Photosynthesis,growth and yield of soybean and maize in a tree-based agroforestry intercropping system on the Loess Plateau

Agroforestry is the most effective way to restore the disturbed lands on the Loess Plateau and to develop the poor local economy. In order to maximize the potential benefits of tree-based intercropping systems, photosynthesis, growth and yield of soybean and corn were studied by measuring photosynthetic active radiation (PAR), plant water deficit and soil moisture in a 4-year-old plantation of walnut (Juglans regia L.) and plum (Prunus salicina) grown at a spacing of 5 m × 3 m on the Loess Plateau. The effects of tree competition significantly reduced PAR, net assimilation (NA), growth and yield of individual soybean or corn plants growing nearer (1 m near tree row) to tree rows. NA was highly correlated with growth and yield of the both crops. These correlations were higher for corn than soybeans, with corn, rather than soybeans being more adversely impacted by tree shading. Plum, rather than walnut had the greatest competitive effect on PAR and NA. Daily plant water deficits were non-significantly and poorly correlated with NA and growth and yield of the both crops. However, soil moisture (20 cm depth) was significantly correlated with biomass and yield of both crops. Possible remediation strategies are discussed to reduce tree competitive interactions on agricultural crops.     

Soil water dynamics in cropping systems containing <Emphasis Type="Italic">Gliricidia sepium</Emphasis>, pigeonpea and maize in southern Malawi

The water dynamics of cropping systems containing mixtures of Gliricidia sepium (Jacq.) Walp trees with maize (Zea mays L.) and/or pigeonpea (Cajanus cajan L.) were examined during three consecutive cropping seasons. The trees were pruned before and during each cropping season, but were left unpruned after harvesting the maize; prunings were returned to the cropping area in all agroforestry systems to provide green leaf manure. The hypothesis was that regular severe pruning of the trees would minimise competition with crops for soil moisture and enhance their growth by providing additional nutrients. Neutron probe measurements were used to determine spatial and temporal changes in soil moisture content during the 1997/98, 1998/99 and 1999/00 cropping seasons for various cropping systems. These included gliricidia intercropped with maize, with and without pigeonpea, a maize + pigeonpea intercrop, sole maize, sole pigeonpea and sole gliricidia. Soil water content was measured to a depth of 150 cm in all treatments at 4–6 week intervals during the main cropping season and less frequently at other times. Competition for water was apparently not a critical factor in determining crop performance as rainfall exceeded potential evaporation during the cropping season in all years. The distribution of water in the soil profile was generally comparable in all cropping systems, implying there was no spatial complementarity in water abstraction by tree and crop roots. However, available soil water content at the beginning of the cropping season was generally lower in the tree-based systems, suggesting that the trees continued to deplete available soil water during the dry season. The results show that, under rainfall conditions typical of southern Malawi, the soil profile contains sufficient stored water during the dry season (ca. 75–125 mm) to support the growth of gliricidia and pigeonpea, and that gliricidia trees pruned before and during the cropping season did not deleteriously compete for water with associated crops. Water use efficiency also appeared to be higher in the tree-based systems than in the sole maize and maize + pigeonpea treatments, subject to the proviso that the calculations were based on changes in soil water content rather than absolute measurements of water uptake by the trees and crops.     

Exploring below ground complementarity in agroforestry using sap flow and root fractal techniques

Indices of shallow rootedness and fractal methods of root system study were combined with sapflow monitoring to determine whether these ‘short-cut’ methods could be used to predict tree competition with crops and complementarity of below ground resource use in an agroforestry trial in semi-arid Kenya. These methods were applied to Grevillea robusta Cunn., Gliricidia sepium (Jacq.) Walp., Melia volkensii Gürke and Senna spectabilis syn. Cassia spectabilis aged two and four years which were grown in simultaneous linear agroforestry plots with maize as the crop species. Indices of competition (shallow rootedness) differed substantially according to tree age and did not accurately predict tree:crop competition in plots containing trees aged four years. Predicted competition by trees on crops was improved by multiplying the sum of proximal diameters squared for shallow roots by diameter at breast height², thus taking tree size into account. Fractal methods for the quantification of total length of tree root systems worked well with the permanent structural root system of trees but seriously underestimated the length of fine roots (less than 2 mm diameter). Sap flow measurements of individual roots showed that as expected, deep tap roots provided most of the water used by the trees during the dry season. Following rainfall, substantial water uptake by shallow lateral roots occurred more or less immediately, suggesting that existing roots were functioning in the recently wetted soil and that there was no need for new fine roots to be produced to enable water uptake following rainfall. This revised version was published online in June 2006 with corrections to the Cover Date.     

Resource competition across the crop-tree interface in a maize-silver maple temperate alley cropping stand in Missouri

In order to improve the management of temperate alley cropping, it is important to study the growth and physiological responses of plants arising from competition across the crop-tree interface. Maize (Zea mays L.) was established between rows of seven-year-old silver maple (Acer saccharinum L.) trees in north-central Missouri, USA with four imposed treatments: (1) an unmodified control with a standard rate of N fertilization (179.2 kg N (as NH₄NO₃) ha⁻¹), (2) trenching with root barrier installed, (3) supplemental fertilization treatment (standard N + 89.6 kg ha⁻¹ N), and (4) a combination of trenching with root barrier and supplemental fertilization. Whereas soil N status had little effect on maize physiology and yield at the interface, competition for soil water was substantial in both years. Without a root barrier, soil water content, predawn and midday water potential, and midday net photosynthesis of maize plants adjacent to the tree row were reduced compared with those of plants in the alley center, but no differences across the maize crop were evident in the presence of a barrier. Grain yield of border row maize plants lacking an adjacent barrier was depressed compared with that for maize plants with a root barrier present (8.42 vs. 6.59 Mg ha⁻¹ in 1997; 5.38 vs. 3.91 Mg ha⁻¹ in 1998). However, the barrier did not completely restore yield to that in the alley center, suggesting that reductions in light near the tree row also limited production. Top ear height showed a similar pattern of response to the presence of a root barrier. Silver maple trees responded to root barrier installation with reduced annual diameter growth and reduced water status on some sample days. This revised version was published online in June 2006 with corrections to the Cover Date.     

Competition for water in a pecan (Carya illinoensis K. Koch) – cotton (Gossypium hirsutum L.) alley cropping system in the southern United States

Understanding the belowground interactions between trees and crops is critical to successful management of agroforestry systems. In a study of competition for water in an alley cropping system consisting of pecan (Carya illinoensis) and cotton (Gossypium hirsutum) in a sandy loam soil (Rhodic Paleudult) in Jay, Florida, root systems of the two species were separated by trenching to 120 cm depth. A polyethylene barrier was installed in half of the plots. Spatial and temporal variations in soil water content, root distribution and water uptake by both species, and leaf area development and height of cotton were measured. Interspecific competition for water was greater in the non-barrier treatment near tree rows than at the alley center. Competition became evident 3 to 4 weeks after emergence of cotton and increased during the following 7 to 8 weeks. Compared with the non-barrier treatment, the barrier treatment had higher soil water content and better growth of cotton (height, leaf area, and fine root biomass). Cotton lint yield in the barrier treatment (677 kg ha^–1) was similar to that in a sole-crop stand, but higher than in the non-barrier (502 kg ha^–1) treatment. Lint production efficiency of plants was higher in the interior rows in the non-barrier treatment (0.197 kg lint per square meter of leaf area, compared to 0.117 kg in the barrier treatment). The results suggest that trenching or even deep disking parallel to the tree row may reduce competition for water, but the impact on tree growth cannot be established from this study. This revised version was published online in June 2006 with corrections to the Cover Date.     

Water use assessment in alley cropping systems within subtropical China

Alley cropping systems may influence soil water movement and the water budget because of its complex interactions between crop and tree rooting systems. The objective of this paper was to evaluate water balance and water competition in an alley cropping system, consisting of deciduous tree wild jujube (Choerospondias axillaris) and economic crop peanut (Arachis hypogaea) within subtropical China. Five treatments (20- by 6-m plots) with three replications were included in this study. The treatments were monoculture peanut cropping (P), monoculture younger trees (T1), monoculture older trees (T2), peanut intercropped with younger trees (T1P), and peanut intercropped with older trees (T2P). A multi-layered water balance model, with water movement between soil layers, was implemented by the measurement of soil water potential using sets of tensiometers during the periods from March 1999 to December 2002. The spatial and temporal variations of soil water regime indicated that the trees used soil water below the 60-cm soil depth and alleviated the water stress. The direction of soil water movement indicated that soil water moved to the tree row, which indicated that trees competed with peanuts for water, especially during the seasonal drought period. Water competition was related to the tree spacing and tree age. Compared to the tree monoculture systems, the alley cropping system significantly influenced water budget components and water use patterns, as indicated by the increased evapotranspiration (6–11%), and decreased net drainage (7–45%), water storage (6–29%), and runoff (50–60%). Furthermore, alley cropping systems encouraged the rapid growth of trees, and depressed the biomass and yield of peanuts by 20–50% associated with tree shading effects. The results suggest that competition for water and light must be taken into account when optimizing the alley cropping system.     

Tree species and pruning regime affect crop yield on bench terraces in SW Uganda

Integration of trees on farms may exert complementary or competitive effects on crop yield. This 4 year study examined novel systems in which Alnus acuminata (alnus), Calliandra calothyrsus (calliandra), Sesbania sesban (sesbania) or a mixture of all three were grown on the degraded upper part of bench terraces in Uganda; beans or maize were grown on the more fertile lower terrace during the short and long rains. Three pruning treatments (shoot, root or shoot + root pruning) were applied to the tree rows adjacent to the crops; shoot prunings were applied as green manure to the woodlot from which they came. Pruning increased survival in calliandra and reduced survival in sesbania; alnus was unaffected. Pruning reduced tree height and stem diameter in alnus, but did not affect calliandra or sesbania. Maize yield adjacent to unpruned calliandra, alnus and sesbania or a mixture of all three was reduced by 48, 17, 6 and 24% relative to sole maize. Shoot pruning initially sustained crop performance but shoot + root pruning became necessary when tree age exceeded 2 years; shoot + root pruning increased maize yield by 88, 40, 11 and 31% in the calliandra, alnus, sesbania and tree mixture systems relative to unpruned trees. Bean yield adjacent to unpruned calliandra, alnus, sesbania and the tree mixture was 44, 31, 33 and 22% lower than in sole crops and pruning had no significant effect on crop yield. The results suggest that sesbania fallows may be used on the upper terrace without reducing crop yield on the lower terrace, whereas pruning of alnus is needed to sustain yield. Calliandra woodlots appear to be unsuitable as crop yield was reduced even after pruning.     

Introduction of Raikhanim (Ficus semicordata) in a Maize and Finger-Millet Cropping System: An Agroforestry Intervention in Mid-Hill Environment of Nepal

Fodder trees are integral part of farming system in the hills of Nepal, but designed agroforestry interventions targeted to particular trees and crops are not widely available. This paper examines the joint productivity of an agroforestry practice in which Raikhanim (Ficus semicordata) is planted in a maize (Zea mays) and finger-millet (Eleusine coracana) cropping system at Keware Bhanjyang of the western mid-hills of Nepal. Raikhanim seedlings were planted in a row on terrace risers 2, 4 and 6 m apart in ordinary farming conditions, in a randomized block design with three replications. Maize and finger-millet were grown on the terraces as intercrops with a control plot without trees on risers in each replicate. Growth parameters of Raikhanim—height, diameter at 30 cm above ground (D₃₀) and survival rate—were recorded annually in December until trees were lopped for fodder biomass, and crop yields were measured to determine tree-crop interaction effects. Tree height and D₃₀ differed significantly between spacings until trees reached the lopping stage 3½ years after planting, with the highest growth in 4 m spacing. Tree lopping checked the height growth but the diameter growth continued to increase and differed among spacings after lopping. Fodder biomass increased with tree age and was highest under 4 m spacing (7.294 t/ha) followed by 6 m (5.256 t/ha) and 2 m (3.84 t/ha). Finger-millet yield in the experimental plots decreased with tree age due to shading effects, while maize yield was not substantially affected. Among spacings, control plots produced the highest finger-millet yield (1,624 kg/ha) while the 6 m spacing produced the highest maize yield (2,463 kg/ha). It is concluded that planting Raikhanim at 6 m intervals will produce additional fodder without significant effect on maize yield and only a moderate effect on finger-millet yield. The agroforestry practice of planting fodder trees on under-utilised terrace risers is a viable option for mid-hill farmers for simultaneous production of fodder and cereal crops while sustaining the hill farming system.     

Water use efficiency and uptake patterns in a runoff agroforestry system in an arid environment

Water is the most limiting factor for plant production in arid to semiarid regions. In order to overcome this limitation surface runoff water can be used to supplement seasonal rainfall. During 1996 we conducted a runoff irrigated agroforestry field trial in the Turkana district of Northern Kenya. The effects of two different Acacia saligna (Labill.) H. Wendl. tree planting densities (2500 and 833 trees per ha), tree pruning (no pruning vs. pruning) and annual intercrops (no intercrop vs. intercrop: Sorghum bicolor (L.) Moench during the first season and Vigna unguiculata (L.) Walp. during the second season) on water use were investigated. The annual crops were also grown as monocrops. Water consumption ranged from 585 to 840 mm during the first season (only treatments including trees). During the second season, which was shorter and the plants relied solely on stored water in the soil profile, water consumption was less than half of that during the first season. Highest water consumptions were found for non-pruned trees at high density and the lowest were found for the annual crops grown as monocrops. Tree pruning decreased water uptake compared to non-pruned trees but soil moisture depletion pattern showed complementarity in water uptake between pruned trees and annual intercrops. The highest values of water use efficiency for an individual treatment were achieved when the pruned trees at high density were intercropped with sorghum (1.59 kg m^–3) and cowpea (1.21 kg m^–3). Intercropping and high tree density increased water use efficiency in our runoff agroforestry trial. We ascribe the observed improvement in water use efficiency to the reduction of unproductive water loss from the bare soil.This revised version was published online in November 2005 with corrections to the Cover Date.     

Defining competition vectors in a temperate alley cropping system in the midwestern USA: 1. Production physiology

With renewed interest in the use of ecologically-designed, sustainable agricultural systems for temperate regions of the world, agroforestry is being proposed as an alternative to intensive production of crops in monocultures. However, the knowledge-base for understanding and managing complex, multi-strata systems worldwide is limited, particularly so for temperate regions. We examined an alley cropping system in the midwestern US where maize (Zea mays L.) is grown in alleys between tree rows of either black walnut (Juglans nigra L.) or red oak (Quercus rubra L.). During a course of ten years, crop yields in rows adjacent to tree rows declined by 50% or more. With the experimental introduction of barriers to separate tree and crop root systems, yields in the rows near trees were equal to those of the center row (and monoculture). Irrespective of a high correlation between photosynthetically active radiation and net photosynthesis, shading did not have a major influence on crop yield. At this stage of system development (11 year old trees), influence of incident PAR on crop yield seems to be minimal. Subsequent papers in this series examine the sharing of belowground resources between trees and crops to quantify the competitive interactions that impact crop yields and their implications for economic return to the farmer.This revised version was published online in November 2005 with corrections to the Cover Date.     

Soil organic carbon under a linear simultaneous agroforestry system in Uganda

The objective of this study was to quantify and compare the amount and distribution of soil organic carbon (SOC) under a linear simultaneous agroforestry system with different tree species treatments. Field work was conducted at Kifu National Forestry Resources Research Institute in Mukono District, Central Uganda, in a linear agroforestry system established in 1995 with four different tree species and a crop only control treatment. Soil samples were collected in 2006 at three depths; 0–25, 25–50, and 50–100 cm, before planting and after harvesting a maize crop. The results indicate that an agroforestry system has significant potential to increase SOC as compared to the crop only control. There was no significant difference in the amount of SOC under exotic and indigenous tree species. Among the exotic species, Grevillea robusta had higher SOC than Casuarina equisetifolia across the entire depth sampled. There is significant difference in SOC among the indigenous species, where Maesopsis eminii has more SOC than Markhamia lutea. Distance from the tree row did not significantly influence SOC concentration under any of the tree species. In selecting a tree species to integrate with crops that will sequester reasonable quantities of carbon as well as boost the performance of the crops, a farmer can either plant an exotic species or an indigenous. In this study, the soil under Grevillea robusta and Maesopsis eminii have the highest potential to store organic carbon compared to soil under other tree species.     

Microclimate of a natural pasture under planted Robinia pseudoacacia in central Appalachia, West Virginia

The conditions under which forages yield more under tree canopies than in open fields are not well understood. This study was conducted to determine how microclimate experienced by forages in central Appalachia is modified by black locust (Robinia pseudoacacia L.) tree canopies. The effect of tree row location relative to forage growing point was evaluated for its impact on soil water, photosynthetically active radiation (PAR), red/far-red ratio, and surface soil temperature. There was no consistent spatial dependency relating tree rows to soil water levels. While daily PAR decreased as the time under shade increased, the level of PAR under tree canopies nearly doubled as cloud cover increased from 0 to 25%. The red/far-red ratio decreased from 1.16 to 0.2 over forage growing between tree rows compared to forage within tree rows. Surface soil temperature remained nearly constant (1.5–2 °C increase) during sunny days under tree canopies but increased 8–12 °C by mid afternoon at unshaded sites depending on soil water levels. Forages under black locust trees experienced less extreme variation in both daily PAR and temperature than unshaded forages, thus reducing the metabolic cost of adaptation to extreme conditions. This revised version was published online in June 2006 with corrections to the Cover Date.     

Water and nitrogen dynamics in rotational woodlots of five tree species in western Tanzania

The objective of this study was to compare the effects of woodlots of five tree species, continuous maize (Zea mays L.) and natural fallow on soil water and nitrogen dynamics in western Tanzania. The tree species evaluated were Acacia crassicarpa (A. Cunn. ex Benth.), Acacia julifera (Berth.), Acacia leptocarpa (A. Cunn. ex Benth), Leucaena pallida (Britton and Rose), and Senna siamea (Lamarck) Irwin & Barneby). The field experiment was established in November 1996 in a completely randomized block design replicated three times. Maize was intercropped between the trees during the first three years after planting and thereafter the trees were allowed to grow as pure woodlots for another two years. Transpiration by the trees was monitored when they were 3 years old using sap flow gauges. Soil water content was measured using the neutron probe approach between November 1999 and March 2001. Soil inorganic N profiles were measured when the trees were four years old in all treatments. The results indicated that the trees transpired more water than natural fallow vegetation during the dry season. The difference was apparent at a depth of 35 cm soil, but was more pronounced in deeper horizons. The water content in the entire soil profile under woodlots and natural fallow during the dry period was 0.01 to 0.06 cm³ cm⁻³ lower than in the annual cropped plots. This pattern was reversed after rainfall, when woodlots of A. crassicarpa, A. leptocarpa, A. julifera, S. siamea and L. pallida contained greater quantity of stored water than natural fallow or continuous maize by as much as 0.00 to 0.02, 0.01 to 0.04, 0.01 to 0.04, 0.01 to 0.03 and 0.00 to 0.02 cm³ cm⁻³, respectively. Natural fallow plots contained the lowest quantity of stored water within the entire profile during this period. Transpiration was greatest in A. crassicarpa and lowest in L. pallida. All tree species examined were `scavengers' of N and retrieved inorganic N from soil horizons up to 2-m depth and increased its concentration close to their trunks. This study has provided evidence in semi-arid environments that woodlots can effectively retrieve subsoil N and store more soil water after rains than natural fallow and bare soil. This revised version was published online in June 2006 with corrections to the Cover Date.     

Growth and yield of maize and timber trees in smallholder agroforestry systems in Claveria,northern Mindanao,Philippines

On-farm experiments were conducted in the Philippines to study over a 4-year period the growth of two timber trees, gmelina (Gmelina arborea R. Br.) and bagras (Eucalyptus deglupta Blume), and their impact on the grain yield of intercropped maize. The experiment consisted of maize monocropping plots (control) and maize intercropped between trees planted in block (2 × 2.5 m), and hedgerow arrangement (1 × 10 m). Three maize crops were planted in the block plots before canopy closure, and seven maize crops were planted in the hedgerow and monocropping plots. Maize grain yield in the hedgerow and in the block arrangement with gmelina were respectively 37% (16.58 tons ha⁻¹) and 68% (8.3 tons ha⁻¹) lower than in monocropping (26.21 tons ha⁻¹). In the plots with bagras, maize grain yield in hedgerow and in block arrangement were respectively 19% (24.8 tons ha⁻¹) and 66% (10.4 tons ha⁻¹) lower than in monocropping (30.6 tons ha⁻¹). For both tree species, the diameter at breast height (dbh) was greater in hedgerow than in block arrangement, with the difference being more pronounced with age. It was estimated that gmelina planted in hedgerows would produce 6–8 m³ ha⁻¹ of merchantable volume more than if planted in block. The study verifies the hypothesis that intercropping between widely-spaced trees rows (planted at 10 m or more) is more profitable and feasible to smallholders than either maize monocropping or woodlots, and concludes with recommendations on how to further improve the productivity of tree-intercropping systems.     

Aboveground and belowground competition between intercropped cabbage and young Eucalyptus torelliana

An expanding market for planted timber in the Philippines is providing a strong incentive for upland farmers to incorporate trees into their farming systems. Farmers often intercrop young timber species with well-fertilized annuals in expectation that inter- species competition for nutrients and light will be minimal while the trees are small, and that the trees will benefit from intensive nutrient and weed management of the intercrop. The relative level of aboveground and belowground competition in a vegetable/timber intercropping system was investigated in the uplands of Mindanao, the Philippines. Eight 5-m² microplots were established containing one nine-month-old Eucalyptus torelliana and four rows of cabbage (two on each of the north and south sides of the tree, 0.5 and 1.0 m from the stem base). The tree canopy shaded north rows. Monocrop cabbage microplots (2 m²) were also installed. Four tree/cabbage microplots and all cabbage-only plots were fertilized with ¹⁵ 5N-labeled ammonium sulfate (100 kg N ha⁻¹); remaining microplots received unlabeled fertilizer. Cabbage yields were reduced by 16% in the north rows when compared to the south rows, and by 15% in rows closer to the tree when compared to rows further from the trees. Belowground competition in the first cabbage row, possibly for moisture, is supported by the high proportion of tree roots found in the top 30 cm of soil. Competition did not appear to be for N or other nutrients. Foliar analyses revealed no row differences in mineral concentrations in cabbage, uptake of applied N, or percent of N derived from fertilizer. The modest amount of 15N found in aboveground tree parts (4.5% of N applied to four cabbage rows) improved overall N-use efficiency in the intercropped plots. An improved understanding of the tradeoffs between improved nutrient efficiency and depressed intercrop growth, as well as management options to reduce competition, will help farmers design systems to improve efficiency without increasing competition. This revised version was published online in June 2006 with corrections to the Cover Date.     

Herbage response to spacing of loblolly pine trees in a minimal management silvopasture in southeastern USA

Loblolly pine (Pinus taeda L.) silvopastures often are established and maintained on sites of poor soil fertility and minimal fertilizer input. Our objective was to determine whether row spacing affected yield, quality, and botanical composition of minimally managed herbage in loblolly pine early in the tree rotation. Plots were randomly located equidistant from bordering tree rows in each of eight alley width treatments that were 2.4, 3.6, 4.9, 7.3, 9.7, 12.2, 14.6 m wide, and no trees. Row spacing affected the yield, quality, and botanical composition of pasture five to six years in the rotation especially at densities exceeding 840 trees ha^–1. Botanical composition shifted from predominantly cool-season to warm-season grasses between annual first- and second-harvests, respectively, which caused seasonal differences in several yield and quality traits. Tall fescue (Festuca arundinacea Schreb.) production appeared to be unsustainable under minimal management. Herbage yield generally increased, but quality and minerals (crude protein, IVDMD, Ca, and P) tended to decrease with spacing. The 4.9 m row spacing was minimally acceptable for herbage yield and quality. System design should seek to balance tree-crop yield and quality within the context of management constraints and site productivity.This revised version was published online in November 2005 with corrections to the Cover Date.     

Tree row spacing affected agronomic and economic performance of Eucalyptus-based agroforestry in Andhra Pradesh, Southern India

The 3 × 2 m spacing currently used for eucalyptus plantations in the state of Andhra Pradesh, southern India does not permit intercropping from the second year. This discourages small landholders who need regular income from taking up eucalyptus plantations and benefiting from the expanding market for pulpwood. Therefore, on-farm experiments were conducted near Bhadrachalam, Khammam district (Andhra Pradesh) for over 4 years from August 2001 to November 2005 to examine whether wide-row planting and grouping of certain tree rows will facilitate extended intercropping without sacrificing wood yield. Eucalyptus planted in five-spatial arrangements in agroforestry [3 × 2 m (farmers’ practice), 6 × 1 m, 7 × 1.5 m paired rows (7 × 1.5 PR), 11 × 1 m paired rows (11 × 1 PR) and 10 × 1.5 m triple rows (10 × 1.5 TR)] was compared with sole tree stands at a constant density of 1,666 trees ha^?1. Cowpea (Vigna unguiculata) was intercropped during the post-rainy seasons from 2001 to 2004, and fodder grasses (Panicum maximum and Brachiaria ruziziensis) were intercropped during both the seasons of 2005. At 51 months after planting, different spatial arrangements did not significantly affect height and diameter at breast height (dbh). Total dry biomass of eucalyptus in different spatial arrangements ranged between 59.5 and 52.9 Mg ha^?1, the highest being with 6 × 1 m and the lowest with 10 × 1.5 TR, but treatment differences were not significant. The widely spaced paired row (11 × 1 PR) and triple row (10 × 1.5 TR) arrangements produced 62–73% of sole cowpea yield in 2003, 59–66% of sole cowpea yield in 2004, and 79–94% of sole fodder in 2005. In contrast, the 3 × 2 m spacing allowed only 17–45% of sole crop yields in these years. The better performance of intercrops in widely spaced eucalyptus was likely because of limited competition from trees for light and water. Intercropping of eucalyptus in these wider rows gave 14% greater net returns compared with intercropping in eucalyptus spaced at 3 × 2 m, 19% greater returns compared with that from sole tree woodlot and 263% greater returns compared with that from sole crops. Therefore, in regions where annual rainfall is around 1,000 mm and soils are fairly good, eucalyptus at a density of 1,666 plants per ha can be planted in uniformly spaced wide-rows (6 m) or paired rows at an inter-pair spacing of 7–11 m for improving intercrop performance without sacrificing wood production.     

Effects of groundcover management on soil properties, tree physiology, foliar chemistry and growth in a newly established Fraser fir (Abies fraseri [Pursh] Poir) plantation in Michigan, United States of America

Incorporating cover crops into Christmas tree plantations may potentially improve soil fertility, tree growth and quality and be an alternative to commercial nitrogen (N) fertilizers. However, cover crops may compete with the trees for water and other nutrients than N. This study was carried out to assess whether soil fertility, tree survival and growth could be improved by incorporating leguminous and non-leguminous cover crops into the Fraser fir (Abies fraseri) production system. Dutch white clover (Trifolium pratense), alfalfa (Medicago sativa) and perennial ryegrass (Lolium perenne) were grown in a newly established Fraser fir plantation using two cover crop management practices; no banding (NB) by growing each cover crop throughout the entire plot and banding (B) by creating a 61 cm-wide bare zone centered on the tree rows. A conventionally-managed system (CONV) was used as a control. The cover crop aboveground biomass and N content were assessed. Soil available N (NO₃ ⁻ and NH₄ ⁺) and N mineralization were measured at 0–15, 15–30 and 30–45 cm soil depths. Tree survival, growth, photochemical efficiency of photosystem II (F_v/F_m), branch water potential (Ψ_w) and foliar nutrients were also evaluated. Biomass production was as high as 13.9, 10.2 and 5.9 Mg DM ha⁻¹ year⁻¹ for clover, alfalfa and ryegrass, respectively. Cover cropping increased soil available N by 1.5- and 2.2-fold relative the CONV in the top soil layer in 2007 and 2008, respectively. Tree seedling survival and growth in the B and CONV systems were similar. In contrast, NB treatments resulted in poor seedling survival and growth relative to the B and CONV plots. Plant Ψ_w and F_v/F_m decreased significantly for A. fraseri seedlings on the NB treatments relative to their counterparts on the B and CONV plots. However, cover cropping had marginal effects on foliar nutrients. Cover cropping with banding can be an efficient strategy for maintaining productivity in Fraser fir Christmas plantations.     

The effect of young Leucaena leucocephala (Lam) De Wit hedges on the growth and yield of maize,sweet potato and cowpea in an agroforestry system in Sierra Leone

An investigation was undertaken at Senehun — Kamajei, a high rainfall region in Sierra Leone, to assess the effects of Leucaena leucocephala, on the growth and productivity of maize, cowpea and sweet potato. The experiment was laid out in a completely randomised design with four replicates of each treatment. The treatments and controls were: (i) pure crops of maize, cowpea, sweet potato, Leucaena clean weeded and unweeded and (ii) intercrops of Leucaena with the food crops, both (iii) with and without applied fertilizers after the first year.The growth of Leucaena was slow but nevertheless tended to reduce grain yields of maize, and tuber and vine yields of sweet potato in the rows in the immediate vicinity of the trees, especially in the nitrogen-treated plots. The maize, in particular, caused an improvement in the early height growth of the trees.     

Spatial heterogeneity of soil organic carbon in tree-based intercropping systems in Quebec and Ontario,Canada

Land use affects the carbon sequestration potential of soils across landscapes. Tree-based intercropping (TBI) systems where annual crops are grown between established tree rows are expected to exhibit spatial heterogeneity in the soil organic carbon (SOC) content due to differences in carbon input and decomposition rates of litter from trees and herbaceous plants. This study aimed to quantify variability in the SOC of TBI systems, compare the SOC content of TBI and nearby conventional agroecosystems, and determine if SOC was related to soil fertility. The TBI research sites were established 4 years (St. Paulin and St. Edouard, Quebec, Canada), 8 years (St. Remi, Quebec, Canada) and 21 years (Guelph, Ontario, Canada) before soil samples were collected for this study. The SOC content was greater within 0.75 m of the tree row than in the intercropped space at the St. Edouard and St. Remi sites. At the Guelph site, the SOC was spatially heterogeneous in plots with Norway spruce (Picea abies L.) but not hybrid poplar (Populus deltoides × P. nigra clone DN-177), probably due to litterfall distribution. Formerly a tree plantation, the TBI system at St. Remi contained 77% more SOC than a nearby conventional agroecosystem, while there was 12% more SOC in the TBI system than the conventional agroecosystem at Guelph. There was no difference in the SOC content of 4-year old TBI sites and nearby conventional agroecosystems. However, an increase in SOC at all TBI sites was positively related to the plant-available N concentrations, indicating the benefit of temperate TBI systems for soil fertility.