A habitat model for brown bear conservation and land use planning in the central Apennines

We investigated the brown bear habitat suitability in an 8000 km² study area encompassing Abruzzo, Latium, and Molise regions in central-southern Italy. Based on long-term field surveys and published records, we classified bear habitat as occupied or unoccupied in 92 out of 320 sample squares (5 × 5 km). For each sample square 36 habitat variables were measured from topographic maps and Corine land-cover III level digital maps. The influence of habitat features on bear presence was investigated by multivariate and one-way analyses of variance and by logistic regression analysis. The logistic model correctly classified 95.5% of sample squares of bear presence and 93.8% of those where bears were absent. Average altitude, deciduous woodlands and ecotone length, showed a positive relationship with bear presence, whereas vineyard-olive groves and shrublands were negatively correlated with bear presence. No specific land management guidelines or strategies exist for bear conservation in central Italy, based on knowledge of habitat-population relationships. The landscape scale habitat model we developed could be useful to predict bear occurrence, to identify critical areas for a brown bear conservation strategy, and to enhance the arrangement of the protected areas network for the conservation of this species.     

Role of human-modified habitat in protecting specialist species: A case study in the threatened Florida Scrub-Jay

Habitat specialists are especially vulnerable to habitat alterations in vegetation structure and composition, making them difficult to protect and restore amidst agricultural landscapes. Protection strategies in such cases require information on species’ ability to survive and reproduce on marginal and modified habitat. We examined reproductive success, survival, and foraging efficiency of the Florida Scrub-Jay (Aphelocoma coerulescens), an extreme habitat specialist, in a human-modified habitat - regenerating pasture - along a pasture-native scrub interface. From 1985 to 2003, Florida Scrub-Jays were equally successful at producing young in regenerating pasture and native scrub. Production of eggs, nestlings, fledglings, independent young, and yearlings were not statistically different between territories containing pasture and those containing only scrub. Similar trends were observed for nest success and survival. When in pasture, individual jays were significantly more efficient at capturing small prey items, but significantly less efficient at capturing medium and large prey items. Availability of small prey items was significantly higher in pasture than in scrub; availability of medium and large prey items was not significantly different between habitat types. We conclude that regenerating pasture provides suitable supplemental habitat for Florida Scrub-Jays when in close proximity to native scrub, suggesting a novel conservation strategy for maintaining and potentially increasing local populations in fragmented agricultural settings. This case study illustrates how a broadened definition of ‘suitable habitat’ may augment existing conservation strategies for specialist species facing substantial and rapid modification of their native habitat.     

Experimental establishment of a declining dry-grassland flagship species in relation to seed origin and target environment

Supporting species persistence may involve (re)connecting suitable habitats. However, for many declining species habitat suitability and drivers of establishment are poorly known. We addressed this experimentally for a declining flagship species of dry grasslands in Germany, Armeria maritima subsp. elongata. In three regions, we sowed seeds from each of eight source populations back to their origin and to eight apparently suitable, but currently unoccupied, habitats close to the source populations. Overall, seeds germinated and seedlings established equally well in occupied and potential sites indicating that suitable habitats are available, but lack seed input. Germination and establishment varied among sowing sites. Moreover, seeds from populations of lower current connectivity established less well in new sites, and establishment was more variable among seeds from smaller than from larger populations, possibly reflecting genetic consequences of habitat fragmentation. Further, establishment across different new environments differed between seeds from different populations. As this was neither related to a home-away contrast nor to geographic or environmental distance between sites it could not clearly be attributed to local adaptation. To promote long-term persistence within this dry-grassland meta-population context we suggest increasing the density of suitable habitats and supporting dispersal connecting multiple sites, e.g. by promoting sheep transhumance, to increase current populations and their connectivity, and to colonise suitable habitats with material from different sources. We suggest that sowing experiments with characteristic species, including multiple source populations and multiple recipient sites, should be used regularly to inform connecting efforts in plant conservation.     

Species groups occupying different trophic levels respond differently to the invasion of semi-natural vegetation by Solidago canadensis

We studied the impact of the invasive plant species Solidago canadensis on the species richness of vascular plants and the abundance, species richness and diversity of butterflies, hoverflies and carabid beetles in herbaceous semi-natural habitats near Ljubljana, Slovenia. The species groups were sampled in sites dominated by S. canadensis and paired nearby sites covered by semi-natural vegetation. Plant species richness and species richness, abundance and diversity of butterfly species were lower in plots dominated by S. canadensis. Hoverfly abundance, diversity and species richness were negatively affected only in July just before the onset of flowering of S. canadensis, but tended to be positively affected in August during the height of flowering of S. canadensis. Only the abundance of carabid beetles was reduced in plots dominated by S. canadensis. The responses of the insect groups seem largely driven by the effects of Solidago on the availability of essential resources like food or larval host plants. Our results suggest that insect species that are closely related to plant species composition are more vulnerable to the effects of invasive plant species than those that are loosely or only indirectly related to plant species composition.     

Impacts of land use, anthropogenic disturbance, and harvesting on an African medicinal liana

African medicinal plant species are increasingly threatened by overexploitation and habitat loss, but little is known about the conservation status and ecology of many medicinal species. Mondia whitei (Apocynaceae, formerly Asclepiadaceae), a medicinal liana found in Sub-Saharan Africa, has been subject to intensive harvesting and habitat loss. We surveyed M. whitei in Kakamega Forest, the largest of three remnant Kenyan forests known to contain the species. In 174 100 m² plots, we quantified the status of M. whitei and investigated its relationships with land use, disturbance and harvesting. With average adult densities of 101 plants/ha, M. whitei is not locally rare in Kakamega. However, the absence of flowers and fruits, together with a spatial disconnect between adults and juveniles, suggests that sexual regeneration is patchy or infrequent. Comparing among habitat types, we found that plants were most abundant in regenerating indigenous forest managed by the Forest Department, which permits some extractive uses. Conversely, plants were largest in indigenous forest managed by the Kenya Wildlife Service, which prohibits extractive uses. Most anthropogenic disturbances were not associated with M. whitei, but plant occurrence and density were higher along paths used by livestock than along other types of paths. Larger individuals appeared to be preferentially harvested, but adult plants were more likely to occur in harvested plots than un-harvested plots. This work emphasizes that restrictions on disturbance and extractive use do not automatically promote medicinal plant conservation. Moreover, harvesting may have important genetic and demographic consequences that are overlooked by studies focused on numerical losses.     

Reconciling the dichotomy between single species and ecosystem conservation: black kites (Milvus migrans) and eutrophication in pre-Alpine lakes

In recent times there has been a growing dichotomy between preservation of single species and broader, ecosystem-based approaches to conservation. Freshwater habitats are among the most highly human-impacted ecosystems. We used a long-term data set on black kites (Milvus migrans), a threatened raptor dependent on aquatic habitats, to explore ways to reconcile single species and biodiversity-driven approaches to conservation. In the pre-Alpine lakes of northern Italy, black kite populations showed medium-low density, extremely low breeding success and widespread declines. Spatio-temporal variations showed density and breeding success to be positively related to ecosystem productivity (as estimated by phosphorus concentrations in lakes), availability of aquatic habitats and grassland, and to be negatively related to extent of farmland and fish harvest by professional fishermen. Fish species richness, used as a surrogate of biodiversity, was highest in one oligotrophic lake, but on average increased with increasing ecosystem productivity (i.e. lake eutrophication). Given expected future declines in ecosystem productivity, kite conservation will be helped by enhancing populations of alternative prey in terrestrial habitats (e.g. through incentives for grassland), and higher regulation of fish harvest. On the other hand, ecosystem management may profit from the use of black kites or other aerial piscivores as indicators of biodiversity and of diffuse ecosystem stress, such as sustainability of fishing practices.     

Habitat use and spatial dynamics of takhi introduced to Hustai National Park, Mongolia

The successful introduction of captive bred takhi or Przewalski’s horse, Equus ferus przewalskii, into Mongolia in the 1990s is a good example of the benefits of ex situ conservation and one of the few examples of the recovery of an animal after it became extinct in the wild. This is also particularly interesting because virtually nothing was known about how takhi lived before they died out, and the introductions have enabled us to study how they have settled, and their ecology and behaviour within their former natural range. In this paper, we describe the movement, home range size and shape, and habitat use of takhi at one of the release areas, the 570 km² Hustai National Park in Mongolia. Harem home ranges varied between 129 and 2399 ha, with 80% core areas of between 61 and 1196 ha. There was no relationship between range size and harem size, or length of time since release. Initially, harems stayed near their release enclosures, but over time they established home ranges further away. There was little overlap between home ranges of different harems, but neither was there evidence of exclusive range use. The more nutritious vegetation at lower elevations was preferentially selected. Thus the present situation looks good, but, as the population continues to grow, we anticipate that there will be potential problems related to intraspecific competition for water and vegetation resources, and the potential for hybridisation with domestic horses belonging to the local people. We consider the time it may take for takhi to reach carrying capacity within Hustai National Park and emphasise that continual monitoring of the population is essential because interventional management is likely to be required in the future.     

Habitat selection of endangered and endemic large flying-foxes in Subic Bay, Philippines

Large flying-foxes in insular Southeast Asia are the most threatened of the Old World fruit bats due to high levels of deforestation and hunting and effectively little local conservation commitment. The forest at Subic Bay, Philippines, supports a rare, large colony of vulnerable Philippine giant fruit bats (Pteropus vampyrus lanensis) and endangered and endemic golden-crowned flying-foxes (Acerodon jubatus). These large flying-foxes are optimal for conservation focus, because in addition to being keystone, flagship, and umbrella species, the bats are important to Subic Bay’s economy and its indigenous cultures. Habitat selection information streamlines management’s efforts to protect and conserve these popular but threatened animals. We used radio telemetry to describe the bats’ nighttime use of habitat on two ecological scales: vegetation and microhabitat. The fruit bats used the entire 14,000 ha study area, including all of Subic Bay Watershed Reserve, as well as neighboring forests just outside the protected area boundaries. Their recorded foraging locations ranged between 0.4 and 12 km from the roost. We compared the bats’ use to the availability of vegetative habitat types, riparian areas, and bat trees. The fruit bats’ locations showed a preference for undisturbed forest types and selection against disturbed and agricultural areas. Bat locations also showed selection for particular fruiting/flowering bat trees. The bats showed strong preference for riparian areas; locations were in riparian areas over four times more than expected. From these results we recommend that management focus flying-fox conservation efforts on undisturbed forest and riparian areas.     

‘Ecological forestry’ and eucalypt forests managed for wood production in south-western Australia

The model of ‘ecological forestry’ has evolved as a part of the development of the concept of ecosystem management. ‘Ecological forestry’ emphasises that manipulation of a forest ecosystem should consider, and as far as practicable work within the limits of, natural disturbance patterns prior to extensive human alteration of the landscape. This paper evaluates the extent to which forest management practices in jarrah (Eucalyptus marginata) and karri (Eucalyptus diversicolor) forests of south-western Australia align with this view of the characteristics and appropriate silviculture of ‘ecological forestry’. Characteristics and appropriate silviculture of ‘ecological forestry’ are evaluated in relation to (i) the stand level decisions of stand structure and harvest timing and (ii) the landscape level decisions of harvest levels and age structures, and spatial patterns of harvest. Forest management in south-western Australia is found to align with appropriate silviculture under this model of ‘ecological forestry’. Additionally, the landscape triad of areas managed to ‘ecological forestry’ principles, conservation reserves and areas managed to production forestry is in place in the south-western forests of Australia. Strengths and weaknesses in the model of ‘ecological forestry’ and the ability to interpret consistency of practices in the forests of south-western Australia with the characteristics of ‘ecological forestry’ are identified.     

Inferring habitat-suitability areas with ecological modelling techniques and GIS: A contribution to assess the conservation status of Vipera latastei

Some snakes are highly vulnerable to extinction due to several life history traits. However, the elusive behavior and secretive habits of some widespread species constrain the collection of demographic and ecological data necessary for the identification of extinction-prone species. In this scenario, the enhancement of ecological modelling techniques in Geographical Information Systems (GIS) is providing researchers with robust tools to apply to such species. This study has identified the environmental factors that limit the current distribution of Vipera latastei, a species with secretive behavior, and has evaluated how human activities affect its current conservation status, identifying areas of best habitat suitability in the Iberian Peninsula. Ecological-niche factor analysis (ENFA) indicated low marginality (0.299) and high tolerance (0.887) scores, suggesting strong tendency for the species to live in average conditions throughout the study area and to inhabit any of the environmental conditions. The analysis also revealed that this viper tends to select particular Mediterranean habitats, although topographic factors (altitude and slope) were the major environmental constraints for the Iberian distribution pattern of the species. The presence of other parapatric viper species in the north of the Iberian Peninsula (V. aspis and V. seoanei) and two human-related variables (landscape transformation and human density) also had a negative relation with the occurrence of V. latastei. All factors can explain its absence in northern Iberia and its fragmented distribution as currently is found mostly in mountains and relatively undisturbed low-altitude areas. The historical destruction and alteration of natural Mediterranean habitats and several life-history traits of the species contribute to its vulnerability to extinction. The ENFA analysis proved to be an outstanding method to evaluate the factors that limit the distribution range of secretive and widespread species such as V. latastei, updating evaluation of their conservation status.     

Using habitat models to determine conservation priorities for pond-breeding amphibians in a privately-owned landscape of northern Idaho, USA

As rural landscapes experience increasing levels of residential development, the persistence of species that rely on them will depend on informed planning and management decisions. An understanding of habitat requirements is essential for setting priorities and developing landscape-level plans for the survival of these species. In many temperate rural landscapes, artificially created ponds may be the only wetlands available for aquatic reproduction by amphibians. The introduction of non-native fish into these ponds reduces survival and prohibits successful reproduction of many native pond-breeding amphibians. We surveyed 105 randomly selected wetlands in a primarily privately-owned, rural landscape in north Idaho, USA, for pond-breeding amphibian larvae in 2004 and 2005. We used an information theoretic multimodel inference and an algorithmic (random forests) approach to model habitat for each species based on local and landscape characteristics. We also used a mail survey to quantify how landowners value fish in their wetlands and their plans for future wetland development and fish stocking. Sixty-seven percent of pond owners reported that fishing in their pond was at least slightly important to them and 36% of owners indicated that they were at least 50% likely to add fish to their ponds in the next 5-10 years. Landscape change predictions for this area indicate that forests will become more open due to thinning; habitat models indicated that this is likely to be detrimental to long-toed salamanders and beneficial to Pacific treefrogs. Habitat models also indicate that Columbia spotted frog breeding sites consist of wetlands on flat ground with high solar insolation and that this species is sensitive to nearby development, indicating that as this landscape becomes further developed, this species may require habitat protection for persistence.     

Habitat use and preliminary demographic evaluation of the critically endangered Mediterranean monk seal (Monachus monachus) in the Cilician Basin (Eastern Mediterranean)

Over the last few decades a dramatic decline in the number of Mediterranean monk seals (Monachus monachus) limited scientific studies on the biology, behaviour, and ecology of the species. This lack of information impaired the effectiveness of conservation strategies. Thus, any further information gathered on the monk seal has utmost value in the work to halt and reverse the plight of the species. A hitherto unknown Cilician Basin colony of the Mediterranean monk seal has been investigated during a 6-year period between 1995 and 2001. Results of direct in-cave surveys and land based seal-watch observations enabled the evaluation of habitat use and preliminary demographic information. A total of 39 caves were discovered among which only three were used for breeding. Scarcity of the breeding caves emphasised the importance of breeding habitat in the survival of the colony. Common features of the breeding caves are found as an entrance with a protective barrier against strong waves; a chamber with a wide beach; and a well-sheltered shallow pool. Supplementary data obtained from infrared monitors installed in three actively used caves were used to assess the number of individuals. Throughout the study period, 25 individuals were identified and 11 newborn pups were found. Evaluation of the results suggests the presence of sub-regions inhabited by an individually identified sub-group of monk seals. Recommendations for conservation guided by this information is a Marine Protected Area consisting of two zones; a restricted core zone prohibiting any human activity securing breeding habitat and a restricted fishery zone securing sustainability of the food source.     

Habitat requirements of passerines and reedbed management in southern France

Reedbeds have high conservation value in Europe. In southern France, they are the major breeding habitat of five passerine species. Yet, habitat management is done primarily by water control to serve socio-economic rather than conservation interests, because we lack information on the species' ecological requirements. Determinants of passerine abundance were assessed through a comparative analysis of water regime, plant structure, and arthropod (food) distribution at 12 sites consisting of at least 10 ha of marsh densely covered with common reed (Phragmites australis). Overall bird abundance estimated through standardised mist netting was positively correlated with food availability (sweep-netted arthropods weighted by their occurrence in birds' diet), which was in turn negatively correlated with duration of ground dryness between June and December. Abundance of four of the five bird species was associated with specific vegetation parameters (reed diameter, dry reed density, growing reed height, etc.), which could be associated with particular management practices, especially with regard to water levels and salinity. Potential impact of socio-economic activities through their water management is addressed, as well as possible ways to minimise these impacts.     

Sampling on private property to evaluate population status and effects of land use practices on the gopher tortoise, Gopherus polyphemus

Although private properties are predicted to play an increasingly significant role in conservation, surveys of species of special concern are rare on these lands. We created a template for a multi-county survey of randomly selected sites and sampled for burrows of the gopher tortoise (Gopherus polyphemus) in south-central Georgia, USA. Current land use was strongly correlated with tortoise population condition. The highest densities of tortoise burrows were found on lands with open-canopied pine stands that were managed with prescribed fire, a practice associated with types of selection forestry and/or wildlife management. Agricultural sites and unburned areas provided poor habitat and pine plantations were only slightly better. Our estimates of tortoise population densities indicated that the current landscape supports less than 20% of the animals present before implementation of modern land use practices. In addition, our estimate for density of active burrows was approximately one third of that projected for the entire state range 20 years ago by Auffenberg and Franz [Auffenberg, W., Franz, R., 1982. The status and distribution of the gopher tortoise (Gopherus polyphemus). In: Bury, R.B. (Ed.), North American Tortoises: Conservation and Ecology (US Fish and Wildlife Service Wildlife Research Report 12). pp. 95-126]. However, some good sites for gopher tortoises remain in south Georgia and our data also suggested that extraordinary conservation actions may not be required if ways can be developed to retain traditional land management practices on private property.     

Lignin and cellulose degradation and nitrogen dynamics during decomposition of three leaf litter species in a Mediterranean ecosystem

Cellulose and lignin degradation dynamics was monitored during the leaf litter decomposition of three typical species of the Mediterranean area, Cistus incanus L., Myrtus communis L. and Quercus ilex L., using the litter bag method. Total N and its distribution among lignin, cellulose and acid-detergent-soluble fractions were measured and related to the overall decay process. The litter organic substance of Cistus and Myrtus decomposed more rapidly than that of Quercus. The decay constants were 0.47 year⁻¹, 0.75 year⁻¹ and 0.30 year⁻¹ for Cistus, Myrtus and Quercus, respectively. Lignin and cellulose contents were different as were their relative amounts (34 and 18%, 15 and 37%, 37 and 39% of the overall litter organic matter before exposure, for Cistus, Myrtus and Quercus, respectively). Lignin began to decrease after 6 and 8 months of exposure in Cistus and Myrtus, respectively, while it did not change significantly during the entire study period in Quercus. The holocellulose, in contrast, began to decompose in Cistus after 1 year, while in Quercus and Myrtus immediately. Nitrogen was strongly immobilized in all the litters in the early period of decay. Its release began after the first year in Cistus and Myrtus and after 2 years of decomposition in Quercus. These litters still contained about 60, 20 and 90% of the initial nitrogen at the end of the experiment (3 years). Prior to litter exposure nitrogen associated with the lignin fraction was 65, 54 and 37% in Cistus, Myrtus and Quercus, while that associated with the cellulose fraction was 30, 24 and 28%. Although most of the nitrogen was not lost from litters, its distribution among the litter components changed significantly during decomposition. In Cistus and Myrtus the nitrogen associated with lignin began to decrease just 4 months after exposure. In Quercus this process was slowed and after 3 years of decomposition 8% of the nitrogen remained associated with lignin or lignin-like substances. The nitrogen associated with cellulose or cellulose-like substances, in contrast, began to decrease from the beginning of cellulose decomposition in all three species. At the end of the study period most of the nitrogen was not associated to the lignocellulose fraction but to the acid-detergent-soluble substance (87, 88 and 84% of the remaining litter nitrogen).     

Density and distribution of the endemic Seram cockatoo Cacatua moluccensis in relation to land use patterns

The endemic Seram cockatoo, Cacatua moluccensis, was placed on Appendix I of CITES in response to declining trade statistics but in the absence of population data. We conducted population surveys and collected data on habitat structure at seven sites on Seram in 1998. Cockatoo densities ranged from 0.93 to 17.25 birds/km² and averaged 7.9 birds/km² across sites. We classified habitat into three types but found that cockatoo densities did not correspond closely to habitat differences across sites. Cockatoo abundance was significantly associated with presence of potential nest trees and strangling figs. Analysis of forest cover and landuse indicates that while most of the island is still covered in lowland forest, only 14% of these forests are protected and almost half the island is classified as logging concession. Conservation recommendations include better information and enforcement of laws protecting Seram cockatoos and resolution of boundary conflicts between parks and logging concessions.     

The effects of land use and climate on red kite distribution in the Iberian peninsula

The main breeding populations of the red kite (Milvus milvus L.), have been declining in the Iberian peninsula during the last decade. However, there is a lack of regional assessments of habitat suitability that identifies limiting ecological factors for the species and areas with conservation problems. In this work we present a regional model for the distribution and abundance of breeding red kites in the Iberian peninsula. The occurrence and estimated abundance in 100 km² UTM squares resulting from road censuses were modelled with broad-scale explanatory variables obtained from satellite imagery, thematic digital cartography, climatic data and spatial coordinates. The occurrence model incorporated mainly climatic variables and had a good discrimination ability, while the abundance model incorporated mainly land-use variables and had a lower explanatory power (r²=0.14). The predictions somewhat overestimated the results of the censuses, and this agrees with the decline of population size and range observed for this species in the Iberian peninsula. These models are relevant in the conservation of the species: first, they suggest the limiting factors for red kite in the Iberian peninsula, and, second, they generate predictive maps pointing out both areas in which conservation problems may be acute (suitable locations that are unoccupied), and areas where no data is available but the red kite is likely to be present (thus guiding further survey and research).     

Soil and climate are better than biotic land cover for predicting home-range habitat selection by endangered burrowing owls across the Canadian Prairies

Statistical models that describe species-environmental relationships are important components within many wildlife conservation strategies. These models are typically developed from studies conducted on small geographic scales (hundreds of square kilometres), representing a relatively small range in environmental conditions. Such local models from local studies are often then extrapolated to predict the suitability of other unsampled regions. The value of many models would be increased by considering larger-scale processes that might be structuring spatial patterns across species distributions. We examined home-range habitat selection by burrowing owls throughout the mixed prairie grassland region of western Canada (180,000 km²) to determine whether owl selection for biotic factors changes along abiotic gradients. Specifically, we classified 37 explanatory variables into five categories (geography, grassland fragmentation, land-use, soil, and climate), created models for each set of variables, and evaluated the predictive ability of each model. We then examined interaction effects to determine if the relationship between land cover variables and the probability of owl home-range selection varied within large-scale abiotic criteria. Our results showed that soil and climate produce the most predictive models of burrowing owl home-range selection and create unique environmental conditions for owls which are independent of land cover at this scale. This study provides new insight into burrowing owl habitat requirements, and strengthens the case for considering large-scale abiotic gradients when prioritizing areas for species conservation.     

Habitat selection of the Eurasian woodcock in winter in relation to earthworms availability

The Eurasian woodcock (Scolopax rusticola) is a game species experiencing high hunting pressure, long-term modifications of its habitats, and with questions regarding its current conservation status. Winter is a season of highest concentration of birds and hunting pressure but woodcock precise habitat requirements are poorly known. It is crucial to assess threats and to develop sustainable management options for the conservation of woodcock populations. During three consecutive winters, we monitored 65 individual woodcocks fitted with radio-tags in Brittany, France. Habitat selection was analysed using GIS and compositional analysis, in relation to vegetation types, soil variables (humus types) and the abundance of their main prey (earthworms). Woodcocks used different habitats diurnally and nocturnally, generally preferring areas with high earthworm biomass. Diurnal habitat selection in forests was associated with humus type (preference for mulls, rich in earthworms) and dense shrub strata (better protection). Hedges with a high density of trees and shrub were also important habitat. At night, grazed meadows were the preferred habitat, containing five times higher biomass of earthworms compared to cultivated fields. Sustainable management of populations requires protection and management of habitats that incorporates food and cover. Forestry practices should preserve rich humus types and coppices by choosing tree species that ameliorate the soil and soil tilling. Changes in landscapes and intensive agricultural practices are current threats to woodcock populations: destruction of hedges, decrease of permanent grazed meadows, impoverishment of soils fauna biomasses from ploughing and chemical applications. However, woodcocks may benefit from the recent development of set-asides, grass field-borders and simplified farm practices (no-tillage and direct sowing).