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1.
Genotype by environment interaction (GEI) for milk yield (ME) wasinvestigated by analyzing 40 140 first lactation records performed in 538 flocks. Flock–year effects from national genetic evaluation were used to define low (L), medium (M) and high (H) yielding environments. Differences observed between adjacent subsets were approximately 20% of the trait overall mean. Sire variance components estimated using multitrait analysis were 58, 82 and 149% of the estimate from the complete dataset (L, M and H, respectively) and genetic correlation coefficient between extreme subsets was 0.66, lower than genetic correlations obtained from randomly sampled subsets. Rank correlation coefficients between estimated transmitting abilities (ETA), obtained independently within L, M and H subsets for 71 AI rams having at least 10 daughters per subset, were lower than one (ranging from 0.42 to 0.56). These results strongly supported the existence of GEI for milk yield in Sarda sheep flocks. The influence of GEI on phenotypic responses in flocks differing in yield level could be due to a reduced expression of genes under conditions of restricted environmental opportunity. Splitting the selection scheme according to farming conditions does not seem a suitable solution for the Sarda breeding program. Thus, to provide farmers with genotypes suitable to exhibit a good phenotypic expression in different conditions, the selection across all environments, in which the descendants are expected to perform, still seems the best option.  相似文献   

2.
3.
Genotype x environment interaction (G x E) effects on live weaning weights of lambs were studied by using the 2 breeds Norwegian White sheep (NWS; heavy, long-tailed) and Spel sheep (Spel; lighter, short-tailed) as genetic groups (G). A total of 37,338 NWS lambs and 30,075 Spel lambs born from 1989 to 1999 on 40 farms that kept both breeds together were included in the analyses. Environment was characterized by farm x year (E). In a mixed linear model framework, significance of the random G x E effect and breed-specific environmental variances were tested by using a log-likelihood approach. Directions and magnitudes of the effect were described through variance component estimates. An across-genotype environmental correlation was also used. There was a significant G x E effect on lamb BW; significant breed differences were found for variance of flock x year effects, indicating different phenotypic plasticities with changing flock x year environments, with the NWS being more sensitive to environmental change. Further, the breed-specific residual variance was greater for NWS, indicating that the effects of environmental variation were larger for the weaning weights of the NWS breed within flock and year. Further, the correlation between flock x year effects for the 2 breeds was significantly different from unity (0.82 +/- 0.02), indicating that the common environment is "perceived" differently in the 2 breeds. The best environment for one breed is not necessarily best for the other breed, and vice versa. Solutions of flock x year effects may be used to describe how environmental characteristics such as climate and topography affect the production of different genotypes, and for clustering of environments, thus facilitating improvement of breeding programs and management schemes for domestic and wild ungulate populations.  相似文献   

4.
In this study the amount of genotype by environment interaction (G×E) for length of productive life of Swedish Red and White dairy cattle was studied. The environmental variables used were the herd-year averages of number of first parity cows, peak yield, protein yield, and productive life. Data were analysed using multi-trait models (using low and high quartile herds) and reaction norm models (using the whole continuous scale). Considerable G×E was found for the trait productive life between environments (herds) with short or long average productive life, and a genetic correlation of 0.74 was found. For productive life in relation to herd size, G×E was negligible, and the genetic correlation was 0.95. For productive life in relation to herd-year average of peak yield or protein yield some G×E was indicated by the reaction norm model, especially between extreme environments. The G×E between herds with short or long average productive life should be studied further and might need to be considered in breeding programmes for dairy cattle.  相似文献   

5.
The objectives of this study were to determine if sires perform consistently across altitude and to quantify the genetic relationship between growth and survival at differing altitudes. Data from the American Angus Association included weaning weight (WW) adjusted to 205 (n = 77,771) and yearling weight adjusted to 365 (n = 39,450) d of age from 77,771 purebred Angus cattle born in Colorado between 1972 and 2007. Postweaning gain (PWG) was calculated by subtracting adjusted WW from adjusted yearling weight. Altitude was assigned to each record based upon the zip code of each herd in the database. Records for WW and PWG were each split into 2 traits measured at low and high altitude, with the records from medium altitude removed from the data due to inconsistencies between growth performance and apparent culling rate. A binary trait, survival (SV), was defined to account for censored records at yearling for each altitude. It was assumed that, at high altitude, individuals missing a yearling weight either died or required relocation to a lower altitude predominantly due to brisket disease, a condition common at high altitude. Model 1 considered each WW and PWG measured at 2 altitudes as separate traits. Model 2 treated PWG and SV measured as separate traits due to altitude. Models included the effects of weaning contemporary group, age of dam, animal additive genetic effects, and residual. Maternal genetic and maternal permanent environmental effects were included for WW. Heritability estimates for WW in Model 1 were 0.28 and 0.26 and for PWG were 0.26 and 0.19 with greater values in low altitude. Genetic correlations between growth traits measured at different altitude were moderate in magnitude: 0.74 for WW and 0.76 for PWG and indicate possibility of reranking of sires across altitude. Maternal genetic correlation between WW at varying altitude of 0.75 also indicates these may be different traits. In Model 2, heritabilities were 0.14 and 0.27 for PWG and 0.36 and 0.47 for SV. Genetic correlation between PWG measured at low and high altitude was 0.68. Favorable genetic correlations were estimated between SV and PWG within and between altitudes, suggesting that calves with genetics for increased growth from weaning to yearling also have increased genetic potential for SV. Genetic evaluations of PWG in different altitudes should consider preselection of the data, by using a censoring trait, like survivability to yearling.  相似文献   

6.
Breeding for host resistance to parasites has become an imperative in many sheep industries. Because of the widespread use of AI in sheep breeding schemes, it is important to understand how the performance of offspring from rams varies in different flock environments, both for resistance to parasites and key production traits. This study used both variance component and reaction norm models to investigate the level of genotype x environment interaction for fecal egg count (FEC) and important Merino production traits in a range of flock environments in Australia. These flocks were linked by the use of common rams in a sire-referencing scheme. Both linear and quadratic polynomial reaction norm models were used. The heritability of these traits and the genetic correlation between them and FEC also was investigated using the reaction norm model. A contemporary group (CG) was defined by a flock, year, age class, sex, and paddock combination. Each CG environment was characterized by the mean value of any given trait for that CG. The recorded data used in the study were analyzed in a standardized form. Standardization for each trait was achieved within a CG by subtracting the CG mean from each observation and dividing by the CG SD. The genotype x environment effect accounted for <0.05 of the phenotypic variance for all traits. In most traits the heritability varied little across environments. The exceptions were FEC, BW, and both greasy and clean fleece weights, which had a higher heritability at the lower end of the environmental range. Fecal egg count also had a higher heritability in high-FEC environments. Genetic correlations between FEC and several key production traits were similar in the flock environments studied. Quadratic polynomial models and models with a variable residual fitted the data better than linear models. The genotype x environment effect for FEC and the genetic correlations between FEC and production traits were effectively zero; thus, sheep breeding programs for increased parasite resistance can be run effectively by ignoring these factors. Some account should be taken of the high heritabilities of FEC and fleece and BW in different flock environments.  相似文献   

7.
Real-time ultrasound technology offers the possibility of estimating carcass characteristics in live animals and represents a potential method for selection of breeding stocks. A total of 745 live lambs born during 2001–2003 into two flocks was used to estimate rib muscle and fat depth by ultrasound. Lambs came from 559 ewes and 97 rams of a fat-tailed breed, known as the ‘Barbarine’ in North Africa. Ultrasound measurements of external fat thickness (UFD) and muscle (UMD), taken at the 12–13th rib and palpation of body conditions (loin and tail scores) were made for 150 days until lambs 520 days old of lamb ages. Main results showed that UMD and UFD had the same trend as live weights from 150 to 240 days old. They decreased from 150 to 200 days old, and then increased. The same trend was seen for loin and tail scores. Average differences in live weights between male and female lambs were 1, 3 and 6 kg at 30, 90 and 120 days old, respectively. The highest muscle depth for males was reached at day 180 and the lowest fat depth was recorded between 180 and 200 days. Average loin scores became greater for males than females from 240 days of age. Male lambs had greater tail scores at all ages, from 150 to 520 days old. Differences became greater for ages more than 180 days, showing that males have a tendency to store more fat in their tails than females. Phenotypic correlation was 0.70 between muscle depth and loin scores, indicating that, at 180 days old, the Barbarine breed has more muscle and less fat. The optimum slaughter age was defined between 180 and 200 days old, producing carcasses with more muscle and less fat. Regression equations estimating carcass traits (UMD, UFD) at 180 and 240 days old were computed.  相似文献   

8.
Economically important traits are usually complex traits influenced by genes, environment and genotype-by-environment (G × E) interactions. Ignoring G × E interaction could lead to bias in the estimation of breeding values and selection decisions. A total of 1,778 pigs were genotyped using the PorcineSNP80 BeadChip. The existence of G × E interactions was investigated using a single-step reaction norm model for growth traits of days to 100 kg (AGE) and backfat thickness adjusted to 100 kg (BFT), based on a pedigree-based relationship matrix (A) or a genomic–pedigree joint relationship matrix (H). In the reaction norm model, the herd-year-season effect was measured as the environmental variable (EV). Our results showed no G × E interactions for AGE, but for BFT. For both AGE and BFT, the genomic reaction norm model (H) produced more accurate predictions than the conventional reaction norm model (A). For BFT, the accuracies were greater based on the reaction norm model than those based on the reduced model without exploiting G × E interaction, with EV ranging from 0.5 to 1, and accuracy increasing by 3.9% and 4.6% in the reaction norm model based on A and H matrices, respectively, while reaction norm model yielded approximately 8.4% and 7.9% lower accuracy for EVs ranging from 0 to 0.4, based on A and H matrices, respectively. In addition, for BFT, the highest accuracy was obtained in the BJLM6 farm for realizing directional selection. This study will help to apply G × E interactions to practical genomic selection.  相似文献   

9.
The genotype of an individual and the environment as the maternal ability of its dam have substantial effects on the phenotype expression of many production traits. The aim of the present study was to estimate the (co)variance components for worm resistance, wool and growth traits in Merino sheep, testing the importance of maternal effects and to determine the most appropriate model for each trait. The traits analyzed were Greasy Fleece Weight (GFW), Clean Fleece Weight (CFW), average Fibre Diameter (FD), Coefficient of Variation of FD (CVFD), Staple Length (SL), Comfort Factor (CF30), Weaning Weight (WWT), Yearling Body Weight (YWT) and Faecal worm Egg Count (FEC). The data were recorded during a 15-year period from 1995 to 2010, from Uruguayan Merino stud flocks. A Bayesian analysis was performed to estimate (co)variance components and genetic parameters. By ignoring or including maternal genetic or environmental effects, five different univariate models were fitted in order to determine the most effective for each trait. For CVFD and YWT, the model fitting the data best included direct additive effects as the only significant random source of variation. For GFW, CFW, FD, SL and CF30 the most appropriate model included direct-maternal covariance; while for FEC included maternal genetics effects with a zero direct-maternal covariance. The most suitable model for WWT included correlated maternal genetic plus maternal permanent environmental effects. The estimates of direct heritability were moderate to high and ranged from 0.15 for log transformed FEC to 0.74 for FD. Most of the direct additive genetic correlation (rg) estimations were in the expected range for Merino breed. However, the estimate of rg between FEC and FD was unfavourable (−0.18±0.03). In conclusion, there is considerable genetic variation in the traits analyzed, indicating the potential to make genetic progress on these traits. This study showed that maternal effects are influencing most of traits analyzed, thus these effects should be considered in Uruguayan Merino breeding programs; since the implementation of an appropriate model of analysis is critical to obtain accurate estimates.  相似文献   

10.
Abstract

Genotype by environment (G×E) interaction effects influence phenotypic expressions of a trait and may be of importance for sheep breeding. Interaction effects are more likely to be present when there are large environmental differences. Norwegian sheep usually graze mountain or forest pastures during summer. In this study, we estimate G×E interactions in Norwegian White Sheep as genetic correlation between area-specific traits (autumn lamb weight) in three ram circles located in two different counties; two in Buskerud in the south and one in Troms to the north of the country. Using data from the National Sheep Recording System, a bivariate animal model was fitted and genetic correlations for each trait were obtained. None of the correlations were significantly different from unity indicating the absence of G×E interaction effect for weaning weight. To gain further insight, studies should include a breeding-goal level aggregation of all traits thought to contribute to profitability.  相似文献   

11.
The objective of this study was to determine the magnitude of genotype by climate interaction (GCI) in the national genetic evaluation for weaning (WW) and yearling (YW) weights of Mexican Braunvieh cattle. The numbers of performance records and animals in the pedigree were 12,364 and 25,173 for WW, and 7,991 and 18,072 for YW, respectively. Performance records were clustered based on climatological variables into: dry tropic (DT), wet tropic (WT), and temperate (TE) climates. Animal models were used to estimate genetic parameters and predict breeding values in each of the climates. Bivariate analyses were carried out for pairwise combinations of climates on each trait, considering the same trait in different climates as a different trait. Criteria to evaluate GCI were genetic correlations (r g), correlations between predicted breeding values (r BV), and frequencies of coincidence (FC) in the ranking of the top 25 sires. Results of comparisons between pairs of climates were variable, depending on specific cases. For WW, the r g, r BV, and FC ranged from ?0.36 to 0.84, ?0.60 to 0.97, and 0.16 to 0.92, respectively; whereas for YW, they fluctuated between 0.23 and 0.99, 0.33 and 1.00, and 0.60 and 1.00, respectively. For both traits, the results suggest absence of GCI between DT and TE; however, GCI was detected in the other pairs of climates, where WT was involved. To maximize genetic progress, the joint genetic evaluation should be performed only for animals with performance data in DT and TE, whereas a separated evaluation is suggested for animals with performance records generated under WT conditions.  相似文献   

12.
With high variation in environmental conditions, especially during the extensive grazing season, genotype by environment interaction is potentially important to lamb production in Norway. Using breed as an indicator of genotype, the interaction between breed and environment, i.e. production level of farm and year calculated as solutions of a random farm*year variable, on 52 farms keeping both long-tailed Dala and short-tailed Spæl during 1989–98 was studied. The dataset included 48,946 Dala lambs and 38,299 Spæl lambs. The breed by environment interaction affected lamb autumn weights, and though Dala lambs were always heavier, the difference was substantially smaller at lower than at higher production levels: 1 kg increase in production level was related to Dala lambs’ weights increasing 0.09 kg more than the Spæl lambs’ weights. The results indicate that breed by environment interaction may be of importance, and potential bias in the data and need for further research is discussed.  相似文献   

13.
Estimates of genetic parameters for growth traits in Kermani sheep   总被引:3,自引:0,他引:3  
Birth weight (BW), weaning weight (WW), 6-month weight (W6), 9-month weight (W9) and yearling weight (YW) of Kermani lambs were used to estimate genetic parameters. The data were collected from Shahrbabak Sheep Breeding Research Station in Iran during the period of 1993-1998. The fixed effects in the model were lambing year, sex, type of birth and age of dam. Number of days between birth date and the date of obtaining measurement of each record was used as a covariate. Estimates of (co)variance components and genetic parameters were obtained by restricted maximum likelihood, using single and two-trait animal models. Based on the most appropriate fitted model, direct and maternal heritabilities of BW, WW, W6, W9 and YW were estimated to be 0.10 +/- 0.06 and 0.27 +/- 0.04, 0.22 +/- 0.09 and 0.19 +/- 0.05, 0.09 +/- 0.06 and 0.25 +/- 0.04, 0.13 +/- 0.08 and 0.18 +/- 0.05, and 0.14 +/- 0.08 and 0.14 +/- 0.06 respectively. Direct and maternal genetic correlations between the lamb weights varied between 0.66 and 0.99, and 0.11 and 0.99. The results showed that the maternal influence on lamb weights decreased with age at measurement. Ignoring maternal effects in the model caused overestimation of direct heritability. Maternal effects are significant sources of variation for growth traits and ignoring maternal effects in the model would cause inaccurate genetic evaluation of lambs.  相似文献   

14.
The ARR allele of the prion protein gene is associated with resistance to classical scrapie, while the ARQ allele is associated with susceptibility. The objective of this study was to estimate the effect, on lamb growth and carcass traits, of replacing an ARQ allele with an ARR allele. Pedigree rams (22 Suffolks, 18 Texels and 18 Charollais) were sourced in pairs, from the same flock (by the same sire where possible), with one member being ARR/ARR and the other being ARQ/ARQ and these rams were progeny tested through single-sire mating in six flocks. Effects of ram genotype on reproductive performance of ewes and on growth and carcass traits of progeny were estimated using mixed model procedures with sire as a random effect. Ram genotype had no effect on litter size at birth or on lamb survival as measured by the number of lambs reared. The differences between the performance of progeny by ARR/ARR and ARQ/ARQ sires provided an estimate of the effect of substituting an ARR allele for an ARQ allele. This allele substitution had no effect on birth weight, growth rate, weight at 5 or 14 weeks, weight or ultrasonic fat depth at 120 days of age or on carcass classification traits (conformation, fatness) for the Suffolk, Texel or Charollais breeds. Substituting an ARR allele for an ARQ allele had a negative effect on ultrasonic muscle depth at 120 days for the progeny of Suffolk rams (P = 0.01) but had a positive effect on ultrasonic muscle depth at 120 days for the progeny of Charollais rams (P = 0.02). The ARR substitution effect was negative for slaughter age in the Suffolk (P < 0.05) and positive for carcass weight in Texel progeny (P < 0.05). The magnitude of these effects was quite small, however, and the overall results support the hypothesis that substituting an ARR allele for an ARQ allele has no important negative effect on lamb growth or carcass traits in any of the breeds examined.  相似文献   

15.
全基因组关联分析(genome-wide association studies,GWAS)是研究家畜复杂经济性状和疾病遗传变异的有效方法,GWAS的核心是挖掘遗传变异与目标表型性状间的关系.随着牛全基因组测序工作完成,海量单核苷酸多态性(single nucleotide polymorphism,SNP)位点被标记...  相似文献   

16.
An understanding of influencing factors and genetic principles affecting the growth traits is needed to implement optimal breeding and selection programs. In this study, heritabilities (direct additive and maternal) of body weights at birth (BW0), 90 days (BW90) and 300 days (BW300) of age and average daily gains from birth to 90 days (ADG0-90), birth to 300 days (ADG0-300) and 90 days to 300 days (ADG90–300) of age in Boer goats were estimated on the basis of 1520 Boer goats at Boer Goat Breeding Station in Yidu, China, during 2002–2007. The parameters were estimated using a DFREML procedure by excluding or including maternal genetic or permanent maternal environmental effects, four analysis models were fitted in order to optimize the model for each trait. Influencing factors such as parity, litter size, kidding year and season, as well as sex of kids and some significant interactions among these factors were investigated as the fixed effects for the models. The results showed that the birth year and maternal genetic effects such as parity and litter size of dam were important determinants of the genetic parameter estimates for pre-weaning growth traits, and environmental effects such as birth year, season and sex of kids had some significant effect on post-weaning growth traits. The mean values and standard errors (SE) of direct additive heritability estimates calculated with the optimum model were 0.17 ± 0.07, 0.22 ± 0.08, 0.07 ± 0.07, 0.10 ± 0.08, 0.30 ± 0.12 and 0.08 ± 0.10 for BW0, BW90, ADG0-90, BW300, ADG0-300 and ADG90–300, respectively. For pre-weaning weights, correlation estimates between direct additive and maternal genetic (ra–m) effect were high and negative ranging from − 0.74 to − 0.86.  相似文献   

17.
The breeding goal for Simmental cattle is derived for intensively managed dairy farms. Its suitability for extensive farms was addressed by analysing possible genotype by environment interaction (G × E) between the management levels for first lactation milk yield traits. A first analysis was performed with the data collected from 300 000 purebred daughters of 278 second crop bulls born in Bavaria in 1993 and 1994. The farms were classified by herd‐year‐effect, using the sum of fat and protein yields into two levels of management, either with 33 or 10% quantiles, corresponding to approximately 100 000 cows and 30 000 cows, respectively. The comparison was based on ‘daughter yield’ deviations (DYD). Correlations between DYD of extensive and intensive environments were 0.90, 0.91 and 0.87 for milk, fat and protein yield (kg) for 33% quantiles, respectively. Corresponding correlations for 10% quantiles were 0.85, 0.83 and 0.77. Despite high correlations, 50 out of 149 sires showed significant differences between DYD in different environments. Bulls with higher DYD for milk yield on intensive farms were superior in all environments. For the second analysis extensive and intensive farms in northern and southern Bavaria were chosen at random. Approximately 20 000 cows in each management class were used for the estimation of genetic parameters. In both regions phenotypic and additive‐genetic variances were higher in the intensively managed herds. Likewise heritabilities were higher for fat and protein yield, but not for milk where higher heritabilities were observed in 33% quantiles. Genetic correlations between extensive and intensive environments were 0.97 and above (33% quantiles). Ten per cent quantiles led to lower genetic correlations (0.90–0.95). Although no serious re‐ranking effects of sires were evident, the scale effect and the differences in genetic parameters should be taken into consideration in practical breeding.  相似文献   

18.
A terminal sire Composite population was formed by mating Columbia rams to Hampshire-Suffolk crossbred ewes. Subsequent generations were produced by inter se mating. The objective was to compare Composite sheep to purebred Suffolks for numerous performance traits. Young ewes lambing at 1 yr of age were evaluated separately from mature ewes (2, 3, 4, and 5 yr old) of each population. Composite and Suffolk populations did not differ significantly in components of litter weaning weight per young ewe joined. Mature Composite ewes produced more wool than Suffolk ewes. Fertility of mature ewes did not differ significantly between populations. The greater prolificacy (P less than .01) of mature Suffolk ewes was offset by a lower survival to weaning (P less than .01), particularly for twin-born lambs. Consequently, mature Composite and Suffolk ewes did not differ for number weaned and litter weaning weight per ewe joined. The average birth weight of lambs born to young and mature Composite ewes was greater (P less than .02) than the average birth weight of Suffolk lambs. The average weaning weight of lambs born to young Composite ewes was greater (P less than .05) than that of lambs born to young Suffolk ewes. Breed differences were not detected for preweaning daily gains, weaning weights, and postweaning daily gains of ewe lambs born to mature ewes. Likewise, weights of ewes at 125 d of age and weights of mature ewes at breeding did not differ significantly between breeds.(ABSTRACT TRUNCATED AT 250 WORDS)  相似文献   

19.
Quantitative trait loci analyses were applied to data from Suffolk and Texel commercial sheep flocks in the United Kingdom. The populations comprised 489 Suffolk animals in three half-sib families and 903 Texel animals in nine half-sib families. Phenotypic data comprised measurements of live weight at 8 and 20 wk of age and ultrasonically measured fat and muscle depth at 20 wk. Lambs and their sires were genotyped across candidate regions on chromosomes 1, 2, 3, 4, 5, 6, 11, 18, and 20. Data were analyzed at the breed level, at the family level, and across extended families when families were genetically related. The breed-level analyses revealed a suggestive QTL on chromosome 1 in the Suffolk breed, between markers BM8246 and McM130, affecting muscle depth, although the effect was only significant in one of the three Suffolk families. A two-QTL analysis suggested that this effect may be due to two adjacent QTL acting in coupling. In total, 24 suggestive QTL were identified from individual family analyses. The most significant QTL affected fat depth and was segregating in a Texel family on chromosome 2, with an effect of 0.62 mm. The QTL was located around marker ILSTS030, 26 cM distal to myostatin. Two of the Suffolk and two of the Texel sires were related, and a three-generation analysis was applied across these two extended families. Seven suggestive QTL were identified in this analysis, including one that had not been detected in the individual family analysis. The most significant QTL, which affected muscle depth, was located on chromosome 18 near the callipyge and Carwell loci. Based on the phenotypic effect and location of the QTL, the data suggest that a locus similar to the Carwell locus may be segregating in the United Kingdom Texel population.  相似文献   

20.
A bio‐economic model including a computer program was developed where a large number of parameters can be modified to describe a great variety of production systems. Economic values for up to 27 traits can be calculated. Seasonal feeding is considered and protein requirement in addition to net energy may be taken into account when calculating feed requirement. Furthermore, the model allows for the evaluation of individual parts of the growth curve. The model was applied to the Czech Suffolk breed used in pure‐breeding. An extensive pasture system with spring lambing and sale of surplus progeny for slaughter after weaning or after a finishing period was assumed. The economic values (in Euro cent per ewe, per year and per genetic standard deviation of the trait) were as follows: 9.9 for birth weight, ?72.6 for mature weight, 142, 32.1 and 16.9 for growth rates till weaning, in rearing of breeding animals and in fattening, respectively, 19.9 and 101 for conception rate of female lambs and ewes, respectively, 690 for litter size per ewe lambing, 172 for productive lifetime of ewes, 402 and 503 for survival rate of lambs at lambing and till weaning, respectively, and 16.5 for fleece weight.  相似文献   

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