The relationship between level of sward height reduction in a rotationally grazed sward and short-term intake rates of dairy cows

The objective of this study was to investigate the relationship between level of sward height reduction (SHR) and short-term intake of herbage by lactating dairy cows offered swards differing in initial bulk density (BD). Three experiments were carried out in which cows were presented with swards representing different levels of SHR (nominally described as ungrazed, low, moderate and high). Experiments 1 and 2 differed with respect to initial sward BD [ungrazed sward 1·7 vs. 2·5 kg dry matter (DM) m^?3 respectively]. Experiment 3 investigated the interaction between BD and SHR. In each experiment, sixteen Holstein/Friesian cows (fitted with excreta collection bags) grazed for a 1-h period in 200-m² plots that had been grazed on the previous day to predetermined sward surface heights (SSH) (= levels of SHR). Herbage intake rates were assessed from changes in live weight, with a correction for insensible weight loss (IWL). Biting rates were recorded from visual observation. High levels of SHR were associated with a significant reduction in SSH, herbage mass and leaf fraction, and a significant increase in sward BD, stem and dead fraction, and DM concentration. Herbage intake, expressed either as DM intake per bite or DM intake per hour, declined as level of SHR increased from low to high. The level of SHR generally had no effect on biting rate. Intake rates varied from 1·9 to 4·4 kg DM h^?1, whereas DM intake bite^?1 ranged from 0·5 to 1·3 g. Pooled regression analysis identified SSH (P < 0·001; r² = 0·94) as the principal determinant of DM intake bite^?1. The regression equation was not significantly improved by the addition of terms for leaf fraction, BD, or herbage mass. In Experiment 3, a significant interaction between level of SHR and sward BD was observed. It is concluded that the principal factor controlling intake (g DM bite^?1 or kg DM h^?1), as swards are progressively grazed down, is SSH, but at a high level of SHR, sward BD also influences intake bite^?1.     

Allowance-intake relations of cattle grazing vegetative tall fescue†

Herbage allowance is one of the important pasture factors in the determination of intake by grazing livestock. Ingestive behaviour of 12 adult Angus cows (Bos taurus) was measured over a range of allowances (0·25 to 0·72 kg dry matter (DM) per 100 kg live weight (LW) for a 1-h period) of vegetative tall fescue (Festuca arundinacea Schreb.). A balanced change-over design was used to estimate direct, residual and permanent effects of herbage allowance on rate of DM intake, rate of biting and herbage DM intake per bite. In Experiment 1, herbage DM intake per meal increased linearly from 0·68 to 1·72 kg (100 kg LW)^?1 as DM allowance increased from 0·25 to 0·72 kg (100 kg LW)^?1 h^?1. Cows grazed at ·30 kg (100 kg LW)^?1 h^?1 and stopped grazing when the sward was reduced to a height about 10 to 12 cm above the soil surface, approximately defined by the tops of pseudostems. In Experiment 2, herbage DM intake rates of 0·29, 0·47 and 0·42 kg (100 kg LW)^?1 h^?1 were recorded as cows grazed allowances of 0·43, 0·70 and 0·90 kg (100 kg LW)^?1 h^?1 for most of the 1-h grazing period. Limiting herbage DM allowances in Experiment 2 were associated with small reductions in rate of biting and herbage DM intake per bite as allowance declined. Sward DM density (>5 cm) was an important variable in the determination of herbage DM intake rates at lower herbage allowances.     

Pattern of herbage intake rate and bite dimensions of rotationally grazed dairy cows as sward height declines

To allow improved prediction of daily herbage intake of dairy cows in rotational grazing systems, intake behaviour was assessed throughout the day in 24‐h paddocks. Herbage intake in 16 lactating Holstein–Friesian cows was assessed using the short‐term (1‐h) weight gain method at four predetermined natural meal times throughout the day (early morning, T1; late morning, T2; mid‐afternoon, T3; and early evening, T4). The study comprised two 4‐day experiments, each with a cross‐over design of four blocks. In both experiments, cows grazed a 24‐h paddock daily, and the effect of the immediately previous grazing experience on intake behaviour was investigated throughout the day, taking account of daily fluctuations in the short‐term physiological condition of the cows. Experiment 1 was carried out to investigate overall grazing behaviour during meals as a sward is progressively depleted during the day, with intake being assessed within the paddock and, hence, on a depleted sward. Experiment 2 similarly investigated the effect of sward depletion and physiological condition throughout the day on intake, but cows were removed to fresh, undefoliated swards during intake measurement periods; thus, intake rate was not influenced by differences in sward condition. Intake behaviour from both experiments was compared to establish the effect on herbage intake of changes in sward state and non‐sward factors. In Experiment 1, sward surface height, available herbage mass, proportion of leaf and green leaf mass declined as the day progressed. Bite mass declined with sward depletion, and mean intake rate was 1·64 kg dry matter (DM) h^–1, which was significantly lower at T3 (P < 0·01) than during other meals. In Experiment 2, plot sward conditions did not change throughout the day, and intake behaviour also remained constant, with a mean intake rate of 2·11 kg DM h^–1. Mean bite depth as a proportion of pregrazing extended tiller height was constant throughout the day (mean 0·32). The results show that, although cows grazed throughout the day on progressively depleted swards, indicative of rotationally grazed paddocks (Experiment 1), bite mass declined linearly and intake behaviour was variable. However, where intake was assessed on high‐quality, undefoliated swards (Experiment 2), intake behaviour was similar regardless of the time of day and the immediately previous experience. There was some indication of an interaction between the effects of the sward and the physiological condition of the animal on herbage intake.     

The performance of summer-calving cows grazing perennial ryegrass swards

The objectives of this experiment were to study the effects of different grazing managements in spring on herbage intake and performance of summer-calving dairy cows and to examine the effects of regrowth in early June on herbage intake and cow performance. Four spring-grazing treatments were applied to predominantly perennial ryegrass swards: Control (C), sward grazed by cows to 6–8 cm sward surface height (SSH); CG16, sward grazed by cows to 3–4 cm SSH in May and allowed to regrow to a target SSH of 16cm in early June; CG8, sward grazed by cows to 3–4 cm SSH in May and allowed to regrow to 8cm in early June; and SG8, sward grazed by sheep to 2–3 cm SSH in May and allowed to regrow to 8 cm in early June, All swards were continuously stocked by summer-calving (May and July) primiparous and multiparous cows from 16 June to 7 September, to a target SSH of 8–10cm. Spring treatments bad marked effects on herbage intakes and milk production. Estimated in July by n alkane analysis, the mean herbage intake ± s.e.d. of cows on each treatment were 1·8, 1·4, 1·4 and 3·0 ± 0·31 kg dry matter (DM) 100 kg live weight (LW)^?1 d^?1 (P < 0·01) for treatments C, CG16, CG8 and SG8 respectively. Measured in August, intakes were 1·8, 20, 2·1 and 2·4 ± O·33kg DM 100kg LW^?1 d^?1 respectively. Severe spring grazing led to increased milk yield and reduced milk fat content from summer-calving cows fed 5·2 kg d^?1 of a proprietary concentrate. Average milk yields for the eleven experimental cows on each treatment were 24·3, 23·4, 26·2 and 29·0 ± 1·20 kgd^?1 (P < 0·01) for C, CG16, CG8 and SG8, and average milk fat contents were 45·4. 42·4, 43·9 and 40·9 ± 1·02gkg^?1 (P<0·05) respectively. The results suggest that severe grazing of swards in early season could improve herbage intake and milk yield of summer-calving cows in mid- and late season. The most favourable spring treatment in this respect was severe grazing by sheep. However, this advantage could be negated in midseason by lax grazing at that time.     

Ingestive behaviour of beef cattle grazing alfalfa (Medicago sativa L.)†

Development of simulation models of grazing beef cattle requires measurement of the components of the ingestive process and the establishment of relationships between these components and the structure of the sward. The ingestive behaviour of eight half-sib Angus steers (live weight (LW), x?= 270 kg) grazing alfalfa (Medicago sativa L.) was studied at three stages of maturity (26, 40 and 47 days of regrowth) and at four allowances of herbage dry matter (DM) (1·0, 1·5, 20 and 2·5 kg per 100 kg LW) at each of two daily grazing sessions. A tethering system of grazing was used in which the experimental unit was a tethered steer and its plot for one grazing session. Grazing sessions commenced at 08.00 and 14.00 h EDT. Intake (DM) increased linearly from 1·98 kg per steer session at a DM allowance of 1 kg (100 kg LW)^?1 to 2 89 kg steer session at an allowance of 2·5 kg (100 kg LW) ^?1 as utilization of herbage declined linearly from 0·69 to 0·43. Herbage DM in take per bite increased from 1 0 g at 1 kg (100 kg LW) ^?1 allowance to 1·5 g at 2·5 kg (100 kg LW) ^?1 allowance. Rates of biting were not affected by herbage allowance and averaged 21 bites min^?1. Dry matter intake increased from 1·77 to 3 41 kg per steer session as the alfalfa matured and herbage mass changed from 1500 to 4656 kg ha^?1. Mean rates of biting were 24 bites min^?1 for steers grazing the youngest alfalfa and 16 bites min^?1 for steers on the oldest forage. Herbage DM intakes per bite were 1·1 g and 1·7 g at the same stages. Rates of DM intake approached 2 kg h^?1 and maximum daily DM intake was estimated at 2 75 kg (100 kg LW) ^?1. Intake of alfalfa was limited by allowance and mass of herbage above a canopy horizon of 20 cm and, to a lesser extent, by the length of fast.     

The effect of initial spring grazing date and subsequent stocking rate on the grazing management, grass dry matter intake and milk production of dairy cows in summer

The objective of this study was to investigate the effects of an early (February; F) or delayed (April; A) primary spring grazing date and two stocking rates, high (H) and medium (M), on the grazing management, dry matter (DM) intake of grass herbage and milk production of spring‐calving dairy cows grazing a perennial ryegrass sward in the subsequent summer. Sixty‐four Holstein‐Friesian dairy cows (mean of 58 d in milk) were assigned to one of four grazing treatments (n = 16) which were imposed from 12 April to 3 July 2004. Cows on the early spring‐grazing treatment were grazed at 5·5 cows ha^?1 (treatment FH) and 4·5 cows ha^?1 (treatment FM) while cows on the late‐grazing treatment were grazed at 6·4 cows ha^?1 (treatment AH) and 5·5 cows ha^?1 (treatment AM). The organic matter digestibility and crude protein concentration of the grass herbage were higher on the early‐grazing treatment than on the late‐grazing treatment. The cows on the FM treatment had significantly (P < 0·001) higher milk (24·5 kg), solids‐corrected milk (22·5 kg), fat (P < 0·01, 918 g) and protein (831 g) yields than the other three treatments. Cows on the FM treatment had a higher (P < 0·001) DM intake of grass herbage by 2·3 kg DM per cow per day than cows on the AH treatment, which had a DM intake significantly lower than all other treatments (15·2 kg DM per cow per day). The results of the present study showed that grazing in early spring has a positive effect on herbage quality in subsequent grazing rotations. The study also concluded that early spring‐grazed swards stocked at a medium stocking rate (4·5 cows ha^?1; FM) resulted in the highest DM intake of grass herbage and milk production.     

Effect of continuous stocking by sheep at four sward heights on herbage mass, herbage quality and tissue turnover on grass/clover and nitrogen-fertilized grass swards

The effects of continuous stocking by sheep at sward surface heights (SSH) of 3, 5, 7 and 9 cm in grass/clover (GC) and nitrogen-fertilized grass (GN) swards were examined in relation to herbage mass and quality, clover content, tiller density and rates of herbage production and senescence in two periods in each of three grazing seasons (1987-89). The GN swards received a total of 300 kg N ha^?1 each year in six equal dressings from March; GC swards received a single dressing of 50 kg N ha^?1 in March each year. Herbage mass measured from ground level increased linearly with SSH with overall mean herbage masses of 0·89, 1·38, 1·78 and 2·12 t OM ha^?1 (s.e.m.0·024, P < 0·001) at SSH of 3, 5, 7 and 9 cm respectively. GN and GC swards had mean herbage masses of 1·58 and 1·51 t OM ha^?1 (s.e.m. 0·051, NS) respectively. Mean N content of herbage on GN swards was greater than that on GC swards and declined with increasing SSH. Crude, fibre (CF) content of herbage was similar for both sward types and increased with increasing SSH. Clover content of GC swards remained low throughout the experiment, ranging from 0·002 to 0·074 of herbage mass. However, from tissue turnover rates it was estimated that its contribution to herbage production was in the range of 0·049–0·219 of net herbage growth. Total growth increased with increasing SSH in both sward types, with maximum growth rates in GN swards of 143 and 130 kg DM ha^?1 d^?1 and in GC swards of 88·2 and 85·4 kg DM ha^?1 d^?1 in Periods 1 (up to early July) and 2 (after July) respectively. Senescence rates ranged between 13·3 and 50·1 kg DM ha^?1 d^?1 and tended to be higher in Period 2 than in Period 1. Net production increased with increasing SSH in Period 1, while in Period 2 net production declined at SSH above 6·5 cm. The increased net herbage production in taller swards was not associated with greater utilized metabolizable energy production at sward heights above 5 cm.     

The effects of poloxalene on ingestion by cattle grazing Lucerne†

The mechanisms that terminate meals of cattle grazing lucerne (Medicago sativa L.) are not well defined. Sub-acute bloat may lead to cessation of grazing and, consequently, surface active substances used in the treatment and prevention of bloat, such as poloxalene, may extend grazing meals and increase herbage intake. Twelve mature Angus cows (Bos taurus) were offered 0, 12·5 and 25·0 g poloxalene in 0·5 kg of crushed maize (Zea mays L.) kernels each day, immediately before two consecutive 1-h measured parts of a grazing meal on 21- to 24-day-old lucerne swards with a herbage dry matter (DM) mass (> 5 cm) of 2·03 t ha^?1 and herbage DM mass allowance of 3·55 kg hd^?1h^?1. Total herbage DM intake was 2·52 kg hd^?1 during the first hour and 1·54 kg hd^?1 during the second hour of the 2-h grazing meal. Differences in herbage intake were attributable to a cessation of grazing. Mean rates of biting were 26·3 and 14·8 bites min^?1 and mean DM intakes per bite were 1·82 and 4·38 g during the first and second part of meals, respectively. Poloxalene treatments caused a small linear decline in grazing time during the first part of meals and a larger increase in grazing time during the second part of meals. Lower rates of DM intake caused by poloxalene were offset by increases in grazing time. It was concluded that poloxalene moderated ingestive behaviour within grazing meals of immature lucerne and this response may have been at least partly due to the relief of sub-acute bloat.     

The effects of sward characteristics and supplement type on the herbage intake and milk production of summer-calving cows

High (H; 27 350 m^?1:) or low (L. 13 300 m-¹) tiller density perennial ryegrass swards were created in the mid- and late grazing season by imposing different sward heights in the spring. Summer-calving cows then grazed these swards from 6 June to 2 September 1992 and were offered 5 kg fresh weight hd^?1 d^?1 of either a barley (S) or a molassed sugar beet pulp (F) based supplement. The factorial combination of sward and supplement types resulted in four experimental swards being grazed by thirteen Holstein/Friesian cows each. Supplement F contained more crude fibre (110 vs. 58 g kg^?1) and less metabolizable energy [12–5 vs. 13–2 MJ kg^?1 dry matter (DM)] than supplement S. Herbage on the H sward contained more metabolizable energy (11–9 vs. 104 MJ kg^?1 DM) and crude protein (232 vs. 205 g kg^?1 DM), had fewer rejected areas f 16–5 vs. 26–9%) and a higher live-dead tiller ratio (4–6 vs. 2–1) than that on the L sward. Sward, but not supplement type, significantly affected the intake of grazed herbage (P<0–001). On average, the herbage intakes of cows grazing II swards were higher than for L swards (14–5 vs. 11 6kg DM d^?1) and those of cows on the S and F supplements were 12–6 and 13–5 kg DM d^?1 respectively. Averaged over the grazing period, sward and supplement had no significant effects on milk yield, milk composition or yield of constituents. When expressed on an average weekly basis, cows grazing an L sward and offered the F supplement on occasions had significantly lower milk yields and higher milk fat contents (P<005) than those grazing an H sward and offered the S supplement. There were no significant effects on cow live weight or condition score change. The results suggested that grazing swards with a high density of live tillers increased herbage intakes and on occasions milk yield, relative to low density swards. However, small increases in energy intake from sward and supplement effects were used primarily to ameliorate liveweight loss.     

Does post‐grazing sward height influence sward characteristics,seasonal herbage dry‐matter production and herbage quality?

Extending the grazing season through the production and utilization of high‐quality forage is a key objective in grassland‐based dairy production systems. Grazing swards to a low post‐grazing sward height (PGSH) is a strategy for improving grass utilization. A grazing experiment conducted in Ireland investigated immediate and subsequent effects of PGSH on sward production, utilization and structural characteristics. Swards were grazed to 2·7 cm (severe; S1) or 3·5 cm (moderate; M1) from 10 February to 18 April 2010 (Period 1; P1). From 19 April, each P1 paddock was halved and grazed to either 3·8 cm (S2) or 4·8 cm (M2), until 30 October (Period 2; P2). The first grazing rotation was +7 d on S1 swards compared with M1 swards (45 d), due to greater herbage utilization (+0·22). Herbage production during P1 was not affected by PGSH but a severe PGSH during this period reduced subsequent herbage production: 13·9 (S1) vs. 15·5 t dry matter (DM) ha^?1 (M1) by the end of the study. Leaf proportion was increased (+0·10) on S2 swards compared with M2 swards, but M2 swards produced 1·2 t DM ha^?1 more herbage during P2. Despite the relatively lower level of sward utilization obtained from moderate grazing in P1 (3·5 cm) and P2 (4·5–5·0 cm), such levels of PGSH increased DM production while maintaining sward quality, compared with severe grazing (2·7 cm in P1 and 3·5–4·0 cm in P2).     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 3. Sward characteristics and animal output, years 4–8

Results for years 4–8 of a long-term grazing experiment on swards of a diploid perennial ryegrass (Lolium perenne), var. Contender (D swards), a tetraploid ryegrass, var. Condesa (T swards) and Condesa with S184 white clover (Trifolium repens) (TC swards), direct sown in May 1987, are presented. The swards were continuously stocked with sheep from 1988 to 1990, as previously reported, and for a further 5 years, 1991–95, at a target sward surface height (SSH) of 4–6 cm. Control of sward height was successfully achieved by variable stocking, except in 1993 when paddocks were set stocked and the resulting mean SSH was 9·3 cm. Grass swards received on average 160 kg N ha^?1 year^?1; grass/clover swards were mainly not fertilized with N with the exception that they were given 30 kg N ha^?1 as a remedial mid-summer application during a period of low herbage mass on offer in 1994 and 1995. Mean white clover content of the swards fell from 18·2% of herbage dry-matter (DM) in 1992 to 8·5% in 1993, whereas stolon lengths fell from 120 to 58 m m^?2. A return to lower sward heights in 1994–95 resulted in an increase in white clover content to 12·8% by the final sampling in August 1995. Perennial ryegrass content of the grass swards remained high throughout (mean 96·7% in 1995). Perennial ryegrass tiller densities recorded in August 1991, 1993 and 1994 showed consistently significant (P < 0·001) sward differences (3-year mean 16 600, 13 700 and 10 100 perennial ryegrass tillers m^?2 for the D, T and TC swards). In 1994, the year after lax grazing, a low perennial ryegrass tiller density (9100 m^?2) and low white clover content (mean 4·3%) in the TC swards resulted in a much lower herbage bulk density than in the grass swards (April–July means 72, 94 and 44 kg OM ha^?1 cm^?1 for the D, T and TC swards). There was a consistent 40 g d^?1 increase in lamb liveweight gain on the TC swards over the T swards, except in 1994. In that year there was a reduction in lamb liveweight gain of 33 g d^?1 on the TC swards and a significant increase in ewe liveweight loss (117 g d^?1) associated with low herbage bulk density despite optimal sward height. Lamb output (kg liveweight ha^?1) on TC swards reflected white clover content, falling from a similar output to that produced from grass given 160 kg N ha^?1, at 18% white clover DM content, down to 60% of grass + N swards with around 5% clover. A 6% greater output from the T than the D swards was achieved mainly through higher stocking rate. The experiment demonstrated a rapid, loss in white clover under lax grazing, and showed that the relationship between performance and sward height is also dependent on herbage density. High lamb output from a grass/clover sward was only achieved when the clover content was maintained at 15–20% of the herbage DM.     

Effect of Lolium perenne sward density on productivity under simulated and actual cattle grazing

The objective of this study was to quantify the effect of perennial ryegrass (Lolium perenne L.; PRG) sward density on seasonal and total DM yield under simulated grazing and animal grazing by cattle, and to assess the effectiveness of visually estimated ground scores (GSs) for predicting sward PRG density. The study incorporated five different seeding rates of PRG, each replicated three times, to simulate swards ranging in PRG density typical of different ages and conditions. There was no significant difference between defoliation managements for total DM yield, but sward PRG density had a significant effect on both the seasonal and total DM herbage yield under both systems. Under simulated grazing, total DM yield ranged from 10·7 to 12·0 t DM ha^?1 with increasing sward PRG density at a GS range of 1·70–4·28 (mean of 2 years’ data, P < 0·01). Under animal grazing, the yield range was from 10·3 to 12·2 t DM ha^?1 for a GS range of 1·50–3·39 (mean of 2 years’ data, P < 0·01). The largest differences in DM yield occurred during the spring period. The relationship between sward DM yield and GS was significant (P < 0·001) for both simulated and animal‐grazed swards. Each unit increase in midseason GS (June) related to an average yield increase of 350 kg DM ha^?1 under simulated grazing and a 721 kg DM ha^?1 increase under cattle grazing. Every unit increase in the GS at the end of the grazing season (December) was associated with a 460 or 1194 kg DM ha^?1 increase under simulated and animal grazing, respectively. These results show that visual estimates of density were an effective tool in describing PRG density and that this could be related to DM yield potential. Further investigations may provide a threshold value below which the renewal of swards could be advised based on a visual GS of PRG.     

Ingestive behaviour of beef cattle offered different forms of lucerne (Medicago sativa L.)†

A change-over design was used to establish the ingestive behaviour of twelve adult Angus cows (Bos taurus), live weight (LW) of 475 ± 18 kg, while grazing luceme (Medicago sativa L.) (T₁), eating from a swath of freshly cut lucerne (T₂), or eating wilted lucerne from a swath (T₃). In the first experiment, the herbage dry matter (DM) allowance was 4-7 kg h^?1 and in T₁ the available herbage DM mass (>5 cm) was 2892 kg ha^?1. Herbage DM intake was 2.5, 1.6 and 2.0 kg h^?1 (0.53,0.35 and 0.41 kg (100 kg LW)^?1h^?1) for T₁, T₂ and T₃, respectively. Cattle grazed at 29 bites min^?1 and ate the swathed lucerne at 6 bites min^?1. Herbage DM intake per bite was 1-2, 4-9 and 6-7 (2-5,104 and 14-4 mg(kg LW)^?1)for T₁, T₂ and T₃, respectively. The DM content of the luceme was 227, 263 and 309 g kg^?1 for the same sequence. In a second experiment with luceme of available herbage DM mass of 5321 kg ha^?1 and an allowance of 7- 3 kg h^?1 cows grazed lucerne at 2-9 kg h^?1, ate freshly cut material at 21 kg h^?1 and wilted herbage at 25 kg h^?1 (0.61, 0.45 and 0.52 kg (100 kg LW)^?1h^?1), respectively. Leaf accounted for 80%, 68% and 54% of intake for T₁, T₂ and T₃, respectively. Swathing lucerne reduced the diet selectivity by forcing cattle to take large bites that required many jaw movements to form a bolus that could be swallowed. It appeared that the mechanics of bolus formation determined ingestive behaviour of cattle eating swathed herbage.     

Production performance and meat quality of grazing lambs finished on red clover, lucerne or perennial ryegrass swards

The combined benefits of a high crude protein concentration, and possible protein protection and growth‐promoting properties, make forage legumes potentially attractive as a natural means of increasing liveweight gain and time to slaughter of lambs in lamb finishing systems. An experiment was conducted to compare the production performance and meat quality of grazing lambs finished on red clover (Trifolium pratense), lucerne (Medicago sativa) or perennial ryegrass (Lolium perenne) swards. Replicate (n = 2) swards of red clover, lucerne and perennial ryegrass were rotationally grazed by ten ram lambs and ten ewe lambs from weaning until selection for slaughter at UK fat class 3L. Lambs grazing the red clover sward had a significantly higher liveweight gain and required significantly fewer days to slaughter than lambs grazing the lucerne sward (305 g d^?1 vs. 243 g d^?1; 38 d vs. 50 d), which in turn had a higher liveweight gain and required fewer days to slaughter than lambs grazing the perennial ryegrass sward (184 g d^?1; 66 d). Lambs grazing the red clover and lucerne swards had significantly higher herbage intakes than those grazing the perennial ryegrass sward (2·06, 1·72 and 1·16 kg DM d^?1 respectively), but in vivo digestibility of herbage was similar. Lambs grazing the red clover and lucerne swards also had significantly higher serum urea concentrations than those grazing ryegrass (12·5, 11·1 and 6·2 mmol L^?1 respectively). Killing‐out percentage was significantly higher for lambs grazing the red clover sward than for lambs grazing the perennial ryegrass sward (48% vs. 46%). There were no significant effects of finishing system on meat flavour, but meat from lambs finished on the lucerne sward was oxidatively less stable than that from lambs finished on the perennial ryegrass sward. Grazing the forage legume swards significantly increased the proportion of linoleic and linolenic acid in muscle tissue, and therefore the proportion of unsaturated to saturated fatty acids (0·19, 0·16 and 0·12 for the red clover, lucerne and perennial ryegrass swards respectively). However, the n?6/n?3 ratio was significantly lower for the muscle of lambs grazing the perennial ryegrass sward compared with those grazing the forage legume swards (1·13, 1·08 and 0·98 for the red clover, lucerne and perennial ryegrass swards respectively). The results indicate that by grazing lambs on forage legume swards it is possible to increase individual lamb performance without compromising meat quality.     

Effects of supplementation on the ingestive behaviour of grazing steers*

Herbage intake is usually depressed when beef cattle grazing abundant pastures are supplemented with energy-rich feedstuffs but relatively little is known about the effects of supplementation on the components of ingestive behaviour. An experiment was conducted to establish the effect of ground corn (Zea mays L.) on the ingestive behaviour of yearling Angus and Angus × Hereford steers (Bos taurus) (mean live weight (LW) of 323 kg) grazing autumn stockpiled Boone cocksfoot (Dactylis glomerata L.) using a tethered grazing system in which the experimental unit was the tethered steer and its grazing area (45 m²) for one grazing session. Herbage dry matter (DM) mass was 1662 kg ha^?1 and herbage DM allowance was 7–5 kg steer^?1 for each grazing session. Herbage DM intake was measured as the difference between herbage DM mass offered and refused. Grazing took place during two daily sessions each of about 2 h duration commencing at 08.00 and 14.00 h for 9 days. Ground corn (0, 1·5, 3·0 and 4·5 kg steer^?1) was fed each day at 12.00 h and had no significant (P < 0·05) effect on rate of DM intake, rate of biting or DM intake per bite. Mean DM intake was 6 2 kg steer^?1 d^?1 (87 mg (kg LW)^?1 min^?1). Steers averaged 4832 bites per grazing session, with a mean DM intake per bite of 644 mg (2·0 mg (kg LW)^?1) and a mean rate of 44 bites min^?1. Data obtained at the beginning and end of each grazing period on ingestive behaviour of one group of four steers (mean LW of 306 kg) fitted with oesophageal fistulae supported data for the two groups of normal steers and showed no response to supplementation. Mean values for rate of DM intake, DM intake per bite and rate of biting established for the fistulated cattle were 73 mg (kg LW)^?1 min^?1, 521 mg bite^?1 (1·7 mg (kg LW)^?1) and 39 bites min^?1, respectively.     

Effect of sward surface height on the performance of ewes and Iambs continuously grazed on grass/clover and nitrogen-fertilized grass swards

In three successive years, sward height was maintained at 3, 5, 7 or 9 cm on grass swards receiving a total of 300 kg N ha^?1 in six equal monthly dressings from April, and on grass/clover swards receiving 50 kg N ha^?1 as a single dressing in early spring. From turnout in April until weaning in July, 64 ewes and their lambs (mean litter size 1·5) were continuously grazed at the four sward surface heights on the two sward types. White clover content of grass/clover swards remained low throughout the experiment ranging from 0·2 to 7·4% of the herbage mass. During the first two years, lamb gains averaged over sward types were 204, 260, 285 and 308 g d^?1 up to weaning, while in the third year gains were 238, 296, 296 and 260 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Ewes lost live weight on 3 cm swards but apart from this sward height had little effect on performance. During the autumn, weaned lambs gained — 27, 87, 147 and 167 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Sward type had only a small effect on the performance of lambs up to weaning but in the autumn, mean gains of weaned lambs were lower on grass/N swards (73 g d^?1) than on grass/clover swards (115 g d^?1). Relative to 3 cm swards, carrying capacities of 5, 7 and 9 cm swards were 0·76, 0·57 and 0·52 respectively from turnout to weaning and 0·66, 0·52 and 0·44 respectively during autumn. Grass/clover swards carried 0·67 of the ewes carried by grass/N swards from turnout to weaning and 0·51 of the live weight carried by grass/N swards during autumn. The reaction of the two sward types to sward height did not appear to differ but in the third year there was evidence of a reduction in white clover content when swards were grazed at 9 cm. The data suggest that lamb growth rates will increase as sward height increases up to 9 cm and the evidence for this was stronger with weaned lambs in autumn than with suckling lambs in spring.     

Effect of sward surface height on intake and grazing behaviour by lactating Holstein Friesian cows

The effect of sward surface height (SSH) on grazing behaviour and intake by lactating Holstein Friesian cows on continuously stocked grass pastures maintained at mean heights of 5, 7 and 9 cm was studied during the growing season. Intake rate was estimated over periods of 1 h by weighing animals before and after grazing, with a correction made for insensible weight loss. Grazing behaviour during that hour and over 24 h was recorded automatically using sensors to measure jaw movements. Although maintained at the overall mean SSH, swards had a patchy appearance with short, frequently grazed areas interspersed with taller, infrequently grazed areas, which is typical of pastures continuously stocked with cattle. Daily organic matter (OM) intake, calculated as the product of daily grazing time and intake rate, was greater at a SSH of 7 cm than at 5 or 9 cm (14·1 vs. 10·5 and 12·1 kg respectively). On the 5-cm sward, OM intake per grazing jaw movement (GJM) was reduced compared with that on the 7-cm sward (0·182 vs. 0·264 g respectively), and because cows were unable either to increase significantly GJM rate (95·8 vs. 90·1 GJM min^?1) or the proportion of GJM that were bites (0·80 vs. 0·81) OM intake rate was reduced (16·9 vs. 23·5 g min^?1). Cows were unable to increase their grazing time significantly (628 vs. 604 min d^?1) to compensate for the reduction in intake rate, and as a result daily intakes were lower. Cows grazing the 9-cm sward also incurred a reduction in OM intake GJM^?1 compared with those on the 7-cm sward (0·237 vs. 0·264 g respectively) and therefore there was a reduction in OM intake rate (21·6 vs. 23·5 g min^?1). These animals did not compensate by increasing the time spent grazing (581 min d^?1), probably owing to an increased ruminating requirement per kg of herbage ingested compared with those on 7 cm SSH (2264 vs. 1780 ruminating jaw movements respectively). The results show that SSH can significantly influence intake rate, but, while the cow's only effective strategy to compensate for any reduction in intake rate is to increase grazing time, this may be limited by the requirement for ruminating and non-grazing, non-ruminating activities, which is influenced by qualitative and quantitative aspects of the herbage ingested, whereas the cows' only effective strategy to compensate for any reduction in intake rate is to increase grazing time.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

Herbage intake and N excretion by sheep grazing monocultures or a mixture of grass and white clover

In 1988 and 1989, swards of grass (G0), while clover (C0) and grass/white clover (GC0) receiving no N fertilizer, and a grass sward supplied with 420 kg N ha^?1 (G420), were grazed by non-lactating sheep to maintain a sward surface height of 6 cm. Herbage organic matter (OM) intakes averaged between 1200 and 1700 g OM ewe^?1 d^?1. For treatments G0, C0, GC0 and G420 respectively, the ewes' live weight gain was 102, 112, 100 and 110 g d^?1 and changes in body condition scores were +0·28, +0·52, +0·36 and +0·44 units season^?1. However, the effect of treatment was not significant for either variable. There were similar levels of output of faecal N ewe^?1 but significantly more urinary N ewe^?1 was excreted on treatments C0 and G420, where the concentrations of N in herbage laminae were also higher. For example, in 1989, total daily N excreted was 39·7, 64·4, 44·0 and 63·3 g N ewe^?1 for G0, C0, GC0 and G420 respectively. Taking into account the mean daily stocking rates, which were 19·4, 26·6, 27·2 and 36·5 ewe ha^?1, the total faeces and urine returns over the season were 161, 358, 249 and 484 kg N ha^?1 for each treatment respectively. The herbage OM intakes ewes^?1 d^?1 measured in September and October were similar for C0 and G420, and so the intake of herbage OM ha^?1 d^?1 was related to stocking rate, i. e. the estimated herbage intake ha^?1 over the growing season for the white clover monoculture was 73% of that for N-fertilized grass. Excretal nitrogen returns to the pasture from grazed mono-cultures of clover were high, and similar to those from a grass sward receiving 420 kg fertilizer N ha^?1. Consequently potential losses of N to the environment are high under these management systems.     

Influence of sward condition on leaf tissue turnover in tall fescue and tall wheatgrass swards under continuous grazing

Two experiments were carried out on a tall fescue sward in two periods of spring 1994 and on a tall wheatgrass sward in autumn 2001 and spring 2003 to analyse the effect of sward surface height on herbage mass, leaf area index and leaf tissue flows under continuous grazing. The experiment on tall fescue was conducted without the application of fertilizer and the experiment with tall wheatgrass received 20 kg P ha^?1 and a total of 100 kg N ha^?1 in two equal dressings applied in March (autumn) and end of July (mid‐winter). Growth and senescence rates per unit area increased with increasing sward surface height of swards of both species. Maximum estimated lamina growth rates were 28 and 23 kg DM ha^?1 d^?1 for the tall fescue in early and late spring, respectively, and 25 and 36 kg DM ha^?1 d^?1 for tall wheatgrass in autumn and spring respectively. In the tall fescue sward, predicted average proportions of the current growth that were lost to senescence in early and late spring were around 0·40 for the sward surface heights of 30–80 mm, and increased to around 0·60 for sward surface heights over 130 mm. In the tall wheatgrass sward the corresponding values during spring increased from around 0·40 to 0·70 for sward surface heights between 80 and 130 mm. During autumn, senescence losses exceeded growth at sward surface heights above 90 mm. These results show the low efficiency of extensively managed grazing systems when compared with the high‐input systems based on perennial ryegrass.