Species interactions in a grassland mixture under low nitrogen fertilization and two cutting frequencies: 1. dry‐matter yield and dynamics of species composition

Four‐species mixtures and pure stands of perennial ryegrass, tall fescue, white clover and red clover were grown in three‐cut and five‐cut systems at Ås, southern Norway, at a low fertilization rate (100 kg N ha^?1 year^?1). Over a three‐year experiment, we found strong positive effects of species diversity on annual dry‐matter yield and yield stability under both cutting frequencies. The overyielding in mixtures relative to pure stands was highest in the five‐cut system and in the second year. Among the possible pairwise species interaction effects contributing to the diversity effect, the grass–grass interaction was the strongest, being significant in both cutting systems and in all years. The grass–legume interactions were sometimes significant, but no significant legume–legume interaction could be detected. Competitive relationships between species varied from year to year and also between cutting systems. Estimations based on species identity effects and pair‐specific interactions suggested that the optimal proportions of red clover, white clover, perennial ryegrass and tall fescue in seed mixtures would have been around 0·1, 0·2, 0·4 and 0·3 in the three‐cut system, and 0·1, 0·3, 0·3 and 0·3 in the five‐cut system.     

Production and nutritional value of sorghum and black oat forages under nitrogen fertilization

The effect of nitrogen (N) fertilization on the dry‐matter (DM) yield and nutritional value of sorghum (Sorghum sp., cv. Jumbo) and black oat (Avena strigosa cv., IPR 61) was investigated in the context of forage and livestock production in southern Brazil. Sorghum was cultivated with 0, 37·5, 75, 150, 225, 300 and 375 kg N ha^?1 during the summer crop seasons of 2010/11 and 2011/12. Black oat received 0, 40, 80, 120, 160, 200 and 240 kg N ha^?1 in the winter of 2011. According to the adjusted polynomial regression, sorghum DM yield increased in response to N up to 288 (12·9 t ha^?1) and 264 kg ha^?1 (5·6 t ha^?1) in 2010/11 and 2011/12 respectively. Crude protein (CP) content of sorghum was highest at 349 and 328 kg N ha^?1, but in vitro dry‐matter digestibility (IVDMD) was highest at 212–207 kg N ha^?1 in 2010/11 and 2011/12 respectively. Sorghum neutral detergent fibre (NDF) and acid detergent fibre (ADF) were not affected by N fertilization. In black oat, the maximum DM yield (6·0 t ha^?1) was obtained with 187 kg N ha^?1; the IVDMD, NDF and ADF were not affected by N fertilization, but the CP content increased up to 220 kg N ha^?1. It is concluded that these forage species can improve the year‐to‐year amount and quality of forage produced but high rates of N fertilizer are required to achieve high yields. Fertilizer N rates of 210–280 kg N ha^?1 in sorghum and 180 kg N ha^?1 in black oat in the crop rotation provide the greatest responses in DM yield consistent with good nutritional quality for livestock production.     

Herbage production and persistence of two tropical perennial grasses and forage sorghum under different nitrogen fertilization and defoliation regimes in a summer‐dominant rainfall environment,Australia

An experiment was conducted in inland northern New South Wales (NSW) to assess the response of tropical perennial grasses Chloris gayana (Rhodes grass) cv. Katambora and Digitaria eriantha (digit grass) cv. Premier and annual forage sorghum (Sorghum bicolor ssp. bicolor × S. bicolor ssp. drummondii hybrid) cv. Sweet Jumbo fertilized with five rates of nitrogen (N; 0, 50, 100, 150 and 300 kg N ha^?1) and defoliated every 2 or 6 weeks over two growing seasons. Tropical perennial grasses were highly responsive to N fertilizer, while there was no significant response by forage sorghum. Herbage production of Rhodes grass increased linearly whereas digit grass had a high response at 50–100 kg N ha^?1. Nitrogen‐use efficiency was highest during the growing season when rainfall was higher. During this season, digit grass had the highest N efficiency (148 kg DM kg^?1 N applied) at 50 kg N ha^?1, and Rhodes grass (66 kg DM kg^?1 N applied) at 100 kg N ha^?1. Plant frequency of both perennial species increased and then stabilized at high levels (>84%, cell size 0·1 by 0·1 m) during the two growing seasons. Plant frequency of Rhodes grass declined over the winter period, but recovered within 6 weeks of commencement of the growing season. Soil nitrate levels indicated that unused nitrate moved down the soil profile during wet winters. Implications of leaching below the rooting zone are discussed.     

Bahiagrass (Paspalum notatum Flugge) management combining nitrogen fertilizer rate and defoliation frequency to enhance forage production efficiency

Bahiagrass (Paspalum notatum Flugge) pastures are widespread in warm climates worldwide and respond to nitrogen (N) fertilizer. Nitrogen fertilization has recently decreased because of increased cost and concerns regarding excessive N in the environment. Responses of bahiagrass to treatments representing three alternative levels of pasture management were assessed. Treatments, each including 56 kg N ha^?1 applied for each growth period, were as follows: (i) six harvests with a total of 336 kg N ha^?1 annually (referred to as intensive management), (ii) three harvests with 168 kg N ha^?1 annually (intermediate management) and (iii) two harvests with 112 kg N ha^?1 annually (extensive management). The intensive management produced the most forage with the highest nutritive value. Intermediate management, with only half the amount of N fertilizer, produced at least 80% of the forage yield each year as the intensive management treatment (4‐year average of 8236 vs. 9122 kg ha^?1 for the intermediate and intensive management treatments respectively) with forage of acceptable nutritive value for some classes of livestock. Limited forage production from the last harvest each year restricts autumn management opportunities, even though crude protein concentration was usually sufficient for some classes of livestock. Extended growth periods, as those that occur with the less‐intensive management treatments, provide opportunities to accumulate forage for late‐season grazing. Matching livestock enterprises to the forage produced, particularly in terms of nutritive value, can contribute to favourable livestock production responses from a range of bahiagrass pasture management approaches.     

High perennial ryegrass seeding rates reduce plant size and survival during the first year after sowing: does this have implications for pasture sward persistence?

Failure of perennial ryegrass swards to persist is a key issue on dairy farms in many areas of the world. This study describes an experiment conducted to test the hypothesis that high ryegrass seeding rates (>18 kg seed ha^?1) reduce plant size and physical survival during the first year after sowing, with negative implications for population persistence. Four cultivars representing four functional types of perennial ryegrass were sown at five seeding rates (equivalent to 6, 12, 18, 24 and 30 kg seed ha^?1) with white clover in three dairying regions of New Zealand. Plant establishment rates, size and survival were measured for the first 13 months after sowing. Herbage accumulation, botanical composition and perennial ryegrass tiller density were also monitored. Increasing seeding rate reduced plant survival in the seven weeks after sowing, reflecting likely differences in germination and very early mortality of small seedlings. Thereafter, plant survival was relatively high and consistent across seeding rates at two sites but was consistently greater in the 6 kg ha^?1 treatment compared with the 24 and 30 kg ha^?1 treatments at one site. Higher seeding rates also increased ryegrass tiller density and the contribution of ryegrass towards total herbage biomass but reduced the contribution of white clover. Very few cultivar × seeding rate interactions were detected for any of the measured variables. Many of the seeding rate effects dissipated by the end of the first year after sowing, indicating that high seeding rates did not predispose swards to poor persistence in the longer term, irrespective of ryegrass functional type.     

Herbage and nitrogen yields,fixation and transfer by white clover to companion grasses in grazed swards under different rates of nitrogen fertilization

In grass–legume swards, biologically fixed nitrogen (N) from the legume can support the N requirements of the grass, but legume N fixation is suppressed by additional fertilizer N application. This study sought to identify a fertilizer N application rate that maximizes herbage and N yields, N fixation and apparent N transfer from white clover to companion grasses under intensive grazing at a site with high soil‐N status. During a 3‐year period (2011–2013), swards of perennial ryegrass and of perennial ryegrass–white clover, receiving up to 240 kg N ha^?1 year^?1, were compared using isotope dilution and N‐difference methods. The presence of white clover increased herbage and N yields by 12–44% and 26–72%, respectively. Applications of N fertilizer reduced sward white clover content, but the effect was less at below 120 kg N ha^?1. The proportion of N derived from the atmospheric N fixation was 25–70%. Nitrogen fixation ranged from 25 to 142 kg N ha^?1 measured using the isotope dilution method in 2012 and from 52 to 291 kg N ha^?1 using the N‐difference method across all years. Fertilizer N application reduced the percentage and yield of fixed N. Transfer of N from white clover to grass was not confirmed, but there was an increased N content in grass and soil‐N levels. Under intensive grazing, the maximum applied N rate that optimized herbage and N yields with minimal effect on white clover content and fixation rates was 60–120 kg N ha^?1.     

Phosphorus efficiency of naturalized Chilean white clover in a grazed field trial

Nine naturalized white clover populations and two cultivars (Huia and Will) were evaluated at two soil phosphorus ( P ) levels (6 and 20 P mg kg^?1) to characterize them for DM production and P efficiency, and to facilitate the selection of suitable genotypes to produce cultivars for P‐deficient marginal soils. The study was carried out in Chile during 2007–2010 under field grazing conditions using a randomized complete factorial block design with three replicates. The clover was grown with perennial ryegrass. Botanical composition and herbage DM production were recorded, phosphorus concentrations in soil and in clover and ryegrass herbage were determined, and P absorption (PAE) and P utilization (PUE) efficiency were calculated. Population 8‐1‐X yielded more DM under the low than under the high soil‐ P level and was the highest yielding clover in the low soil‐ P treatment. It also had the highest PAE (0·6 kg P ha^?1 per mg kg^?1 Olsen P) under low P availability. Dry‐matter yield of 9‐2‐X was statistically similar to 8‐1‐X and Huia at low P level; however, because of its lower P absorption, it had the highest PUE (455 kg DM ha^?1 per kg ha^?1 absorbed P). Furthermore, the highest yield of ryegrass occurred when grown with 8‐1‐X, and this combination gave the maximum total yield. These two populations have potential for inclusion in breeding programmes.     

Effect of dairy slurry application rate and forage type on production,soil nutrient status and nitrogen‐use efficiency

Seven forage types (diploid and tetraploid perennial ryegrass, Italian ryegrass and hybrid ryegrass, a low‐input mixture of perennial ryegrass, cocksfoot, timothy and meadow fescue, a mixture of perennial ryegrass and white clover, and monoculture of red clover) were sown in late July 2004. Each received one of four rates of dairy cattle slurry in three annual applications by trailing shoe, which supplied average nitrogen (N) inputs of 0·0, 114·9, 204·8 and 301·2 kg N ha^?1 annum^?1. Treatments were cut either three or four times annually over four years. Average dry‐matter yield (DM) response to slurry N was 15·6 kg DM kg^?1 N. Lowest recovery of slurry N was in the second application each year (after first cut). The data suggest that slurry applied to Italian ryegrass, and also to swards containing legumes on soils with high phosphorus content, will produce a lower DM response to slurry N and result in a lower slurry N recovery than on swards of perennial ryegrass or cocksfoot‐dominant low‐input mixtures. Apparent recovery of slurry N was low at the second cut, especially when first‐cut yields had been high. To maximize slurry N recovery, application to regrowths with potentially slow rates of growth or high legume content should be avoided.     

Effects of 5 years of digestate application on biomass production and quality of cocksfoot (Dactylis glomerata L.)

Anaerobic digestion of biomass produces biogas for combustion and also provides a residual digestate. Although sometimes regarded as a waste product, the nutrient‐rich chemical composition of digestate makes it a potential organic fertilizer for agriculture. The goal of this study was to evaluate the effectiveness of digestate as a fertilizer on the biomass yield and chemical composition of cocksfoot (Dactylis glomerata L.). In a 5‐year small‐plot field experiment digestate fertilization treatments supplying 90, 180, 270, 360 and 450 kg N ha^?1 were compared with untreated plots and plots fertilized with 180 kg N ha^?1 of mineral N fertilizer. Swards fertilized with digestate produced higher biomass yield compared with the control. The same rate of nitrogen fertilizer (180 kg N ha^?1) supplied as digestate and from mineral fertilizers gave similar results on biomass yield. Herbage in swards fertilized with digestate contained less nitrogen, but the C:N ratio was much more suitable for biogas production. Digestate fertilization resulted in higher concentrations of cellulose and hemicellulose in biomass and lower contents of the inhibitors of anaerobic digestion—sulphur, calcium, magnesium and phosphorus—compared with those of swards receiving mineral fertilizers.     

‘MaxClover’ grazing experiment: I. Annual yields,botanical composition and growth rates of six dryland pastures over nine years

Six dryland pastures were established at Lincoln University, Canterbury, New Zealand, in February 2002. Production and persistence of cocksfoot pastures established with subterranean, balansa, white or Caucasian clovers, and a perennial ryegrass‐white clover control and a lucerne monoculture were monitored for nine years. Total annual dry‐matter (10.0–18·5 t DM ha^?1) and sown legume yields from the lucerne monoculture exceeded those from the grass‐based pastures in all but one year. The lowest lucerne yield (10 t ha^?1 yr^?1) occurred in Year 4, when spring snow caused ungrazed lucerne to lodge and senesce. Cocksfoot with subterranean clover was the most productive grass‐based pasture. Yields were 8·7–13·0 t DM ha^?1 annually. Subterranean clover yields were 2·4–3·7 t ha^?1 in six of the nine years which represented 26–32% of total annual production. In all cocksfoot‐based pastures, the contribution of sown pasture components decreased at a rate equivalent to 3·3 ± 0·05% per year (R² = 0·83) and sown components accounted for 65% of total yield in Year 9. In contrast, sown components represented only 13% of total yield in the ryegrass‐white clover pastures in Year 9, and their contribution declined at 10·1 ± 0·9% per year (R² = 0·94). By Year 9, 79% of the 6.6 t ha^?1 produced from the ryegrass‐white clover pasture was from unsown species and 7% was dead material. For maximum production and persistence, dryland farmers on 450–780 mm yr^?1 rainfall should grow lucerne or cocksfoot‐subterranean clover pastures in preference to ryegrass and white clover. Inclusion of white clover as a secondary legume component to sub clover would offer opportunities to respond to unpredictable summer rainfall after sub clover has set seed.     

Species interactions in a grassland mixture under low nitrogen fertilization and two cutting frequencies. II. Nutritional quality

Mixtures and pure stands of perennial ryegrass, tall fescue, white clover and red clover were grown in a three‐cut and a five‐cut system in southern Norway, at a low fertilization rate (100 kg N ha^?1 year^?1). The nutritional quality (annual weighted averages) of the dried forage from the two‐first harvesting years was analysed. There was no significant effect of species diversity on crude protein (CP) concentration. In the three‐cut system, we found a significant species diversity effect leading to 10% higher concentrations of acid detergent fibre (ADF), 20–22% lower concentrations of water‐soluble carbohydrate (WSC) and 4% lower net energy for lactation (NE_L) concentrations in mixtures compared with pure stands (averaged across the two‐first years). In the five‐cut system, similar effects were seen in the first year only. This diversity effect was associated with a reduction in WSC and NE_L concentrations and an increase in ADF, NDF and CP concentrations in the grass species, and not in red clover, when grown in mixtures. This is thought to be a combined result of better N availability and more shading in the mixtures. Species diversity reduced the intra‐annual variability in nutritional quality in both cutting systems.     

Effect of a stay-green locus on the dry-matter yield, nitrogen yield and shoot density of perennial ryegrass

A rare stay‐green allele transferred from meadow fescue (Festuca pratensis L.) to perennial ryegrass (Lolium perenne L.) has improved both the colour of turf and the nutritive value of herbage. In this study its effect on shoot density and forage yield was assessed. Equivalent populations of perennial ryegrass were constructed with and without the stay‐green allele, following eight generations of backcrossing to perennial ryegrass. The stay‐green population, the normal population and the cv. AberStar were compared over two harvest years (2005 and 2006) in a field experiment with six application rates of N fertilizer (100, 200, 300, 400, 500 and 600 kg ha^?1 annually). There were no significant interactions between level of N fertilizer and population in any of the traits measured. The mean annual dry‐matter (DM) yield over all populations and fertilizer levels was 6·45 t ha^?1 lower in the second harvest year. Mean annual DM yields over all fertilizer levels of the normal population were higher than, or equal to, AberStar while those of the stay‐green population were significantly (proportionately 0·10–0·13) lower than the normal population. In 2005, the mean total yield of N in the herbage of the stay‐green population was 0·09 lower than that of the normal population and the mean concentration of N over all harvests was 1·5 g kg^?1 DM higher. The shoot density of the stay‐green population after the last harvest in November 2006 was 0·18 lower than that of the normal population (3689 and 4478 shoots m^?2 respectively).     

The efficiency of nitrogen in cattle manures when applied to a double‐annual forage cropping system

The use of cattle manure (CM) for fertilization presents challenges for optimizing nitrogen (N) use. Our work aimed to assess N efficiencies, in a 6‐year experiment with three biennial rotations of four crops: oat–sorghum (first year) and ryegrass–maize (second year) in a rainfed humid Mediterranean area of Spain. Fertilization treatments included the following: control (no N), 250 kg mineral N ha^?1 year^?1 (250MN), three CM rates (supplying 170, 250 and 500 kg N ha^?1 year^?1) and four treatments where the two lowest CM rates were complemented with either 80 or 160 kg mineral N ha^?1 year^?1. Treatments were distributed randomly in each of three blocks. Maximum dry‐matter yield (~44–49 t ha^?1 rotation^?1) was achieved in the third rotation, and only the control and the 170CM yielded significantly less. Within the limitations of the EU Nitrate Directive, the N steady state supply of 170CM always requires a complement of mineral N (80 kg N ha^?1) to maximize N agronomic efficiency. The maximum N‐fertilizer replacement value (250CM vs. 250MN) was 0·67, without significant differences between the two treatments in other N‐related efficiency indexes, which indicates that plants took advantage of residual‐N effects. Nitrogen losses by leaching in the 250CM treatment were around 5–7% of the N applied. This reinforces the sustainability of manure recycling in long cropping seasons.     

Comparative effects of food processing liquid slurry and inorganic fertilizers on tanner grass (Brachiaria arrecta) pasture: grass yield,crude protein and P levels and residual soil N and P

This small‐plot field study evaluated food processing liquid slurry (FPLS) as a potential fertilizer for tanner grass (Brachiaria arrecta) production on an acidic loam soil. The treatments, arranged in a randomized complete block design with three replicates, consisted of an unfertilized control, inorganic fertilizer applied at 50 and 200 kg nitrogen (N) ha^?1 with and without phosphorus (P) at 50 kg P ha^?1, and FPLS applied at 50 and 200 kg N ha^?1. Compared to the unfertilized control, the FPLS applied at 200 kg N ha^?1 significantly increased grass dry‐matter yield (DMY), herbage crude protein (CP) and P content, and N and P uptake in the second of two trials and P uptake in both trials. However, DMY and contents, of CP and P were generally lower for the FPLS treatments compared to the inorganic fertilizers. Apparent N recovery was higher for the inorganic fertilizer treatments than FPLS treatments in trial 1, while apparent P recovery was similar among all treatments in both trials. The FPLS treatments did not significantly increase soil NO₃‐N and P concentrations, but increased NH₄‐N in the 0–15 cm layer. The results suggest that application of FPLS to tanner grass pastures is an alternative to its disposal in landfill.     

Milk yield and nitrogen excretion of dairy cows grazing binary and multispecies pastures

Three grazing experiments were carried out in late spring (early lactation), summer (mid‐lactation) and autumn (late lactation) to compare the effects of perennial ryegrass cultivar or grass species, sown in binary or multispecies mixtures, on milk yield and nitrogen excretion of dairy cows. Replicated groups of multiparous Holstein Friesian × Jersey cows were offered either a control or high‐sugar perennial ryegrass (Lolium perenne) or tall fescue (Festuca arundinacea) base grass in a binary mixture with white clover (Trifolium repens) or in a multispecies mixture with additional legumes, bromegrass (Bromus willdenowii) and forbs. During each 9‐day experiment, botanical composition, milk production and faecal and urine composition were measured. Milk solid (MS) yield for the control ryegrass, high‐sugar ryegrass and tall fescue grass types averaged, respectively, 1.53, 1.64 and 1.70 kg MS cow^?1 day^?1 for a binary mixture sward, compared with 1.65, 1.54 and 1.53 kg MS cow^?1 day^?1 for a multispecies sward. Legume content influenced milk production more than the number of species present in a mixture. There was lower urine N concentration from a multispecies sward compared with a binary mixture. Urine N concentration of cows grazing the control ryegrass, high‐sugar ryegrass and tall fescue grass types averaged, respectively, 4.6, 5.3 and 6.8 g N L^?1 for a binary mixture, compared with 4.1, 3.9 and 3.9 g N L^?1 for a multispecies mixture. Feeding dairy cows on multispecies swards containing forbs presents an opportunity to reduce N losses without compromising milk yield.     

Effects of cutting frequency on plant production, N-uptake and N2 fixation in above- and below-ground plant biomass of perennial ryegrass–white clover swards

Nitrogen (N), accumulating in stubble, stolons and roots, is an important component in N balances in perennial ryegrass–white clover swards, and the effects of cutting frequency on the biomass of above‐ and below‐harvest height were studied during two consecutive years. Total dry matter (DM) and total N production, and N₂ fixation, were measured at two cutting frequencies imposed in the summers of two years either by cutting infrequently at monthly intervals to simulate mowing or by frequent cutting at weekly intervals to simulate grazing. Total DM production harvested was in the range of 3000–7000 kg DM ha^?1 with lower DM production associated with the frequent cutting treatment, and it was significantly affected by the different weather conditions in the two years. The higher cutting frequency also reduced the biomass below harvest height but the different weather conditions between years had less effect on stubble and, in particular, biomass of roots. The biomass of roots of white clover was significantly lower than that of roots of perennial ryegrass and remained at a relatively constant level (200–500 kg DM ha^?1) throughout the experiment, whereas the biomass of perennial ryegrass roots increased from 2400 kg DM ha^?1 in the year of establishment to 10 200 kg DM ha^?1 in the infrequent cutting treatment and 6650 kg DM ha^?1 in the frequent cutting treatment by the end of the experiment, giving shoot:root ratios of 4·7–16·6 and 0·5–1·6 for white clover and perennial ryegrass respectively. Annual N₂ fixation was in the range of 28–214 kg N ha^?1, and the proportion of N fixed in stolons and roots was on average 0·28. However, as weather conditions affect the harvested DM production and the shoot:root ratio, care must be taken when estimating total N₂ fixation based on an assumed or fixed shoot:root ratio.     

Dynamics of perennial ryegrass and white clover in sown swards in NW Greece

Seasonal dynamics of white clover and perennial ryegrass were examined in sown perennial ryegrass/white clover swards subject to a 2 × 2 factorial treatment combination of defoliation (rotational grazing by sheep and cutting) and nitrogen fertilizer application (0 and 40 kg N ha^–1 year^–1) in NW Greece. Sward surface height and percentage cover were measured before and after five defoliation periods in 1996 within permanent microplots (30 × 30 cm, divided into nine cells) in which white clover was either initially present or absent. Both white clover and perennial ryegrass achieved maximum height and cover in April–May. Defoliation treatment and whether white clover was present initially significantly affected height and cover of both species. Total plant cover was similar prior to all defoliation periods except in July, a time of drought. Cover of perennial ryegrass was greater where white clover was initially absent, but total plant cover was greater in microplots containing white clover and the extent of the differences varied during the year. In contrast, N fertilizer application had little effect on species cover, other than small reductions in white clover cover. When white clover was present in April, it was found in virtually every microplot cell until July, but if it was absent in April there was little colonization of the microplot.     

A comparison of the herbage productivity of Bromus wildenowii cv. Grasslands Matua with four cultivars of Lolium perenne when grown in association with Trifolium repens

In a small-plot trial five grass varieties bromegrass cv. Grasslands Matua, perennial ryegrass CVS. Melle (diploid) and Bastion, Condesa and Meltra (tetraploid) were established as grass/white clover swards with white clover cv. Menna. Productivity was measured under 6-weekIy cutting both without N fertilizer (N_o) and with 100 kg N ha^?1 applied in spring (N₁₀₀) Evaluation was made over 2 harvest years, 1986–87. Total mean annual production of herbage dry matter (DM) in the first harvest year at No and N_o was 5·07 t ha^?1 and 6·93 t ha^?1 respectively. In year 2, corresponding values were 11·81 and 12·67 t ha^?1. In year 1, Matua swards at N_o and N₁₀₀ yielded 5·08 and 6·65 t DM ha^?1 compared with 507 and 70 t DM ha^?1 for the mean of the four ryegrass varieties. In year 2, corresponding values were 12·90 and 12·29 for Matua and 11·54 and 12·78 for the four ryegrasses. In year 1, the digestable organic matter in the dry matter (DOMD) of the Matua swards was lower than that of Melle, Bastion and Condesa at N_O, particularly at the first cut. In year 2, differences in DOMD between treatments and varieties were not significant. The proportion of white clover was found to be higher in the N_o than the N₁₀₀ treatment, and also higher in year 2 in most treatments. For the _No treatment Matua swards had the highest proportion of white clover in year 1 (32% compared with 24% for the mean of the ryegrass varieties) but the lowest proportion in year 2 (27% compared with 60% for the ryegrasses). For the N_o treatment in year 1 clover production was also 43% higher, on average, from the tetraploid treatments than with Melle as the companion grass; for this comparison in year 2 the differences were not significant. It is concluded that Matua bromegrass/white clover swards receiving no N fertilizer may have a good potential under cutting management. However, the evidence from this trial is that in the second year the proportion of white clover is lower with Matua swards than with perennial ryegrass as the companion grass.     

Stocking rate effects on liveweight gain of ewes and their twin lambs when grazing subterranean clover–perennial grass pastures

Two grazing experiments were conducted on non‐irrigated tall fescue–subterranean clover and cocksfoot–subterranean pastures subject to summer‐dry conditions in Canterbury, New Zealand, to measure the effect of low (8·3–10 ewes and their twin lambs ha^?1) vs. high (13·9–20 ewes and their twin lambs ha^?1) stocking rates (SR) on lamb and ewe liveweight gain in spring. In tall fescue–subterranean clover pasture, lambs grew faster at low (374 g per head d^?1) than high (307 g per head d^?1) SR, but total liveweight gain per ha was greater at high (12·3 kg ha^?1 d^?1) than low (7·5 kg ha^?1 d^?1) SR. In successive years in spring in cocksfoot–subterranean clover pastures, lambs grew faster at low (327, 385 g per head d^?1) than high (253, 285 g per head d^?1) SR but total liveweight gain per ha was greater at high (7·26, 7·91 kg ha^?1) than low (5·43, 6·38 kg ha^?1) SR. These studies indicate that in summer‐dry areas, subterranean clover‐based pastures will support high twin lamb growth rates in spring, with lower SR leading to higher lamb growth rates and more lambs reaching slaughter weights before the onset of dry conditions.     

The value of timothy (Phleum pratense L.) in ryegrass — timothy mixtures managed to simulate intensive grazing

The performance of timothy in mixtures with perennial ryegrass was assessed under a simulated intensive grazing management over two harvest years in 1974–75. Three seed rates of S23 perennial ryegrass were factorially combined with three rates of Scots timothy and compared with pure stands of each grass. All were sown with Huia white clover. When cut six times at monthly intervals and with an annual N input of 350 kg ha^?1, there were no significant differences in total DM production in either year. The 2-year mean DM yield for the nine mixtures and six pure swards was 9·77 t ha^?1 (range 9·34–10·16). Compared with the pure ryegrass swards, in both years the ryegrass-timothy mixtures produced earlier spring growth but were significantly lower yielding at the second cut. Over the first five cuts the proptortion of timothy in the three mixtures with 22·4 kg ha^?1 ryegrass seed averaged 26% in the first year and 37% in the second. Corresponding calculated mean DM yields of timothy were 2·75 and 3·00 t ha^?1. It is concluded that an early timothy variety is capable of competing with a late-heading perennial ryegrass in frequently cut swards managed to simulate intensive grazing. The strong development of timothy in the dry summer of 1975 suggests that in mixtures of late perennial ryegrass varieties, an early variety of timothy should be beneficial for its spring growth in grazed swards.