winter,freezing from December to next February.Heavy grazing largely reduced soil water content by 43%-48% and plant available water by 46%-61% as compared to the CG site.During growing season net water flux was nearly similar between HG(242 mm) and CG(223 mm) sites between 5 and 20 cm depths.However,between 20 and 40 cm depths,the upward flux was more pronounced at HG site than at CG site,indicating that water was depleted by root uptake at HG site but stored at CG site.In semi-arid grassland ecosystem,grazing intensity can affect soil water regime and flux,particularly in the growing season.     

Effects of prescribed burning and seasonal and interannual climate variation on nitrogen mineralization in a typical steppe in Inner Mongolia

Fires in grasslands significantly alter nutrient cycling processes. Seasonal climatic changes can interact with fire to further modify nutrient cycling processes. To investigate the effects of fire on soil nitrogen transformation processes and their seasonal change and interannual variability in a typical steppe in Inner Mongolia, we determined the rates of net nitrogen mineralization and nitrification over two growing seasons and a winter following a prescribed spring fire in May 2006. Fire significantly decreased rates of both net nitrogen mineralization and net nitrification during the first growing season and winter following burning. Cumulative net nitrogen mineralization in unburned and burned plots in the 2006 growing season was 133% and 183% higher, respectively, than in the drier 2007 growing season. Nitrogen mineralization apparently occurred in winter and the cumulative net nitrogen mineralization from October 2, 2006, to April 27, 2007 in unburned and burned plots amounted to 1.18 ± 0.25 g N m⁻² and 0.51 ± 0.08 g N m⁻², respectively. Cumulative net nitrogen mineralization was higher in a wet 2006 than in a dry 2007 growing season, indicating that the net N mineralization rate was sensitive to soil moisture in a dry season. Our study demonstrated that a one-time prescribed fire decreased net N mineralization rates only for a short period of time after burning while interannual variation in climate had more significant effects on the process of nitrogen mineralization.     

Effects of cover crops on soil hydraulic properties during commodity crop growing season

Cover crops (CCs) can improve soil hydraulic properties prior to termination, but their effects on soil hydraulic properties during the growing season are less known. The objective of this study was to investigate the influence of no-till CC on the soil hydraulic properties during the commodity crop growing season in Murfreesboro, USA. The CCs included hairy vetch (Vicia villosa Roth.), crimson clover (Trifolium incarnatum L.), winter wheat (Triticum aestivum L.), winter peas (Lathyrus hirsutus L.), oats (Avena sativa), triticale (Triticale hexaploide Lart.), barley (Hordeum vulgare L.) and flax (Linum usitatissimum L.). The cash crop grown was corn (Zea mays). Soil samples were collected using a cylindrical core (55 mm inside diameter, 60 mm long) at 0–10, 10–20, and 20–30 cm depths during April (prior to CC termination), May, June and July. Results showed that soil bulk density (Db) was 23%, 12%, 11% and 10% higher under no cover crop (NCC) compared with CC management during April – July, respectively. This suggests a lower rate of soil consolidation under CC management even after several rainfall events. Four months after CC termination, macroporosity and total porosity were 306 and 50% higher, respectively, under CC compared with NCC management. Therefore, saturated hydraulic conductivity (K_sat) during July was two times higher under CC management compared with NCC management and this can affect increase water infiltration and conservation during the growing season. Due to CC root-induced improvement in macroporosity, CCs had 64% higher volumetric water content (θ) at saturation during July compared with NCC management. Cover crops can improve soil hydraulic properties and these benefits can persist for up to four months after termination.     

放牧和刈割对草甸草原中小型土壤动物群落的影响

为了解放牧和刈割对草甸草原中小型土壤动物群落结构的影响,于2019~2020年在内蒙古呼伦贝尔市鄂温克旗贝加尔针茅草甸草原采用自由放牧、刈割和围封3种不同草地利用方式下,研究了内蒙古草甸草原中小型土壤动物群落结构特征。调查研究在草地返青期、生长期和枯黄期3个时期进行,共捕获中小型土壤动物284只,隶属于12目34个类群。结果表明:放牧导致中小型土壤动物数量、类群数和多样性指数降低,而刈割影响相对较轻;返青期、生长期、枯黄期对应的春、夏、秋三季中小型土壤动物群落结构存在明显差异,土壤动物个体数、类群数和多样性指数均在草地枯黄期最多。放牧和刈割利用方式下改变了草甸草原中小型土壤动物在草地生长期Shannon-Weiner指数、Margalef指数、Pielou指数的变化趋势。中小型土壤动物群落表聚性在刈割利用方式中表现最为典型,放牧利用方式下0~10 cm土层中小型土壤动物有向10~20 cm 土层移动趋势;土壤pH和速效钾含量与中小型土壤动物类群有密切关系。     

Arbuscular mycorrhizal fungi associated with wild forage plants in typical steppe of eastern Inner Mongolia

A preliminary investigation was conducted on the arbuscular mycorrhizal (AM) status of the dominant and common wild forage plants in typical steppe of eastern Inner Mongolia, a major semi-arid grassland region in China. Fifty-four wild forage plant species were collected and examined, and 27 of these were colonized by AM fungi. Some plants belonging to families that are presumed to lack mycorrhizas (Cyperaceae, Caryophyllaceae and Chenopodiaceae) were also found to be mycorrhizal. Higher proportions of arbuscular mycorrhizal plants were found in perennial (56.1%) and monocotyledonous (64.7%) forage species. However, neither percentage of root length colonized nor spore density varied significantly between the two life forms or cotyledon types. Twenty-seven species belonging to 7 genera of AM fungi were identified in total according to the morphological characteristics of the spores from field soil and trap cultures, and the results indicate that Glomus was the dominant AM genus and Glomus geosporum (Nicolson & Gerdemann) Walker and Glomus mosseae (Nicolson & Gerdemann) Gerdemann & Trappe were the dominant species in field soil and trap cultures, respectively. Glomus intraradices Schenck & Smith, Glomus etunicatum Becher & Gerdemann, Glomus claroideum Schenk & Smith emend Walker & Vestberg, Glomus clarum Nicolson & Schenck and Scutellospora callospora (Nicolson & Gerdemann) Walker & Sanders also occurred with high frequencies.