Development of lowland weed management and weed succession in Japan

Since the introduction of rice production in Japan, lowland areas have been managed for rice production with the purpose of better rice growth, as well as lesser weed infestation. Rice is cropped every year in lowland fields by repeated cultivation of a single crop, with high yields and without soil sickness usually being observed in upland fields. This is probably because the irrigation water supplies various nutrients for healthy rice growth and the drainage washes out and removes harmful factors. However, until recently, the wet or flooded conditions of lowland fields in the Asian monsoon region never have allowed humans to cultivate useful summer crops, except rice or some aquatic plants. Therefore, the management of lowland areas in the Asian monsoon region has been significantly different from European field management, where crop rotation has been the traditional standard practice. Paddy weeds are aquatic plants or hygrophytes that have adapted to lowland fields. Traditionally, tillage and puddling were practiced seasonally in lowland fields on a regular schedule every year. Rice cultivation technology was developed and supported by regional irrigation systems that created stable environments for typical paddy weeds to complete their life cycle. After the introduction of chemical weed control, rice fields became very severe habitats for these paddy weeds, where they could not grow and reproduce without strategies for survival under herbicide exposure. Even so, many of the traditional paddy weeds survived because of their accumulated or uneradicated seed banks, although several aquatic plants were listed as endangered or threatened species. The important weed species changed, sometimes rapidly and sometimes slowly, depending both on their reproductive system and their biological response towards field management and weed control systems. Very recently, the level of perennial weeds, herbicide‐resistant weeds, and weedy rice has increased in paddy fields that are highly dependent on herbicide use. In addition, several hygrophyte species have invaded paddy fields. In order to address these issues, the improvement and application of integrated weed management methods are expected to be critical.     

The effect of crop,cultivation and seed addition for birds on surface weed seed densities in arable crops during winter

Weed seeds present an agronomic threat, but are also an important food resource for wildlife in winter. Weed seed densities on the soil surface in winter were examined from 1999 to 2002 in 105 fields on three different farms in UK. The effect of the preceding crop, cultivation, position within the field and the application of seed for birds (bird seed) on surface seed abundance and species composition was tested. Six or fewer species comprised c. 80% of the weed seeds. By January of each study year, the densities of seeds important for farmland birds (key seeds) were 73% or 87% lower compared with early winter on two of the farms, but were stable on the third where seeds were incorporated through cultivation. At the edge and mid‐field, seed densities only exceeded 400 m^?2 in 17%, 10% and 12% of fields for total, key and dicotyledonous seeds respectively. The preceding crop only affected seed densities at one site; stubbles of winter barley had fewer seeds compared with winter wheat or spring barley. Seed densities varied between the edge and mid‐field, but trends were inconsistent between sites. The density of the larger seeds (Atriplex patula, Viola arvensis, Polygonum aviculare and Chenopodium album) were reduced in fields receiving bird seed. The objectives of weed control and conservation may not be mutually exclusive because seed return was most reduced where the ground remained uncultivated through the winter, yet this also provided the best foraging opportunities for surface feeding seed predators.     

Weed communities respond to changes in the diversity of crop sequence composition and double cropping

Agricultural intensification, besides increasing land productivity, also affects weed communities. We studied weed shifts in cropping sequences differing in the identity and number of crops grown. We also evaluated whether dissimilar weed communities in different cropping systems converge towards more similar communities, when the same sequence is cropped during 2 years. In three locations in the Rolling Pampa, Argentina, field experiments were conducted including five cropping systems in the first year (winter cereal/soyabean, field pea/soyabean, and field pea/maize double crops, and maize and soyabean as single crops), while the same sequence was grown in the following 2 years (wheat/soyabean double crop and maize). Changes in weed community composition and structure were analysed through multivariate analyses and frequency–species ranking plots. Weed communities differed first among sites, while weed shifts within each site were mainly associated with growing season and crop type. Differences among crop sequences were higher in the first year, mostly related to specific crop grown, rather than to the number of crops in the sequences. Differences were reduced when the same sequence was grown during two consecutive seasons. Frequency of highly common weeds was negatively associated with the number of days with high crop cover. Our findings contribute to understand weed shifts in consecutive growing seasons, which may help readapting crop sequences to reduce the occurrence of abundant weed species.     

Suppression of weeds and weed seeds in the soil by stubbles and no-tillage in an arid maize-winter wheat-common vetch rotation on the Loess Plateau of China

Reduced tillage provides ecological and economic benefits to arable land on the Loess Plateau of China, where soil erosion has long been a serious problem and soil water availability is largely restricted. However, high abundances of weeds in reduced tillage systems cause significant yield losses. In this study, we explored the effects of no-tillage and stubble retention on the number and density of weeds and weed seeds in a 12-year maize-winter wheat-common vetch rotation on the Loess Plateau. Four treatments including conventional tillage, no-tillage, conventional tillage+stubble retention and no-tillage+stubble retention were designed and applied. We found that no-tillage increased the number of weed species and weed density in most of the crops, while stubble retention decreased weed density in maize and tended to suppress weeds in both no-tillage treatments(no-tillage and no-tillage+stubble retention). No-tillage led to an increase in the number of weed species in the weed seedbank and tended to increase seed density during the spring growth of winter wheat, but it decreased seed density during post-vetch fallow. Stubble retention tended to reduce seed density during the spring growth of winter wheat and post-vetch fallow. We concluded that no-tillage can promote weeds in the experimental crop rotation, while stubble retention suppresses weeds in untilled fields. The combined effects of stubble retention and no-tillage on weed suppression varied among the three crops. Based on these results, we recommend stubble retention in untilled legume-crop rotations on the Loess Plateau to improve the control of weeds.     

南繁水稻田杂草发生与化学防治

在我国冬季南繁育制种作物中,水稻所占面积最大,但是由于不同地域杂草群落组成的差异,继续沿用原产地的除草剂品种和技术后,除草效果并不理想甚至会造成水稻药害。为了解决此类问题,连续3年对南繁区水稻田冬季杂草发生情况进行调查,明确现用地的杂草种类与发生规律,分别按照杂草群落中禾本科、阔叶杂草和莎草科主要危害的种类指导科学合理地选择除草剂种类,避免盲目用药,从而实现减药又高效的除草目标。     

Ecophysiological study on weed seed banks and weeds in Cambodian paddy fields with contrasting water availability

Weed infestations are a major cause of yield reduction in rice (Oryza sativa) cultivation, particularly with direct‐seeding methods, but the relationship between weed dynamics and water availability in Cambodian paddy fields has not been documented previously. We surveyed the weed abundance and weed seed banks in the soil of paddy fields with inferred differences in their water regime in 22 farm fields in three provinces of Cambodia in the 2005 and 2006 rainy seasons. We studied rain‐fed lowland fields in upslope and downslope topographic positions and fields at different distances from the irrigation water source inside an irrigation rehabilitation area. The weed seed banks were estimated by seedling emergence in small containers and weed abundance and vigor were estimated by a simple scoring system. The estimated weed seed bank in the top 5 cm of soil ranged from 52.1 to 167 × 10³ seeds m^?2 (overall mean of 8.5 × 10³ seeds m^?2) and contained a high proportion (86%) of sedge species, such as Fimbristylis miliacea L. and Cyperus difformis. Several fields had particularly large seed banks, including one near the reservoir. No clear difference was found in the weed seed banks between the irrigated fields that were located close to (upstream) and distant from (downstream) the water source or between the irrigated and rain‐fed lowland fields, but the weed scores were larger in the rain‐fed fields and the downstream fields within the irrigated area. A water shortage during the late growing season in 2005 led to a proliferation of weeds in some fields and an associated increase in weed seedbank size in 2006. However, the weed scores in 2006 were more strongly associated with that year's water conditions than with the weed seedbank size.     

Effect of tillage, cover crop and crop rotation on the composition of weed flora in a sandy soil

The development of integrated weed management strategies requires knowledge of mechanisms that influence compositional changes in weed flora. A 9-year study was initiated in 1988 at Delhi, Canada, on a loamy sand soil to evaluate the effect of tillage systems [conventional (CT) and no-till (NT)] and cover crops (only in NT) on weed density, species composition and associations, and crop yield in a winter wheat ( Triticum aestivum L.)/bean/winter wheat rotation. Three bean types: soyabean ( Glycine max L. Merr.), white bean ( Phaseolus vulgaris L.) and kidney bean ( P . vulgaris L.) were included. The NT system included variations: rye ( Secale cereale L.) or maize ( Zea mays L.) cover crop, volunteer wheat disked after harvest and wheat stubble. Data were collected in 1994, 1995 and 1996. Tillage systems, cover crops and crop type had differential effects on weed densities, species composition and associations. Weed densities were not affected by tillage or cover crops in wheat but, in the beans, densities were greater in the CT than in the NT systems. Various associations of weed species with tillage system, cover crop and crop type were observed. Crop yields were not affected by tillage type or cover crop, except that soyabean yields were highest in plots with cover crops.     

Development of chemical weed control and integrated weed management in China

More than 200 species of weeds are infesting main crop fields in China, among which approximately 30 species are major weeds causing great crop yield losses. About 35.8 million hectares of crop fields are heavily infested by weeds and the annual reduction of crop yields is 12.3–16.5% (weighted average). Along with rural economic development, approximately 50% of the main crop fields undergo herbicide application. Chemical weed control has changed cultural practices to save weeding labor in rice, wheat, maize, soybeans and cotton. At the same time, continuous use of the same herbicides has caused weed shift problems and weed resistance to herbicides. Consequently, integrated weed management in main crops is being developed.     

The impact of management on weeds and aquatic plant communities in Hungarian rice crops

This study assessed the cultural and weed management factors influencing the weed communities of Hungarian rice fields. Hungary is situated at the northern limit of rice production with a history of about 300 years of rice culture. We surveyed the weed flora and 25 background variables in 100 active rice fields. Using a minimal adequate model containing 11 terms, 48.5% of the total variation in weed species data could be explained. The net effects of nine variables on species composition were significant. Crop cover was found to be the most important explanatory variable, which was followed by the herbicides penoxsulam and azimsulfuron, tillage depth, phosphorous and potassium fertilisers, years after last rotation, water depth in May, sowing type, pendimethalin and water conductivity. Filamentous algae, as the most abundant group of weeds, were positively associated with deep tillage, deep water and surface sowing. Echinochloa crus‐galli, one of the most troublesome grass weeds, was associated with low rice cover, shallow water and later years after crop rotation, while weedy rice favoured high crop cover, deep water and soil sowing. These findings can be used to design improved weed management strategies. The occurrence of red list species and charophytes in diverse micro‐mosaic patterns deserves attention from a conservation perspective, as well. The maintenance of these unique charophyte communities can be facilitated by shallow tillage without soil inversion.     

Diversity in weed seed production and the soil seed bank: Contrasting responses between two agroecosystems

This study was carried out to compare the diversity in seed production and the soil seed bank in a dryland and an irrigated agroecosystem in the dry tropics. Both agroecosystems showed a comparable number of species, but only 25% and 38% similarity during the winter and rainy cropping seasons, respectively. In the irrigated agroecosystem, the amount of seed production diversity was almost double in the winter season, compared to the rainy season. The weed seedbank diversity was low but was sensitive to cropping practices and seasons in both agroecosystems. A considerably smaller soil seedbank size in the irrigated agroecosystem (cf. dryland) was related to lowered weed seed production. The dryland agroecosystem showed a greater accumulation of the seeds of broad‐leaved weeds, whereas the irrigated agroecosystem accumulated more seeds of the grasses or sedges. About three‐fourths of the seeds during the winter season were accounted for by Anagallis arvensis and Chenopodium album in the dryland agroecosystem and by C. album and Melilotus indica in the irrigated agroecosystem. However, during the rainy season, Ammannia baccifera, Echinochloa colona and Cyperus rotundus dominated in both agroecosystems. The changes in the weed seed bank and its diversity are mainly attributed to differences in water management, which tends to reduce species diversity, especially at a lower depth, but leads to the dominance of some potentially noxious weeds (e.g. Phalaris minor and M. indica). Approximately double the soil seedbank size and a greater diversity at a lower depth might indicate an adaptive mechanism in the storage of weed seeds in the dryland agroecosystem.     

Effects of human-mediated processes on weed species composition in internationally traded grain commodities

International trade is a major route by which non‐indigenous organisms are introduced into new habitats. Various kinds of weed seeds have been introduced through grain trade. The objectives of this study were to understand the factors that affect the initial assemblage of plant species introduced by the international grain trade and to extract their general attributes. We surveyed weed seed contamination of spring wheat imported from Canada to Japan and analysed the effects of the field abundance of each weed and of harvesting and cleaning on the quantity of weed seed included in the imported wheat. The field abundance was positively correlated with the weed seed quantity. Seeds of short weeds and seeds with a pappus were eliminated from the wheat by the harvesting or cleaning process. Many other crop plants contaminated the wheat. Because various transportation vehicles, temporary storage sites and port elevators are used commonly with all exported crops and it is difficult to remove all residues from them, other crops might be carried over into the wheat commodity. These relationships also apply to other grains.     

长江中下游小麦田杂草发生组成及群落特征

为明确我国长江流域冬小麦田杂草发生情况,采用倒置"W"方法取样对江苏、安徽和湖北省945个冬小麦田杂草进行了抽样调查.结果表明,长江流域冬小麦田共发现杂草93种(变种),隶属于28科72属,其中禾本科杂草最多,有15种;其次是菊科杂草有12种;蓼科杂草9种;十字花科7种;豆科和车前科各6种;石竹科5种.其中优势杂草有猪...     

An on-farm approach to investigate the impact of diversified crop rotations on weed species richness and composition in winter wheat

Weed species diversity may benefit from organic farming due to enhanced temporal diversification of crop species in a rotation and omission of herbicide applications. However, in intensively managed conventional systems, little evidence exists as to what extent diversified crop rotations contribute to higher weed species richness. Using an on-farm approach, the effect of crop rotation (organic, conventional diverse (CD) and conventional simple (CS) crop rotations) and weed control (with vs. without) on weed species richness, cover, community composition and crop biomass, was analysed in 24 winter wheat fields. Weed species with beneficial functions for invertebrates and birds were analysed separately. Weed species richness was higher in the organic crop rotation, but did not differ between CD and CS crop rotations. Weed control treatment reduced species richness in both conventional rotations, but not in the organic one. Redundancy analyses revealed that crop rotation intensity accounted for the largest part of the explained variation in weed species composition. Results from the study indicate that the maintenance of weed species richness and conservation of species with important ecological functions requires not only temporal diversification of crop species in the rotation, but also an adjustment of weed control strategies.     

Correlation between the soil seed bank and weed populations in maize fields

Annual weed populations establish every year from persistent seed banks in the soil. This 3 year study investigated the relationship between the number of weed seeds in the soil seed bank and the resultant populations of major broadleaf and grass weeds in 30 maize fields. After planting the crop, 1 m² areas were protected from the pre-emergence herbicide application. Soil samples were collected soon after spraying to a depth of 100 mm and the weed seeds therein were enumerated. The emerged weed seedlings in the field sampling areas were counted over the following 8 weeks. Up to 67 broadleaf species and five grass weeds were identified, although not all were found at every site and some were specific to a region or soil type. For the most abundant weeds in the field plots, on average 2.1–8.2% of the seeds of the broadleaf species and 6.2–11.9% of the seeds of the grass weeds in the soil seed bank emerged in any one year, depending on the species. Overall, the results showed a strong linear relationship between the seed numbers in the soil and the seedling numbers in the field for all the grasses and for most broadleaf weeds. For some species, like Trifolium repens , only a weak relationship was observed. In the case of Chenopodium album , which had the largest seed bank, there was evidence of asymptotic behavior, with seedling emergence leveling off at high seed numbers. An estimate of the soil seed bank combined with knowledge of the germination and behavior of specific weed species would thus have good potential for predicting future weed infestations in maize fields.     

麻类作物田杂草种类与防除技术

杂草常伴随麻类作物滋生,与其争光争肥争水,严重影响其生长、降低产量和品质。因此,有效防除麻田杂草尤为必要。文章概述了我国麻类作物生产现状,主要麻类作物田杂草危害、杂草种类及防除技术。     

Competition between weeds and wet season transplanted paddy rice for nitrogen use, growth and yield in the central and northern regions of Laos

To quantify the effect of weeds on the nitrogen (N) use, growth and yield of wet season paddy rice in the central and northern regions of Laos, we surveyed the paddy fields in these regions in October 1999 and November 2000. We found 13 weed species in total, but there were few major weeds abundant at the survey sites. In the infertile soils under rainfed conditions, weed growth was poor. Rough rice yield, the number of panicles, the number of seeds per square meter, the above-ground biomass of paddy rice and the amount of N accumulated in the above-ground biomass of paddy rice (amount of N in rice) were suppressed by competition with weeds. However, harvest index (HI) and nitrogen use efficiency (NUE) of paddy rice were not suppressed by competition with weeds. The amount of N in rice was suppressed by competition with weeds, the number of panicles decreased as the amount of N in rice decreased, and the number of seeds per square meter decreased as the number of panicles decreased. As a result, rough rice yield was suppressed by competition with weeds. The weeds competed with paddy rice for N uptake during the productive tillering stage. However, the ability of paddy rice to compete for N uptake with weeds was not reduced under rainfed lowland conditions. When the weeds were completely removed, the amount of N in rice increased. Rough rice yield may be increased by 10% under rainfed lowland conditions and by 17–19% under irrigated conditions.     

Arbuscular-mycorrhizal fungi: potential roles in weed management

The importance of interactions between arbuscular-mycorrhizal fungi (AMF) and weeds of agro-ecosystems is reviewed. Considerable evidence suggests that AMF can affect the nature of weed communities in agro-ecosystems in a variety of ways, including changing the relative abundance of mycotrophic weed species (hosts of AMF), and non-mycotrophic species (non-hosts). These effects may merely change the composition of weed communities without affecting the damage that these communities cause. However, it is quite plausible that interactions with AMF can increase the beneficial effects of weeds on the functioning of agro-ecosystems. Through a variety of mechanisms, weed:AMF interactions may reduce crop yield losses to weeds, limit weed species shifts, and increase positive effects of weeds on soil quality and beneficial organisms. If beneficial effects of AMF on the composition and functioning of weed communities can be confirmed by more direct evidence, then AMF could provide a new means of ecologically-based weed management. Intentional management will be required to increase diversity and abundance of AMF in many cropping systems, but these actions (e.g. conservation tillage and use of cover and green-manure crops) typically will confer a range of agronomic benefits in addition to potential improvements in weed management.     

Reducing intrarow weed numbers in row crops by means of a biennial cultivation system

Intrarow weed emergence in row crops can be reduced by means of a biennial cultivation system that is based on an exhaustion of the weed seedbank in the upper soil layer of crop‐free bands established in a preceding cereal crop, year I, before the row crop, year II. In this study, a series of field experiments is presented focusing more specifically on the influence of prevention of weed seed shedding and of different cultivation tactics in the system on intrarow weed emergence in the bands in year II. As the system excludes inverting soil tillage between years I and II, the prevention of weed seed shedding in year I turned out to be a key factor in achieving the desired effect of the system. This was seen with spring barley grown in year I, but not with winter wheat, where the system actually resulted in an increase in intrarow weed numbers in spite of the measures taken in year I. Cultivating the crop‐free bands in spring barley in year I with conventional goose‐foot shares in order to enhance seed germination and mortality further had no significant effects regardless of time and intensity. Results from two other experiments, in which freshly shed weed seeds were incorporated into the soil, supported the finding that cultivating the bands was of no advantage.     

Development of arable weed seedbanks during the 6 years after the change from conventional to organic farming

All of the arable land of a farm in southern Germany was investigated during an 8-year period, starting 2 years before and ending 6 years after the change to organic farming. The first 3 years after the conversion, the total seed number in the soil increased from 4050 to 17 320 m⁻². From the fourth to the sixth year, it dropped back to 10 220 m⁻². The number of seeds increased particularly at sites with a low crop cover and a high density of weed plants at the soil surface. The increase predominantly occurring on the more fertile soils may have been caused by the rotation commencing with less competitive crops. Crops which increased the seedbank by 30–40% were winter cereals, sunflowers and lupins. Potatoes and sown fallow caused no significant change and grass–clover mixtures even reduced the number of seeds by 39%. Among 44 species occurring frequently enough for statistical analysis, 31 increased and only 3 decreased. The change of management particularly increased summer annual, perennial and dicotyledonous weeds. This can be attributed to both operations which are characteristic of organic farming (e.g. replacing herbicide applications by mechanical weed control) and to general modifications of the management practice which may also occur in non-organic farming systems (increasing the percentage of broad leaved and spring sown crops in crop rotation). The present study confirms investigations into the aboveground vegetation that indicate that arable organic farming favours plant species diversity and provides evidence that the conversion need not encourage the dominance of a few noxious weeds.     

Increasing weed flora in Danish arable fields and its importance for biodiversity

A number of political initiatives have been implemented to meet growing concerns about negative side effects of the intensification of land‐use practice over the past 25 years. Declining species diversity and abundance and pollution of the environment have been reported worldwide. In order to assess the overall effect on wild flora of the changing management of arable land, national surveys are necessary. Surveys of the weed flora in Danish arable fields were conducted in 1987–89 and 2001–04. The frequency of common weed species in spring barley, spring rape, winter rye, winter wheat and grass leys was compared between time periods. Weed frequency increased overall during the interval. Species that are of particular importance as food sources for invertebrates and birds increased considerably in some of the crops. The frequency of some grass species increased remarkably, possibly because of increased areas of winter crops. Some perennial weeds also became more frequent, probably caused by less mechanical weed control and a reduction in the use of specific herbicides. The dominant weed species in 2001–04, with frequency higher than 10%, were largely the same as in 1987–89, but included three new species in 2001–04. Only a few species occurred less frequently in 2001–04 than in 1987–89 and only in some crops. As common weed species make up an important food reservoir for wildlife, new trends in the management of arable land in Denmark seem to have led to more biodiversity.