The silage yield potential of hybrid ryegrass over a 5-year period in Northern Ireland relative to other ryegrasses and timothy

The yield and persistency of grass species within a managed sward are a major consideration when determining species mixtures for either long-term or short-term sward management systems. Perennial ryegrass is lower yielding but more persistent than Italian ryegrass. Hybrid ryegrass, an interspecies cross between perennial and Italian ryegrass, may be higher yielding than perennial ryegrass and more persistent than Italian ryegrass. Therefore, the yield and persistency of hybrid ryegrass was investigated in Northern Ireland in five experiments, each harvested over five consecutive years, and compared with that of Italian ryegrass, perennial ryegrass and timothy. The performance of the five grass species were in the expected order; Italian ryegrass was higher yielding than hybrid ryegrass, which was higher than perennial ryegrass, with timothy the lowest, whereas perennial ryegrass and timothy had significantly higher sward densities than the hybrid and Italian ryegrasses. In addition, it was found that the rate of decline in yield and sward density with age was the same for all species. Consequently, this study indicated that the higher yielding Italian and hybrid ryegrasses could continue to out-perform perennial ryegrass for 5 years, opening the possibility of more extensive farming use of these species, particularly in Northern Ireland.     

High perennial ryegrass seeding rates do not negatively impact pasture sward persistence

Poor persistence of perennial ryegrass swards is a common problem; however, there is a lack of long‐term studies to understand the mechanisms associated with poor persistence. This study describes an experiment to test the hypothesis that high ryegrass seeding rates (>18 kg seed per ha) reduce long‐term population persistence because of smaller plant size and poorer survival during the first year after sowing. Four cultivars, representing four functional types of perennial ryegrass, were sown at five seeding rates (equivalent to 6, 12, 18, 24 and 30 kg seed per ha) with white clover in three regions of New Zealand. Swards were monitored for 5 years. No evidence was found to indicate a lack of persistence of ryegrass‐based swards sown at higher seeding rates. During the first year, swards sown at higher seeding rates had greater herbage accumulation (except at the Waikato site), greater ryegrass tiller density and greater ryegrass content. This initial impact of high seeding rates had largely dissipated by the fourth year, resulting in swards with similar annual herbage accumulation, tiller density and botanical composition. Similarly, there were relatively few differences among cultivars for these variables. Although high seeding rates did not negatively impact sward persistence, geographical location did, with strong evidence of ryegrass population decline at the Waikato site for all treatment combinations, some decline in Northland, and stable populations in Canterbury. It is possible that productive perennial ryegrass pastures can only be sustained for 4–5 years in some situations, even when the best ryegrass technology and management practices are used.     

THE USE OF PERENNIAL RYEGRASS SOWN AT 1 LB. PER ACRE IN A STANDARD SEEDS MIXTURE

An attempt was made in a survey-type trial to combine perennial ryegrass with cocksfoot, meadow fescue and timothy together in the same sward, without the association being dominated by the ryegrass. It is concluded from this work that such non-dominant ryegrass swards can be obtained by limiting the seeding rate of the ryegrass to 1 lb. per acre in a seeds mixture sown at the rate of about 20 lb. per acre.     

Productivity of mixtures of Italian ryegrass and red clover

Two red clover ( Trifolium pratense ) cultivars, Red Head (tetraploid) and Kuhn (diploid), were sown at a seed rate of 13 kg ha⁻¹ either alone or in mixture with Italian ryegrass ( Lolium multiflorum ) cv. RvP sown at seed rates of 10, 15, 20 or 30 kg ha⁻¹. RvP was also sown alone at a seed rate of 30 kg ha⁻¹ and received nil or 300 kg ha⁻¹ fertilizer a⁻¹ fertilizer N. All plots were established using the barley cultivar Midas sown at a seed rate of 100 kg ha⁻¹ as a nurse crop.
Neither clover cultivar nor ryegrass seed rate significantly influenced either dry matter harvested or botanical composition over the 3 harvest years. On average over all years the grass-clover mixtures produced 75% of the yield of the N-fertilized RvP, 125% of the clover monocultures and 225% of the unfertilized RvP. The red clover contribution to the total dry matter harvested of the mixtures averaged 45–60%. The dry matter concentrations of the mixtures were considerably higher than those of the pure clover stands. In the third year yields were markedly reduced in comparison with those in the first and second years.
It was concluded that Italian ryegrass can be a suitable companion grass for red clover. Its superior yielding capacity over other grasses such as perennial ryegrass or timothy under a conservation management can be coupled to advantage with red clover to give a sward which Is essentially stable, at least over a 2- to 3-year cropping period, although giving slightly reduced yields in the third year. Italian ryegrass-red clover mixtures, without the use of fertilizer N, can produce high DM yields of good quality herbage.     

The effects of separated cow slurry liquor on soil and herbage nitrogen in Phalaris arundinacea and Lolium perenne

Simulated swards of reed canary grass ( Phalaris arundinacea L ) and ryegrass received applications of separated cow slurry liquor in winter, at rates up to the equivalent of 500 m³ha⁻¹. Subsequent measurement of oxidized nitrogen in the soil under the different species indicated that lower levels of soluble forms of nitrogen were found under the reed canary grass than were under the ryegrass sward. At the first harvest after liquor application, the dry weight yield of reed canary grass responded positively to increasing applications of slurry liquor, but the ryegrass sward appeared to be damaged by very high rates of application. At later harvests, yields of the two species were similar, but over the season, apparent recovery of the nitrogen applied to the swards was greater in the reed canary grass than in ryegrass, suggesting that this species could be of value in the control of losses of nutrients from livestock wastes.     

CLOVER NITROGEN AND SWARD PRODUCTIVITY

Under mowing conditions in the absence of the animal, ultra-simple swards based either on ryegrass or cocksfoot gave large increases in yield of dry matter and nitrogen from the inclusion of white clover in the seeds mixture. Raising the clover seed rate from 1 to 3 lb./acre gave a further response in yield.
As the swards aged, the continuing effect of increased clover seed rate was dependent on applied nitrogen.
The rate of transference of nitrogen from clover to the soil depended upon the species of the associated grass, the clover seed rate, the age of the sward and the degree of reduction of the clover in the sward.
Suppression of clover led to an increase in soil nitrogen, particularly with high seed rates of clover.
Dry matter response to applied nitrogen (69 lb./acre/annum) was greater in cocksfoot than in ryegrass. Fertilizer nitrogen had little effect on the yield of nitrogen from the sward except where the high seed rate of clover had been used.     

Component yields and quality of binary mixtures of lucerne and perennial, Italian or short rotation hybrid ryegrass

Studies were conducted to characterize sward dynamics of lucerne ( Medicago sativa L.) and perennial, Italian or short rotation hybrid rye-grass ( Lolium perenne L. and L. multiflorum Lam.) binary mixtures under hay management. Lucerne/timothy ( Phleum pratense L.) mixtures were included as controls. The study was conducted on two farms in central Pennsylvania, USA on Hagerstown silt loams (fine, mixed, mesic hapludalf). Total dry matter yield, grass yield and yield of digestible dry matter were higher for tetraploid perennial and tetraploid Italian ryegrass mixtures than for timothy mixtures. Seasonal production differed for the three kinds of ryegrass. Italian ryegrass dry matter yields were higher in cuts 2 and 3 than perennial ryegrass yields. Grass yields represented 34%, 24% and 12% of total yields of the Italian, perennial or short rotation, and timothy mixtures, respectively. However, herbage in vitro dry matter digestibility of Italian was lower than that of perennial ryegrass.
The tetraploid perennial ryegrasses Grimalda and Reveille had high-ranking N concentrations more frequently than other cultivars, while Bar-vestra and Reveille tetraploid perennials and Bison short rotation ryegrass had high-ranking dry matter digestibility values more frequently than the other cultivars. Perennial ryegrass cultivars and Lipo tetraploid Italian ryegrass persisted well for 4 years and produced higher yields with a lower weed content than timothy mixtures. It is concluded that perennial ryegrasses and Lipo Italian ryegrass should be recommended over timothy, as companion grasses with lucerne.     

Effect of dairy slurry application rate and forage type on production,soil nutrient status and nitrogen‐use efficiency

Seven forage types (diploid and tetraploid perennial ryegrass, Italian ryegrass and hybrid ryegrass, a low‐input mixture of perennial ryegrass, cocksfoot, timothy and meadow fescue, a mixture of perennial ryegrass and white clover, and monoculture of red clover) were sown in late July 2004. Each received one of four rates of dairy cattle slurry in three annual applications by trailing shoe, which supplied average nitrogen (N) inputs of 0·0, 114·9, 204·8 and 301·2 kg N ha^?1 annum^?1. Treatments were cut either three or four times annually over four years. Average dry‐matter yield (DM) response to slurry N was 15·6 kg DM kg^?1 N. Lowest recovery of slurry N was in the second application each year (after first cut). The data suggest that slurry applied to Italian ryegrass, and also to swards containing legumes on soils with high phosphorus content, will produce a lower DM response to slurry N and result in a lower slurry N recovery than on swards of perennial ryegrass or cocksfoot‐dominant low‐input mixtures. Apparent recovery of slurry N was low at the second cut, especially when first‐cut yields had been high. To maximize slurry N recovery, application to regrowths with potentially slow rates of growth or high legume content should be avoided.     

Milk-production potential of different sward types in a temperate southern Australian environment

The milk-production potential of different sward types was measured in each of the four seasons of the year in a replicated experiment in south-west Victoria, Australia. Dairy cows were offered ad libitum allowances of a 'short-term winter' sward, based on Italian ryegrass (treatment STW), a 'long-term winter' sward, based on a winter-active tall fescue (treatment LTW), a 'long-term summer' sward, based on a summer-active tall fescue (treatment LTS) and a Control sward (perennial ryegrass) in four seasons (days in milk in parentheses): spring (November dairy cows, 124), summer (February, 227), autumn (May, 234) and winter (August, sixty-four). A 'short-term summer' sward, based on chicory and white clover (treatment STS), was also included in summer. There was a significant season × treatment interaction caused by a more gradual decline in milk yield from peak for cows grazing treatment LTS compared to the Control treatment in the transition period from spring to summer. In summer, cows grazing treatment STS produced more milk (1·41 kg fat plus protein per cow d⁻¹) than cows grazing all other treatments (0·92 kg per cow d⁻¹) because of the superior nutritive value of herbage of pre-grazing pasture and higher apparent dry-matter intakes. Swards based on alternative species to perennial ryegrass are capable of supporting milk production that is at least comparable over an annual cycle.     

The response of mini-swards of perennial ryegrass/white clover to simulated rainfall following slurry application

Perennial ryegrass/white clover mini-swards were used lo determine whether sward responses to slurry were modified by subsequent simulated rainfall (SR). Combinations of two SR volume rates (6.5 and 13 mm), three SR timings (3, 24 and 48 h after slurry application) and two slurry types (cattle, pig) were compared with a fertilizer control (no SR). Scorch, smother and growth of marked stolons and tillers were monitored after slurry application, and yields of both species were recorded. Slurry coverage of plastic squares indicated that smother may be greater on horizontally orientated leaves (clover). Reduced grass leaf growth and numbers of clover growing points in the week following slurry application were attributed mainly to smother, and possibly other phytotoxic effects of slurry. Reasons for the superior clover growth with cattle slurry arc unknown, although its higher P and K content may have contributed. Timing of rainfall is important in influencing sward responses to slurry application. Results suggest that application of slurry may be most beneficial when it immediately precedes rainfall and that longer intervals between slurry application and rainfall reduce grass and clover growth through either greater ammonia loss or slurry negative effects.     

The effect of sward structure as influenced by ryegrass genotype on bite dimensions and short-term intake rate by dairy cows

Abstract The effects of genotypic variation in ryegrasses on sward structure, bite dimensions and intake rate by dairy cows were investigated. Two experiments were conducted. In Experiment 1, swards were in a vegetative state whereas, in Experiment 2, they were partly reproductive and were taller with higher herbage mass but lower leaf proportion than in Experiment 1. Applicability of relationships between sward structure and bite characteristics, previously established from artificial or hand‐constructed swards, to field conditions were tested. Additional short‐term intake rates and/or sward structural characteristics were considered as indicators of potential intake for use in protocols for the evaluation of grass varieties. Four cultivars were studied: AberElan, Twins (diploid and tetraploid perennial ryegrasses respectively), Polly, a hybrid ryegrass (perennial × Italian ryegrass) and Multimo (Italian ryegrass), each established in 200‐m² plots in four replicated blocks. Herbage intake rate was determined by short‐term liveweight change (taking account of insensible weight loss) using 16 dairy cows allocated to four balanced groups with each plot grazed by one group for a 1‐h assessment period. One block was grazed per day, over a 4‐d experimental period, with each group grazing each variety in a complete crossover design. Sward characteristics and bite rate were also measured in both experiments. Bite dimensions were subsequently estimated, with bite depth being determined as a function of extended tiller height (ETH) in both experiments. Within both experiments, bite mass and intake rate did not differ significantly between swards of different cultivars despite swards containing Multimo generally having a higher ETH and water‐soluble carbohydrate concentration and lower green leaf mass, sward bulk density and neutral‐detergent fibre concentration than the other swards. However, bite depth was significantly higher (P < 0·01) in swards containing Multimo swards than in the others and, in Experiment 1, bite depth, as a proportion of ETH, was higher in swards containing Multimo and lower in those containing Twins than in the other two cultivars, whereas there was no difference in Experiment 2. Taking both experiments together, the mean bite depth was 0·5 of ETH with sward bulk density accounting for almost half the variance in the relationship between bite depth and ETH. The bulk density of the bite (bite mass per unit bite volume), measured in Experiment 2, followed a similar pattern to sward bulk density, increasing in the order Multimo, Polly, AberElan and Twins. It is concluded that the relationships between sward characteristics and bite dimensions, derived from artificial swards, are applicable to field swards, although the range in natural ryegrass sward characteristics is usually not as wide as in experiments using artificial swards. Lack of precision in the measurement of short‐term intake and in sward‐based measurements is likely to preclude their use in the evaluation of grass varieties.     

The effect of sward height on responses of mini-swards of perennial ryegrass/white clover to slurry application

The influence of sward height at the time of slurry application on sward responses to slurry was investigated using perennial ryegrass ( Lolium perenne )/white clover ( Trifolium repens ) mini-swards under greenhouse conditions. Pig slurry, cattle slurry and a fertilizer control were applied either to: swards cut 1 d before slurry application to heights of 2, 4 or 8 cm (CH2, CH4 or CH8); or to swards cut to 4 cm with regrowth intervals of 1, 4 or 8 d before slurry application (RI1, RI4 and RI8). Scorch, smother and growth of marked clover stolons and grass tillers were monitored after slurry application. Dry-matter yields of both species were recorded over two harvests. Electrolyte leakage from leaves was used to assess leaf damage. Both slurries increased leakage, which was greatest from clover leaves, with biggest increases caused by pig slurry. Yield responses to slurry application varied with plant species and sward height. Reduced grass growth after slurry application to long swards (CH8, RI4, RI8) was probably caused by greater smother. In short swards (CH2), grass leaf growth was not depressed following slurry application but grass yields were reduced. Ammonia volatilization losses may have been greater in both long and short swards, contributing to yield reductions. Yield responses to slurry were greatest in swards cut to 4 cm, 1 d before application.     

THE EFFECT OF CUTTING HEIGHT AND CUTTING FREQUENCY ON THE PRODUCTIVITY OF AN ITALIAN RYEGRASS SWARD

Three cutting heights, 1, 3 and 5 in. (2.5, 7.6 and 12.7 cm), and three cutting frequencies, 3, 6 and 10 weeks, were imposed on a pure sward of Italian ryegrass (Lolium multiflorum) cv. Irish. Lowering the cutting height increased the yield of DM, digestible DM, crude protein and crude fibre. Increasing the interval between cuts increased the yield of DM, digestible DM and crude fibre. The results are discussed in relation to other research findings and to the management of short-term leys of Italian ryegrass.     

Effects of different rates and timing of application of nitrogen as slurry and mineral fertilizer on yield of herbage and nitrate-leaching potential of a maize/Italian ryegrass cropping system in north-west Portugal

Efficient use of cattle-slurry to avoid nitrogen (N) leaching and other losses is important in designing intensive dairy systems to minimize pollution of air and water. The response in dry-matter (DM) yield of herbage and nitrate-leaching potential to different rates and timing of application of N as cattle slurry and/or mineral fertilizer in a double-cropping system producing maize ( Zea mays L.) silage and Italian ryegrass ( Lolium multiflorum Lam.) was investigated in north-west Portugal. Nine treatments with different rates and combinations of cattle slurry, and with or without mineral-N fertilizer, applied at sowing and as a top-dressing to both crops, were tested and measurements were made of DM yield of herbage, N concentration of herbage, uptake of N by herbage and amounts of residual soil nitrate-N to a depth of 1 m, in a 3-year experiment. Regression analysis showed that the application of 150 and 100 kg of available N ha⁻¹ to maize and Italian ryegrass, respectively, resulted in 0·95 of maximum DM yields of herbage and 0·90 of maximum N uptake by herbage. Residual amounts of nitrate-N in soil after maize ranged from 48 to 278 kg N ha⁻¹ with an exponential increase in response to the amount of N applied; there were higher values of nitrate-leaching potential when mineral-N fertilizer was applied. The results suggest that it is possible in highly productive maize/Italian ryegrass systems to obtain high DM yields of herbage for maize silage and Italian ryegrass herbage with minimal leaching losses by using slurry exclusively at annual rates of up to 250 kg available N ha⁻¹ (equivalent to 480 kg total N ha⁻¹) in three applications.     

PRODUCTIVITY OF ITALIAN AND PERENNIAL RYEGRASS MIXTURES

Italian ryegrass cv. Leda and perennial ryegrass cv. Presto were sown alone and in five mixtures containing different proportions of viable seed and compared with hybrid ryegrass cv. Manawa and Italian ryegrass cv. S22. Yields and herbage composition were measured five times in 1962 and 1963, with particular reference to the contribution from perennial ryegrass in the first year and the effects of Italian ryegrass on second-year production. Italian ryegrass dominated first-year production wbile perennial ryegrass dominated second-year production. Manawa was the top yielder in the first year, but both Manawa and S22 bebaved as Italian ryegrasses and were badly frosted and weedy in the second year. The mixtures were generally intermediate in yield between the components and tended towards the yield of the more vigorous component. Since the more vigorous component changed from cut to cut or from season to season, the sums of harvests sbowed some advantage for certain mixtures. Certain pairs of herbage varieties can be sown togetber witb yield advantage. Furtber physiological investigations are warranted and the breeder has a part to play in a search for complementary varieties.     

Impact of seeding rate on annual ryegrass performance

Annual ryegrass (Lolium multiflorum Lam.) is a primary forage resource for livestock producers throughout the south‐eastern USA during the winter‐growing season. It is important for livestock producers to begin grazing annual ryegrass as early as possible and any management practices maximizing early season production could be beneficial. To assess the impact of seeding rate on subsequent yield, yield distribution, quality, seedling density, and end‐of‐season plant and tiller density, a 2‐year study was initiated at four locations in Louisiana. Three annual ryegrass cultivars, varying in seed size, were established at four seeding rates based on pure live seed (PLS) rates of 400, 800, 1200 and 1600 PLS m^?2. There was no advantage in total yield from increasing seeding rates beyond 800 PLS m^?2. However, first‐harvest yields increased from 360 to 930 kg dry matter (DM) ha^?1 as seeding rate increased from 400 to 1600 PLS m^?2. Crude protein and neutral‐detergent fibre concentrations, and in vitro DM digestibility, were not affected by seeding rate. Seedling density and end‐of‐season plant numbers increased as seeding rate increased. However, stems per plant decreased as seeding rate increased, indicating compensatory tillering for the reduced plant numbers observed at the lower seeding rates. These results indicate first‐harvest yield can be increased by planting at higher seeding rates but total yields are not increased.     

THE EFFECT OF A RYEGRASS COMPANION GRASS AND THE VARIETY OF RED CLOVER ON THE PRODUCTIVITY OF RED-CLOVER SWARDS

An experiment was conducted to measure the effects of differrat ryegrass companion grasses and red-clover varieties on the productivity of red-clover swards. Three silage harvests per year were taken over a 2-year period. The addition of a companion grass increased total herbage yields; S24 perennial ryegrass gave the highest herbage yield over the two years, followed by Reveille perennial ryegrass. Because of lack of persistence, Tetila Italian ryegrass yielded poorly the second year. A companion grass had little effect on red-clover yields in the mixed swards but improved percentage digestibility of the OM of the total herbage and lowered the CP percentage. Its presence also reduced the ingress of unsown species. The variety of red clover used had little effect on total herbage yields or red-clover yields in the first harvest year. In the second year, Hungaropoly and Tilo persisted better and so gave higher total herbage yields and red-clover yields than Dorset Marl or Essex. The fall in total herbage yields from the first to the second year was entirely due to a fall in red-clover yield since yields of the ‘non-red clover’ fraction of the total herbage increased. A red-clover/grass sward may have advantages over a pure red-clover sward nutritionally, for silage-making and for its effect in diluting the oestrogenic activity of a pure clover sward. A major reappraisal of the role and potential of red-clover swards in the UK is warranted because of their many valuable attributes, particularly their ability to give high herbage yields of high nutritive value without the addition of fertilizer N. The improved persistency of some of the tetraploid varieties of red clover enhances the value of the plant.     

Tillering and leaf area index in grasses in the vegetative phase

The relationship between tillering of a vegetative grass stand and Leaf Area Index (LAI) has been studied during sward establishment of perennial ryegrass and Italian ryegrass with different seedling densities and nitrogen fertilizer rates and on established swards of tall fescue during winter and early spring after different dates of last cut in autumn.
The different experiments on different species led to the conclusion that in every instance the tillering rate slows down as soon as the LAI reaches a value of 3 and terminates rapidly at higher LAI, which corresponds with the almost complete extinction of light at the level of the tiller buds. The results confirm the direct role of light on morphogenesis previously demonstrated by several authors. The existence of such a limit of LAI helps to explain some contradictory effects of temperature or nitrogen on tillering.     

The diet ingested by sheep grazing swards differing in white clover and perennial ryegrass content

A series of twenty-four swards containing different proportions of white clover (0·20-0·25) and perennial ryegrass were created by using different seed mixtures, herbicide applications and previous cutting Frequencies. These swards were used to study the diet of oesophageally-fistulated wether sheep which grazed the various swards for a 30-min period after 1, 2 and 3 weeks of regrowth.
The proportion of white clover in the diet was generally greater than that in the sward. Fifty-seven percent of the variation in the proportion of white clover in the diet could be attributed to the proportion of white clover in the sward. White clover and perennial ryegrass leaf and stem were grazed to the same height and the proportion of white clover in the grazed horizon of the sward explained 83% of the variation in the proportion of white clover in the diet. The proportion of white clover in the diet was greater than the proportion in the grazed horizon of the sward in week 3 of regrowth, but not in weeks 1 and 2, and greater when the proportion of white clover in the grazed horizon was lower than 0·20. Both these observations were interpreted as indicating selection for white clover by the sheep within the grazed horizon.
There was a positive and linear relationship between the depth of the grazed horizon and sward height which, together with the relationship between the proportion of white clover in the grazed horizon and in the diet, would allow the prediction of the proportion of white clover of the diet from the height and the white clover content of the grazed horizon of the sward.     

How does the vertical and horizontal structure of a perennial ryegrass and white clover sward influence grazing?

Mixtures of perennial ryegrass ( Lolium perenne L.) and white clover ( Trifolium repens L.) sown in alternate rows or in a thoroughly mixed matrix were grazed by sheep, either continuously or during short grazing tests, and were used to investigate the influence of the vertical and horizontal components of the sward structure on defoliation by sheep.
In an experiment under continuous grazing, the defoliation intensity was greater for white clover compared with perennial ryegrass leaves (0·80 and 0·58 respectively). In spring, perennial ryegrass leaves were more defoliated than white clover leaves, whereas the reverse was observed in summer. The ratio of the proportion of white clover to perennial ryegrass leaves grazed was negatively correlated with the difference between the surface height of the perennial ryegrass and white clover rows in spring. In both spring and summer, white clover leaves of the same extended leaf length had a higher proportion of them grazed than perennial ryegrass leaves.
In another experiment, during short grazing tests with perennial ryegrass–white clover swards that were grazed at the same sward surface height and at the same white clover content as in the previous experiment, there were no significant differences in the proportion of white clover and perennial ryegrass leaves grazed between strips of the two species and thoroughly mixed structures. The proportion of white clover leaves grazed was higher than that of perennial ryegrass leaves.
These results show that the differential defoliation by sheep of perennial ryegrass and white clover leaves varies according to their vertical distribution in the mixed canopy, but is little affected by their horizontal distribution. Even small differences in sward surface height between mixed perennial ryegrass and white clover can affect diet selection by sheep to a rather large extent.