The role of coleoptile on barban sensitivity between wild oat and wheat

Differential sensitivity of wild oat (Avena fatua L.) and wheat (Triticum aestivum L.) to barban (4-chloro-2-butynyl-m-chlorocarbanilate) was highest when barban was applied to the coleoptile. The coleoptile acts as a physical and physiological barrier to reduce the concentration of free barban in the stem section where the sensitive meristematic sites are located. Metabolism of barban was higher ïn the coleoptile of tolerant wheat than in that of susceptible wild oat. Free barban concentration was higher in the stem of wild oat than in the stem of wheat after 24 hr, but after 48 hr, concentrations were similar. The coleoptile appears to enhance the differential sensitivity to barban between wild oat and wheat.     

Weed control in wheat with pyroxasulfone and its combinations with other herbicides

Field and pot investigations were conducted to determine the effectiveness of pyroxasulfone alone and its combinations with other herbicides against diverse weed flora of wheat (Triticum aestivum L.) including multiple herbicide-resistant (MHR) littleseed canarygrass (Phalaris minor Retz.). Applications of pyroxasulfone 100–127.5 g/ha as pre-emergence (PE) or early post-emergence (EPOE) @ 63.75 g/ha at 21–23 days after sowing (1 day before irrigation) were highly effective for control of grass weeds namely P. minor and wild oat (Avena ludoviciana Dur.). It was poor for control of broad-leaved weeds (Medicago denticulata Willd. and Rumex dentatus L.). However, pyroxasulfone in tank-mix combination with metsulfuron 4 g/ha, triasulfuron 20 g/ha, and pyroxsulam 18 g/ha effectively controlled (96.5%–99.8%) the diverse weed flora and improved the wheat grain yield (69.5%–285.9%) over untreated weedy control. Also, the pre-mix of pyroxasulfone + pendimethalin applied as PE was superior to either of these applied alone for weed control and grain yield. Pyroxasulfone 100–127.5 g/ha had yield gain of 119.6%–125.4% and 10.1%–26% over untreated control and pendimethalin 800–1250 g/ha, respectively. In pot studies, straw burnt ash drastically reduced the pyroxasulfone efficacy against P. minor and A. ludoviciana. Pyroxasulfone was also effective in pot studies for control of MHR P. minor having resistance against acetyl-coA carboxylase (ACCase), acetolactate synthase (ALS), and photosynthesis at the photosystem-II site-A (PS-II) inhibitor herbicides (clodinafop, sulfosulfuron, and isoproturon, respectively). The studies indicate that pyroxasulfone as PE or EPOE can be an alternative grass weed control herbicide in wheat in particular for the control of MHR P. minor.     

SELECTIVE ACTION OF TRIAZINES FOR CONTROL OF WILD CANARYGRASS IN WHEAT

Summary. The selective action of triazine herbicides for control of wild canarygrass ( Phalaris spp.) in wheat was studied in four field experiments. Canarygrass was controlled by pre- and post-emergence applications of methoprotryne and terbutryne. The phytotoxicity of methoprotryne to wheat was higher than that of terbutryne. Addition of small amounts of simazine to either one of the methylthiotriazines considerably increased damage to wheat. The most resistant stage of wheat after emergence was that of spike initiation, whereas the most sensitive stage was that of earing. Increase of yield as a result of control of canarygrass was higher in semi-dwarf wheat than in normal-height cultivars. Application of 625 g/ha of terbutryne to a semi-dwarf cultivar pre- and post-emergence increased grain yield by 56% and 29%, respectively.
Action selective des triazines pour la lutte contre les Phalaris dans le blé     

Interactions between barban and benzoylprop etbyl, flamprop metbyl or flamprop isopropyl applied at two growtb stages for wild oat control in cereals

Barban was combined with benzoylprop ethyl, flamprop methyl, or flamprop isopropyl, and the eflects of the combinations on wild oat control and crop yield were assessed, tn greenhouse experiments, the interaction in all three combinations was synergistic, following application at the 2- or 4-leaf stage of wild oats. Under field conditions, combinations of barban + benzoylprop ethyl applied at the 2- or 4-leaf stage of wild oats were as eflective for wild oat control and wheat yield response as barban at 028 kg/ha applied at the 2-leaf or benzoylprop ethyl at 14 kg/ha applied at the 4-leaf stage of wild oats. Mixtures of barban + flamprop methyl applied at the 2- or 4-leaf stage of wild oats provided excellent wild oat control and wheat yield increases. Mixtures of barban + flamprop isopropyl applied at the 2- or 4-leaf stage of wild oats provided comparable wild oat control and barley yield increases to barban (028 kg/ha) applied at the 2-leaf or flamprop isopropyl (14 kg/ha) applied at the 4-leaf stage of wild oats.     

The interaction between planting depth of four winter wheat cultivars, Alopecurus myosuroides Huds. and Bromus sterilis L. and their susceptibility to post-emergence applications of isoproturon and chlorotoluron

Seeds of four winter wheat cultivars, Slejpner, Galahad, Avalon and Penman, were sown at depths ranging from 6–75 mm in soil in pots, and isoproturon or chlorotoluron was then applied to the soil surface. For chlorotoluron-treated plants (both pre- and post-emergence) the dose required to produce a 50% effect (ED₅₀) was unaffected by depth of planting. In contrast, for isoproturon applied pre-emergence, the ED₅₀ for both Avalon and Slejpner was strongly affected by sowing depth. Although chlorotoluron was much more active in a second experiment when applied post-emergence to Slejpner wheat, the ED₅₀ for both herbicides increased with greater depth of sowing. Protection of wheat from isoproturon damage by deeper planting was enhanced if the adsorption capacity of the soil was raised from K_d 0.5 to 2.0 by incorporation of activated charcoal in the soil. Isoproturon entry into plants (as measured by the effect on rate of photosynthesis) was slower in those that had been sown deeper and were growing in more adsorptive soils, but there was no obvious relationship between these observations and isoproturon distribution in the soil profile. In nutrient culture the four wheat cultivars responded similarly to a range of doses of isoproturon. The chlorotoluron-sensitive cultivars, Slejpner and Galahad, were damaged by much lower doses of chlorotoluron than were Avalon and Penman. Bromus sterilis L. responded similarly to wheat with regard to its interaction with isoproturon and planting depth. Alopecurus myosuroides Huds., however, was less damaged by isoproturon when the zone above the seed was protected from the herbicide by growing the shoot through a plastic straw.     

Optimizing the performance of diclofop-methyl, cycloxydim, and clodinafop-propargyl on littleseed canarygrass (Phalaris minor) and wild oat (Avena ludoviciana) control with adjuvants

Optimizing the herbicide dose by the addition of adjuvants is an acceptable way to reduce the risk of side-effects from herbicides. Therefore, to detect a suitable adjuvant for diclofop-methyl, cycloxydim, and clodinafop-propargyl against littleseed canarygrass ( Phalaris minor ) and wild oat ( Avena ludoviciana ), six dose–response experiments were conducted. The treatments consisted of diclofop-methyl at 0, 112, 225, 450, 675, and 900 g ai ha⁻¹, cycloxydim at 0, 15, 30, 60, 90, and 120 g ai ha⁻¹, and clodinafop-propargyl at 0, 8, 16, 32, 48, and 64 g ai ha⁻¹ with and without the adjuvants of Frigate, olive oil, and castor oil at 0.2% (v/v) in order to control both littleseed canarygrass and wild oat. Tested herbicides performance was enhanced by all adjuvants against littleseed canarygrass and wild oat. The addition of Frigate and the vegetable oils had the lowest and the highest effect on the performance of all of the herbicides on both littleseed canarygrass and wild oat, respectively, which confirms the solubilizing nature of the cuticular waxes by vegetable oils. A comparison between the two vegetable oils revealed that olive oil exerted a greater control of littleseed canarygrass than did the castor oil. In contrast, castor oil exerted a greater control of wild oat than did the olive oil, which can be related to differences in the leaf surface micromorphology of the weeds.     

Chemical constituents of the essential oils of wild oat and crabgrass and their effects on the growth and allelochemical production of wheat

The interference of allelopathic weeds with crop plants might be mediated by volatile allelochemicals. In this study, the essential oil constituents of two weeds, wild oat (Avena fatua) and crabgrass (Digitaria sanguinalis), were investigated in relation to their effects on the growth and allelochemical production of wheat (Triticum aestivum). Subsequently, by means of gas chromatography and gas chromatography‐mass spectrometry, 52 compounds were identified from the crabgrass essential oil, particularly a signaling compound called methyl jasmonate, while 28 constituents were detected in the wild oat essential oil. Both essential oils were rich in terpenoids and inhibited the growth of wheat in air, filter paper and soil media but their inhibition varied with the growth medium and the weed species. In both the air and the filter paper media, there were not significant differences in the dry weight of wheat between the wild oat and the crabgrass essential oils. However, there was a greater reduction in the dry weight of the wheat root and plant with the wild oat essential oil than with the crabgrass essential oil in the soil medium. Furthermore, the production of the allelochemical, 2,4‐dihydroxy‐7‐methoxy‐1,4‐benzoxazin‐3‐one, in wheat was induced by the essential oils. The results suggest that allelopathic interference with wheat by wild oat and crabgrass affects not only the biomass allocation, but also the allelochemical production, of wheat.     

The effect on yield of mixtures and sequences of herbicides for the control of Alopecurus myosuroides Huds. and broad-leaved weeds in winter cereals

Mixed infestations of Alopecurus m yosuroides Huds. and broad-leaved weeds in winter cereals were controlled with a commercial formulation of ioxynil + bromoxynil + mecoprop. used in mixtures or as sequences with isoproturon or clofop-isobutyl. As well as controlling A. myosuroides the mixtures, applied in the autumn, gave good control of Veronica persica Poir., Galium aparine L. Lilhospermum arvense L. and Viola arvensis Murr. weeds which were resistant to isoproturon used alone. A. myosuroides had the major influence on crop yield in three experiments. High infestations of A. myosuroides reduced the broad-leaved weeds so that crop yields were only slightly improved when all the weeds were controlled compared with the control of A. myosuroides alone. In the remaining experiment. V. persica was very competitive with wheat; additional control of A. myosuroides improved yields considerably. Ioxynil + bromoxynil + mecoprop gave better final control of broad-leaved weeds when applied in the spring rather than the autumn but the sequence of a spring application following isoproturon or clofop-isobutyl in the autumn did not improve yields compared with the mixtures applied in the autumn. In all experiments, the highest yields were obtained from controlling both A. myosuroides and broad-leaved weeds in the autumn, provided that good control of A. myosuroides was obtained.     

Anwendung von Isoproturon, allein und in Kombination mit anderen Wirkstoffen, im Nachauflauf-Winter (NA-W) und Nachauflauf-Frühjahr (NA-F) zu Winterweizen und Wintergerste

Use of isoproturon, alone and in combination with other compounds, post-emergence in winter and spring on winter wheat and winter barley Isoproturon, alone and combined with dinoseb acetate and bifenox, was applied post-emergence in winter and spring to winter barley and winter wheat. The experiment was on two sites (sandy and clay soils) and lasted for two years. Although after winter application the initial herbicide level in the soil was higher than after spring treatment, there were no differences at the end of the growing season. The DT₅₀ (time to 50% disappearance) value for isoproturon was between 12 and 33 days in both years for both soils and the DT₉₀ value varied from 34 to 68 days. Soil residues of isoproturon were not affected by the presence of the other compounds. Winter application gave better weed control and higher crop yields.     

Physiological effects of methyl 2-[4(2,4-dichlorophenoxy)phenoxy] propanoate on oat,wild oat,and wheat

Methyl 2-[4-(2,4-dichlorophenoxy)phenoxy]propanoate (dichlofop-methyl) is a selective herbicide for wild oat (Avena fatua L.) control in wheat (Triticum aestivum L.). Dichlofop-methyl inhibited IAA-stimulated elongation of oat and wheat coleoptile segments by 51 and 13%, respectively, at 10 μM concentrations. Dichlofop-methyl alone had no auxin activity at concentrations of 0.1, 1.0, and 10 μM. The inhibitory effect of dichlofop-methyl was overcome partially by increasing the IAA concentration or by application of 3,6-dichloro-o-anisic acid (dicamba), a herbicide with weak auxin activity. The de-esterified free acid metabolite, 2-[4-(2,4-dichlorophenoxy)phenoxy]-propionic acid (dichlofop), at 10 μM inhibited auxin-stimulated oat coleoptile elongation by 23%, but it did not affect wheat coleoptile elongation at the same concentration. Both dichlofop-methyl and dichlofop inhibited root growth in excised shoots and seedlings of wild oat but had no effect on wheat. Dichlofop was a more effective inhibitor of root growth than dichlofop-methyl. The results suggest that dichlofop-methyl functions as a strong auxin antagonist, while the metabolite, dichlofop, inhibits root growth and development by another mechanism. The herbicidal effect of dichlofop-methyl may be the net effect of two biologically active forms of the compound each with a different mode of action acting at different sites within a susceptible plant.     

Influence of time of emergence of wild oat on competition with wheat

The influence of time of emergence of wild oat (Avena fatua L.) on its competition with wheat (Triticum aestivum L.) was studied, using boxes that allowed separation of root and shoot competition. The relative yield total for mixtures of wild oat and wheat, grown under different forms of competition and with different wild oat sowing times, was very close to unity, indicating that the two species competed fully for limiting resources. Wild oat was more competitive than wheat when the two species were sown simultaneously, due largely to its greater root competitive ability; the two species had similar shoot competitive abilities. When wild oat was sown 3 or 6 weeks later than wheat, wheat was more competitive than wild oat and the production of wild oat panicles was prevented. This was mainly the result of greater root competitive ability of the wheat, although shoot competitive ability was also greater. The effects of root and shoot competition were additive. It is concluded that in order to prevent the return of wild oat seeds to the soil, and hence obtain long-term benefit, it is necessary to control the wild oat seedlings emerging within the first 3 weeks after drilling a wheat crop.     

河北省冬小麦田杂草群落特征

为了明确河北省冬小麦田杂草的群落组成和结构,采用倒置"W"取样法对河北省7个地区146块冬小麦田的杂草进行了调查和物种多样性测度。结果显示,河北省冬小麦田有61种杂草,隶属于21科53属,播娘蒿、打碗花、荠菜、麦瓶草和麦家公是河北省冬小麦田的优势杂草。保定地区麦田杂草群落的物种丰富度最大;沧州地区麦田杂草群落的Shannon-Wiener指数和Pielou指数最大;廊坊地区麦田杂草群落的Gleason指数、Shannon-Wiener指数和Pielou指数最小,优势草种比较突出。经聚类分析和主成分分析,河北省冬小麦田杂草群落分为3组:石家庄、保定、邢台和廊坊的麦田杂草群落中耐旱杂草的相对多度较高;沧州和衡水的麦田杂草群落中耐盐杂草的相对多度较高;邯郸的麦田杂草群落中喜湿杂草的相对多度较高。说明土壤的盐度和湿度是影响河北省冬小麦田杂草群落组成的主要生态因子。     

Differential inhibition of potassium ion absorption by difenzoquat in wild oat and cereals

Over a concentration range of 5.0 × 10^?6?7.5 × 10^?4M, the selective herbicide difenzoquat (1,2-dimethyl-3,5-diphenyl-1H-pyrazolium) caused more pronounced inhibition of potassium ion (K⁺) absorption by excised seedling roots of susceptible wild oat (Avena fatua L.) compared to those of tolerant barley (Hordeum vulgare L. cv. Bonanza) or wheat (Triticum aestivum L. cv. Neepawa). At 2.5 × 10^?5M difenzoquat, the relative inhibition of K⁺ (⁸⁶Rb) absorption by wild oat root segments inceased from 30% with a 10-min uptake period to 75% with an uptake period of 90 min, whereas no inhibition at all was evident for wheat root segments even after a 90-min exposure to the herbicide. An ion efflux compartmental analysis procedure demonstrated that difenzoquat did not affect the passive permeability properties of the plasma membrane of wild oat root cells. The experimental findings indicated that difenzoquat interfered directly with the process of active ion transport across the plasma membrane of root cells.     

Weed Control in Wheat and Barley in the Middle East

Abstract

Wheat and barley constitute the bulk of agricultural production in the Middle East. Unfortunately, these crops are grown on marginal soils which are infested with annual and perennial weeds. The principal weed species include Avena sterilis L., Convolvulus arvensis L., and Sinapis arvensis L. and it has been estimated that yield losses due to weeds are between 20 and 30%. Control is mostly by hand pulling, hoeing or mechanical tillage, the use of herbicides being limited to about 2–5%, of the cereal area. The chemical used is usually 2,4-D but there are a few applications of barban. With the increase in wages, and unavailability of agricultural labour it is suggested that herbicide application should be widely used in the Middle East. Mixtures with benzonitriles and dicamba should be used in addition to the economical phenoxy herbicides to widen the spectrum of control of broadleaved weeds. Glyphosate may also be used after grain harvest for the control of perennial weeds and tri-allate, chlortoluron and other chemicals for the control of wild oats and other grasses. Plant breeders should study the genetic response of newly developed varieties to the most widely used herbicides.     

Differential effects of isoproturon on the photosynthetic apparatus and yield of two varieties of wheat and L. rigidum

The effect of isoproturon on the ultrastructure of the photosynthetic apparatus, ribulose bisphosphate carboxylase activity, protein and chlorophyll content, and the grain yield was investigated in two wheat cultivars (Triticum sativum L. cvs Castan and Esquilache) and a weed (Lolium rigidum Gaud.). Field experiments used applications of 1–65 and 2–5 kg a.i. ha^?1 isoproturon post-emergence, and growth chamber experiments used nutrient solution with the addition of isoproturon (1·7 × 10^?4 M). The ultrastructure of the photosynthetic apparatus of the cv. Esquilache was much affected by the herbicide. In the case of cv. Castan, slight disorganization of the grana and intergrana was observed. Isoproturon decreased the activity of ribulose bisphosphate carboxylase. A decrease in protein and chlorophyll content was also observed in the cv. Esquilache and in L. rigidum. These alterations were much less evident in the cv. Castan, where, moreover, no loss of protein occurred. The yield of the treated cv. Castan plants was slightly greater than that of the control plants in two consecutive years. However, the yields of the cv. Esquilache were significantly less when the herbicide was applied in the first year at commencement of tillering in a dry season but not when applied at an advanced stage as in the second year in a wet season.     

Survey of the presence of wild oat and blackgrass in parts of the United Kingdom

The survey was carried out over an area representing about 47% of the cereal acreage of the United Kingdom. Estimates of the acreage of fields containing wild oat (Avena fatua L. and A. ludoviciana Dur) ranged from 15% (Northern Ireland) to 74% (England, South West Region). The degree of infestation over the total acreage was 58%‘light’, 34%‘medium’, and 8%‘heavy’. Blackgrass (Alopecurus myosurodies Huds.) was found only in the central and southern regions, estimates of the infested cereal field acreage ranging from 2% (Wales) to 23% (South East England). Nine per cent of the fields were rogued for wild oat. On average, only 10% of the infested acreage received herbicide treatment for wild oat and blackgrass though, in some regions, up to 23 % was treated. The timing of treatment and the herbicide used were recorded. Information on the extent and type of herbicide used for broad-leaved weeds was also recorded. Data on sowing dates of the crops, the previous cropping, previous infestations, the soil type and the state of field drainage were obtained and were correlated with the incidence of wild oat and blackgrass where relevant.     

Littleseed canarygrass (Phalaris minor) resistance to clodinafop‐propargyl in wheat fields in north‐western India: Appraisal and management

Littleseed canarygrass (～canarygrass) evolved populations that are resistant to isoproturon during the early 1990s in north‐western India. Clodinafop‐propargyl (～clodinafop) was recommended for controlling these populations. It has been used extensively in wheat for the last several years. Recently, poor or no control of canarygrass by clodinafop has been observed in large areas, which could be related to cross‐resistance or multiple resistance. This study was designed to test whether resistance has evolved in canarygrass populations against clodinafop and to explore control of the resistant populations with sulfosulfuron and pinoxaden. Among the 311 canarygrass populations that were tested, 86, 55 and 34 showed variable phytotoxicity (0–99%) due to 0.030, 0.060 and 0.120 kg ha^?1 clodinafop, respectively. Based on the resistance index, 11 populations were “highly resistant”, 60 were “resistant” and the rest (240) were “susceptible” to clodinafop. Five and six clodinafop‐resistant populations showed slight resistance to 0.0125 kg ha^?1 sulfosulfuron and 0.025 kg ha^?1 pinoxaden, respectively. But, sulfosulfuron at 0.025 and 0.050 kg ha^?1 and pinoxaden at 0.050 and 0.100 kg ha^?1 controlled all the canarygrass populations. Clodinafop used for 4 years increased the chance of resistance evolving, whereas its rotation with sulfosulfuron reduced the chance of resistance evolving. This study showed that considerable canarygrass populations have evolved a low‐to‐high degree of resistance against clodinafop. The further use of clodinafop would lead to the spread of resistance in larger areas through the dispersal of resistant seeds. Clodinafop should be replaced with 0.025 kg ha^?1 sulfosulfuron or 0.050 kg ha^?1 pinoxaden. Besides, where canarygrass has not evolved resistance, the yearly rotation of sulfosulfuron with clodanafop or pinoxaden might delay the evolution of resistance.     

Competition between vegetative plants of wild oat (Avena fatua L.) and wheat (Triticum aestivum L.)

Competition between wild oat (Avena fatua L.) and wheat (Triticum aestivum L.) was studied in two experiments; a replacement series model and a technique for separation of root and shoot systems. Wild oat and wheat in association resulted in a relative yield total very close to unity showing that the two species were‘crowding for the same space’(or competing for the same resources) and were‘mutually exclusive'. Wild oat was more competitive than wheat, as shown by its aggressivity relative to wheat, relative yields, shoot dry weights and other plant attributes. The greater competitive ability of wild oat was predominantly due to its greater root competitive ability, while the two species had similar shoot competitive ability. Root competition had a much greater effect on the relative performance of the two species than did shoot competition. The effects of root and shoot competition were additive.     

A comparison of the dynamics of root growth and biomass partitioning in wild oat (Avena fatua L.) and spring wheat

The dynamics of early root growth and dry matter partitioning were compared in spring wheat (Triticum aestivum L.) and wild oat (Avena fatua L.) grown in solution culture. Total root length was greater in wheat than wild oat throughout the experiment; a result of a greater number of seminal axes and greater production of lateral root length per axis. The final number of adventitious roots was greater in wheat than in wild oat, but their length was similar. Relative growth rates were also similar as was shoot:root dry weight ratio and rate of root respiration. However, wheat used the dry matter partitioned to its roots more efficiently, producing a greater specific root length (SRL, length per unit weight). Caution must be exercised when relating these results to plants growing and compet-ing in the field, but three general points are raised. First, the initial number of seminal axes can have a profound effect on the rate of early root development; second, the adventitious root system of wild oat is not inherently more vigorous than that of wheat; and third, future studies should compare SRL of wheat and wild oat in the field. If differences similar to those in the present study are found they may contribute to the greater competitive ability of wheat.     

Residues of Mecoprop in Post-emergence-Treated Wheat and Oat

The dissipation of mecoprop in wheat (Triticum aestivum L.) and oat (Avena sativa L.) was monitored over a growing season following post-emergence application of the dimethylamine salt of mecoprop to each crop at 1·1 kg ha^?1. Residues of mecoprop, as its methyl ester, were determined gas chromatographically using electrolytic conductivity detection. Initial residues in wheat (119 (±20) mg kg^?1) and oat (95·3 (± 10·0) mg kg^?1) on the day of application (four-leaf stage of wheat and four- to five-leaf stage of oat) decreased to 0·1 to 0·2 mg kg^?1, respectively, within six weeks. Residues were non-detectable in the mature seed of both crops. Recoveries of mecoprop were in the order of 90% from the green tissue and seed of both crops fortified at 0·05 mg kg^?1.