兴安落叶松天然林林木直径与树高分布的分层特征

[目的]通过分析大兴安岭呼中林区兴安落叶松天然林胸径和树高的一元、二元分布特征,揭示脆弱地带森林群落的林木大小分布特征。[方法]在大兴安岭呼中林场设置3种不同类型兴安落叶松林固定样地,利用整体和分层的方法,对样地林分胸径和树高的变化规律以及两者之间的关系进行研究。[结果]表明:(1)草类-落叶松林直径分布为单峰分布,其他2块样地为双峰分布; 3块样地林木直径分布均呈左偏。杜鹃-落叶松林树高分布为多峰分布,其他2块地均为单峰分布; 3块样地林木树高分布均呈右偏。(2) 3块样地上、下层林木的直径分布均为单峰分布,上层木直径分布均呈左偏,下层木直径分布草类-落叶松林右偏,杜鹃-落叶松林左偏,杜香-落叶松林为近似正态。杜鹃-落叶松林的树高上、下层分布为双峰,其他2块样地均为单峰,3块样地上层木树高分布均左偏,而下层木树高分布均右偏。(3)草类-落叶松林和杜鹃-落叶松林小径阶林木株数占绝大多数。3块样地林分中树高与胸径的比值在小径阶林木中明显大于大径级林木。草类-落叶松林和杜鹃-落叶松林上层木中小径阶林木株数所占比例稍大,杜香-落叶松林和杜鹃-落叶松林下层木中大径阶林木株数比例稍大,但3块样地上层和下层木树高与胸径的比值在大小径阶林木中均无明显差异。[结论]高密度的杜鹃-落叶松林平均树高要高于低密度的杜香-落叶松林和草类-落叶松林平均树高,而平均胸径无明显差异。3块样地林分中树高与胸径的比值小径阶林木中明显大于大径级林木,但其差异在上层和下层林分中均表现不明显。     

A phytosociological study onBetula platyphylla forests in Daxing’an Mountains of China

For the first time in this paper the forests ofBetula platyphylla in Daxing’an Mountains are subdivided by phytosociological methods. Three community types ofB. platyphylla forest have been differentiated, e.g.,Rhododendron dahuricum-Betula platyphylla community,Corylus heterophylla-Betula platyphylla community andArtemisia stolonifera-Betula platyphylla community. The distributed elevation, stand height, differential species, major composition species of tree layer, shrub layer and herb layer for each community were detail described Yang Yongfu, male, born in Sept., 1961; Senior engineer of Central Nursery of Xinlin Forestry Bureau, Daxinganling, P. R. China Responsible editor: Chai Ruihai     

大兴安岭不同区域落叶松相容性材积方程及异方差研究

[目的]对不同区域立木相容性材积方程以及不同异方差校正方法进行详细对比分析,建立相容性材积方程预估大兴安岭不同区域落叶松的立木材积。[方法]以大兴安岭3个不同区域的落叶松为研究对象,采用误差变量联立方程组的方法构建不同区域立木相容性材积方程。采用非线性额外平方和的方法(F检验)进行区域性检验。使用多种权函数分别对3个区域存在异方差的材积方程进行加权回归。[结果]表明:任何2个区域的立木材积方程都有显著不同(P0.000 1),区域1和区域3的材积相差较大,区域2与区域1和区域3的材积相差较小。不同区域立木材积方程的错误应用会导致较大的预测误差。在参数稳定性和评价指标方面,加权估计会优于普通最小二乘估计。基于平均相对误差(MRE)和总相对误差(TRE),区域1(-0.11、0.97)、区域2(0.04、0.08)和区域3(1.04、0.93)的最优权函数分别为1/F(x)、1/D4.99、1/D3.38。[结论]立木材积方程是森林调查和林分生长与收获模型的主要组成部分,本文所构建3个区域的相容联立方程组模型预测误差均不超过±3%。建立相容性立木材积方程时应考虑其异方差的影响。最优权函数没有统一的形式。为尽可能得到稳定的参数估计,在加权回归估计过程中应选用多种权函数进行对比分析。     

Effects of differences in forest floor and canopy vegetation on ectomycorrhizas ofBetula platyphylla var.japonica: A test using seedlings planted into soils taken from various sites

Effects of different forest floor vegetation types in secondary forest and of conversion to plantation on the quality and quantity of ectomycorrhizas are mostly unknown.Betula platyphylla var.japonica seedlings were used for bioassays of ectomycorrhizal fungal inoculum using soils from four 50-year-oldB. platyphylla var.japonica forests that had different types of forest floor vegetation: two with shrub types, one with aSasa type, and one with a grass type. Seedlings were also grown in soil from a nearby monospecific plantation ofChamaecyparis obtusa. Ectomycorrhizas formed 13 to 26% of root length of seedlings grown in soil from the five different sites. The maximum percentage of ectomycorrhizal formation was obtained from the grass-type forest. The dominant type of ectomycorrhiza in the two shrub-type forest soils was the same as that in theSasa-type forest soil. The dominant types of ectomycorrhizas in the grass-type forest soil and in theC. obtusa plantation soil were different from that in the two shrub-type forest soils and in theSasa-type forest soil. The results of this investigation suggest that the type of forest floor vegetation, accompanied with changes in thickness of the A₀ horizon, might affect the ectomycorrhizal fungi in the soils ofB. platyphylla var.japonica forests. Establishment of artificial plantations ofC. obtusa might change the ectomycorrhizal fungi that could associate withB. platyphylla var.japonica seedlings in soil.     

针叶林标准化地位指数在大兴安岭林区立地质量评价中的应用(英文)

本文采用两点法原理和方法，对大兴安岭地区的落叶松和樟子松人工林地位指数表进行分析，建立了大兴安岭人工针叶林标准化生长模型，按模型导出各生长指数级和各龄阶理论值，并建立了标准化地位指数表。理论值检验结果表明，该标准化地位指数表符合精度要求。本项研究解决了大兴安岭地区针叶林同一立地类型不同树种地位指数和不同树种不同立地类型地位指数间的相互转换。     

大兴安岭主要森林类型林分空间结构及最优树种组成

以2011—2012年大兴安岭的44块固定样地调查数据为基础,以常用的林分空间结构参数(角尺度、大小比、混交度、林分空间结构指数和林分空间结构距离)为切入点,对大兴安岭主要森林类型的林分空间结构特征及最优树种组成进行研究。结果表明:天然落叶松林、针叶混交林以及针阔混交林的林木水平分布格局均表现为随机分布,而白桦林则为明显的聚集分布;4种林型中树木的生长整体处于中庸状态,落叶松处于明显优势地位,其他伴生树种则处于不同程度受压状态;落叶松混交程度随着其优势程度的增加而降低,其他伴生树种则具有较高的混交度,整体上天然落叶松林、白桦林的树种空间隔离程度较低,而针叶混交林、针阔混交林的林分平均混交度相对较高;大兴安岭4种主要森林类型林分空间结构的优劣表现为针阔混交林针叶混交林天然落叶松林天然白桦林;天然落叶松林最优的树种组成为8落,其次为7落。     

Relationship between resistance to vole browsing and content of ether extract in the bark of larch species and hybrids

We determined the survival rate of tree families produced by the intra- and interspecific crossing of larch species damaged by browsing of vole (Clethrionomys rufocanus bedfordiae Thomas) and the contents of ether extracts (EE) in their bark. The average survival ofLarix gmelini var.japonica Pilg. (L. gmelini)×L. gmelini (G×G) was highest, followed by the hybridL. gmelini×L. leptolepis Gordon (G×L). The families of the hybrid G×L, which is considered suitable for silviculture in Hokkaido, Japan, showed significant differences in survival rates, suggesting that it is possible to produce improved varieties of the hybrid which are more resistant to vole browsing. The EE contents of the bark were under strong hereditary control, and the rank of the EE content among the various families hardly fluctuated between the location of the test fields. The correlation coefficient between the survival rates and the EE contents was highly significant (r=0.89,n=24,p<0.01), indicating that the EE content in the bark may be used as a possible index of resistance or susceptibility to vole browsing in larch plantations. A part of this report was presented at the 106th Annual Meeting of the Japanese Forestry Society, at Sapporo, April, 1995.     

Characteristics of reproductive biology forLarix originating in Japan

The paper summarized the life cycle, environmental features and distributions ofLarix origination in Japan. The time of blooming and fruiting and the ways of cone collection and seeds storage were introduced. The treating methods of seeds germination and the sowing time for both Japanese larch (Larix kaempferi Carr) and Dahurian larch (Larix gmelinii var.japonica Pilger) were discussed. Responsible Editor: Zhu Hong     

Population characteristics ofDendrolims superans in Daxing’an Mountains

Dendrolims superans produced one generation every two years. Large larvae and small larvae exist at the same time. It broke out in Daxiang an Mountains in 1990, and seriously destroyed growth ofLarix gmelini Rupr. In order to controlD superans, the population properties ofD superans were studied from 1991 to 1992, including sexratio, age distribution, pattern etc.. The sex ratio ofD. superans population is ♀∶♂ − 1.06:L. The larvae age distribution indicates that larvae over 5 instar is less than younger larvae under 4 instar in Yongqing forest farm, but the younger larvae under 4 instar is more than larvae over 5 instar in Hanjiayuanzi Forest Farm. The population ofD. superans in Yongqing Farm has been declining. and mereasing in Hanjiayuanzi. Pupae and eggs are mainly distributed in middle and lower crowns layer of trees respeetively account for 66% and 59.5% of total individuals. The pupae in higher crown layer is lightly regular distribution and clumping distribution middle and lower crowns layer of trees. The eggs in higher crown is light clumping distribution and random in middle and low crown of tree. (Responsible Editor: Zhu Hong)     

The pulping and papermaking characteristics of youngL.gmelini andB.platyphylla

5, 10, 15, and 20 year oldLarix gmelini andBetula platyphylla were used as pulping raw material in the research. The examination of youngL. gmelini andB. Platyphylla fibrous structures showed that the fiber length and ratio of length to width of 5 years oldL. gmelini noticeably differed from that of matureL. gmelini. but that of 5 year oldB. platyphylla were nearby equal to that of matureB. platyphylla. The research results indicated that 15 years oldB. platyphylla and 20 year oldL. gmelini were best suited to pulping among the eight experimental raw materials; 10 years oldB. platyphylla and 15 year oldL. gmelini were optimal to the ratio of beating degree increment and corresponding beating time. The pulp characteristics of 15 year old B. platyphylla and 20 year oldL. gmelini were similar to those of mature wood, and it was feasible to consider 15 years toB. platyphylla and 20 years toL. gmelini as their rotation period; 10 years and 15 years as their thinning cycles for the papermaking forest base.     

兴安落叶松老龄林分幼苗天然更新及微生境特点

对落叶松老龄林幼苗天然更新及微生境研究采用样线法，沿样线设置样方，调查样方内幼苗数、测定环境变量。结果表明，原始老龄林各环境变量中地表层的作用最突出，光照的作用最小，灌木层居中。一般，枯落物、根系盘结层和苔藓的作用较大，其中枯落物对苔藓的发育有抑制作用。抑制兴安落叶松更新的主要因素：在丛桦落叶松林中主要是草本植物、赤杨落叶松林为苔藓厚度、杜香落叶松林为苔藓厚度和根系盘结层厚度。     

Rainfall redistribution of a virgin Pinus koraiensis forest and secondary Betula platyphylla forest in Northeast China

A virgin Pinus koraiensis forest in the Xiaoxing’an Mountains was selected to study its rainfall redistribution effect via 97 rainfall occurrences during a growing season. The following results were obtained: 1) The canopy interception of the P. koraiensis virgin forest amounted to 98168 mm during a growing season (May to September), which was 19.6 per cent of the total rainfall and 1.3 times that of a secondary Betula platyphylla forest. Compared with other forest types in China (11.4%–36.5%), the ratio of the canopy interception in the virgin pine forest was at a medium level. 2) The throughfall of the virgin pine forest was 395.77 mm, which accounted for 78.7% of total precipitation, and the stem-flow was 8.78 mm, accounting for 1.74% of total precipitation. Compared with the secondary birch forest, the virgin pine forest had lower throughfall but higher stem-flow. 3) Cubic regression equations (p < 0.01) which describe the relation between throughfall, stem-flow and canopy interception in the virgin pine forest and rainfall in an open field were fitted. A linear regression equation (p < 0.01) was found to be a better fit for the relationship between throughfall of the secondary birch forest and rainfall outside the forest. Factors affecting throughfall and stem-flow were analyzed, with results providing a good reference to the study of rainfall redistribution in coniferous and broadleaved mixed forests. __________ Translated from Science of Soil and Water Conservation, 2006, 4(6): 61–65 [译自: 中国水土保持科学]     

Growth variation and early selection ofBetula platyphylla in natural stands

The stem analysis ofBetula platyphylla in natural forest at the Maoershan Forest Experimental Station of Northeast Forestry University, showed that the growth variation of the trees, including height and diameter breast height, decreases with the increase of the age. There is the turning point for the growth variation of the tree from acute change to relative stability when the trees are in eighteen years old. There are significant differences among the height, diameter and volume growth of the trees at that time. Therefore, the optimal age for early selection of this species in natural stands is eighteen years old (18 a). Diameter is used as main index for selection and height growth as auxiliary one. (Responsible Editor: Zhu Hong)     

不同干扰类型下的白桦次生林土壤理化性质研究

研究内蒙古大兴安岭地区不同干扰类型下白桦次生林土壤理化性质特征,在内蒙古根河市选取3种干扰类型的白桦次生林样地的不同土层土壤作为研究对象,对土壤理化性质进行测量分析,探究不同干扰类型对土壤理化性质的影响。研究结果表明,白桦次生林在不同干扰方式下土壤理化性质存在显著差异。其中人工条带改造后的白桦次生林土壤pH值增大,火烧后白桦次生林容重增大,土壤有机碳、全氮、全磷、全钾、速效磷、速效钾含量在人工条带改造后显著增高,采伐干扰样地白桦次生林样地土壤含水量、速效氮含量最高。不同干扰类型的白桦次生林样地土壤理化性质间相关性不同。人工条带改造干扰样地白桦次生林土壤综合指数较高,火干扰样地白桦次生林土壤综合指数较低。     

长白落叶松纸浆林培育措施对经济效益的影响(英文)

根据长白落叶松纸浆林的培育实践,以经济指标内部收益率和净现值为基础,分析了培育措施对经济效益的影响。结果表明:轮伐期越短,经济效益越高,立地级越高,经济效益亦越高。造林密度对经济效益的影响也较明显。在保证造林成活率和保存率的基础上,用于整地和幼林抚育方面的经费越少,经济效益则越高。选定的6项措施对财务指标的影响顺序为:轮伐期→立地条件→造林密度→管理费水平→幼林抚育强度→整地方式     

Population genetic diversity and regional differentiation of Chinese forest frogs (Rana chensinensis) in Heilongjiang Province

The Random Amplified Polymorphism DNA (RAPD) markers were used to study the intra-specific diversity and regional differentiation of the Chinese forest frogs (Rana chensinensis), which were sampled from the fields of 8 regions in, Heilongjiang Province. Totally 78 polymorphic DNA loci were amplified by 10 RAPD primers. By genetic distance analysis and phylogenetic tree reconstruction with the Neighbor-Joining (NJ) method the results showed that the populations of Chinese forest frogs distributed in 8 regions existed great differentiation (Average Fst=0.347, SD=0.235) while, there was the paradox between geographic distances and genetic distances. Based on geographic and geological data, a hypothesis was posed that it is very possible that the hilly lands in the downstream of the Songhua River and the Heilong River were the center of the origin of the Chinese forest frogs in Heilongjiang Province. And mainly through the Songhua River system, the Chinese forest frogs dispersed into the Songneng Plain from the Sanjiang Plain whereas the Fangzheng region became a sub-center for the western dispersion. Foundation item: This project is supported by the Natural Science Foundation of Heilongjiang Province (C9726). Biography: XIAO Xiang-hong (1960-), female, professor in College of Wildlife Resources, Northeast Forestry University, Harbin 150040, P.R. China. Responsible editor: Zhu Hong     

Control of Lymantria dispar L. by biological agents

The experiment on control ofLymantria dispar L by using different kinds of biological measures, including nuclear polyhedrosis virus (NPV) ofLymantria dispar L., BtMP-342, sex-attractant as well as botanical insecticide, was carried out in the forest regions of Inner Mongolia in 2003. Two concentrations (2.632×10⁶ PIB·ml⁻¹ and 2.632×10⁷ PIB·ml⁻¹) ofLymantria dispar L. NPV were sprayed on the 2rd-instar-larvae ofL. dispar and 70% and 77.8% control effect were obtained respectively. BtMP0-342 was applied to the 3rd- and 4th-instar larvae and the control effect was around 80%. The sex-attractant provided by Canada Pacific Forestry Research Center also showed a good result in trappingL. dispar adults. The self-produced botanical insecticide, which was extracted from a kind of poisonous plant distributed in Daxing'an Mountains, China, exhibited a good control result in controlling the larvae ofL. dispar, and 82% mortality was observed when spraying primary liquid of the botanical insecticide on the 3rd–5th-instar-larvae in lab. Foundation item: This paper was supported by Natural Science Foundation of Heilongjiang Province (C0235) and Youth Foundation of Harbin City (0061218024). Biography: ZHANG Guo-cai (1964), male, Ph. Doctor Candidate, associate professor in School of Forestry, Northeast Forestry University, Harbin 150040, P. R. China. Responsible editor: Song Funan     

大老岭自然保护区日本落叶松林凋落物分解及养分释放研究

[目的]研究外来引进树种日本落叶松林凋落物对土壤养分的影响。[方法]采用分解袋法分别对18年生和24年生日本落叶松林以及周围针阔混交林凋落物的分解和养分释放规律进行了研究。[结果]凋落物分解和养分释放速率均表现为针阔混交林日本落叶松纯林;24年生日本落叶松林18年生日本落叶松林。其中不同林分的凋落物残留率与时间呈指数相关,凋落物年分解系数(K)也表现为针阔混交林(0.555 6)24年生日本落叶松林(0.445 0)18年生日本落叶松林(0.366 2)。凋落物分解速率与初始N元素含量呈极显著正相关,而与C/N比呈显著负相关,高的木质素含量对凋落物的分解有一定影响。C元素、K元素表现为直接释放模式,而研究中C/N比和C/P比相对较高,使N元素和P元素均表现为先富集后释放的模式。各养分元素的残留率总体呈现出18年生日本落叶松林24年生日本落叶松林针阔混交林的格局。[结论]不同林分凋落物分解和养分释放速率差异较大。凋落物年分解系数表现为针阔混交林24年生日本落叶松林18年生日本落叶松林。     

华北落叶松液流速率的优势度差异及其对林分蒸腾估计的影响

将实测的杨树液流速率和林木边材厚度(或胸径)经尺度扩展得到林分蒸腾,已成为常用的野外测定方法,但此法没有考虑其他树形因子的影响,当林分密度大、光竞争激烈时,会导致蒸腾估计误差偏大。为认识主要树形因子对液流速率的影响,并为改进样树液流向林分蒸腾的尺度扩展方法提供依据,在六盘山北侧半干旱区华北落叶松人工林内,利用热扩散探针对5株不同优势度样树的液流速率进行了连续观测,研究了生长季中期(叶面积指数达到峰值并保持稳定)不同土壤水分条件下不同优势度树木的液流速率差异。结果表明:优势度越大的树木,其液流在日内的启动越早,结束越晚,到达峰值越早,峰值也越大;日均液流速率明显比优势度小的树木高;液流速率对太阳辐射和饱和水汽压差瞬时变化的响应敏感性比优势度小的树木高,而对土壤水分条件的响应敏感性则弱于后者,但整体上对环境条件的响应趋势一致,不同优势度树木间液流速率的相对差异比较稳定。相关分析表明:液流速率与优势度(或相对树高)、树高呈极显著正相关,与冠长、胸径显著正相关,而与冠幅、边材面积正相关但不显著。利用拟合的优势度与液流速率之间的线性关系(R²=0.95)计算了样地内所有树木的液流速率及其平均值,即林分平均液流速率,该值比常用方法计算结果低16%。建议今后在利用样树液流速率测定结果进行尺度扩展计算林分液流速率和蒸腾时,增加考虑优势度等主要树形因子的影响。     

兴安落叶松天然林不同分化等级林木树干液流对综合环境因子的响应

[目的]以大兴安岭北部典型寒温带针叶林优势建群树种兴安落叶松为对象,分析不同分化等级林木树干液流对环境因子的综合响应,构建不同分化等级林木树干液流模型。[方法]利用热扩散式液流监测系统和通量塔的梯度气象系统连续监测树干液流及环境因子的变化。[结果]表明:1)观测期间,优势木具有较强的蒸腾能力,其平均液流密度分别为中等木和被压木的1.9倍、2.5倍。总体上,分化程度越高的林木日树干液流持续时间越长,液流密度峰值出现时间越早,液流密度的峰值也越高。2)利用主成分分析将降雨、净辐射、空气温度、空气湿度、风速、土壤温度、土壤含水量和水汽压亏缺降维为蒸发需求因子(EDI)、土壤水热因子和降水因子。EDI(与净辐射、温湿度、水汽压亏缺显著相关)是影响该地区林木树干液流的关键环境要素,其携带环境数据信息量的45%;土壤水热因子和降水因子分别携带20%和13%。3)各分化等级林木树干液流密度对EDI呈顺时针时滞,对净辐射和水汽压亏缺则分别呈逆时针、顺时针时滞,且EDI的时滞效应明显较小。不同分化等级林木液流密度对EDI和水汽压亏缺的时滞表现一致,对净辐射的时滞则以优势木最小。4)各分化等级林木树干液流密度对EDI的响应均符合"S"型模型,即液流升高到阈值后,不再随蒸发需求的增加而增大。模型中,中等木(0.458)和被压木(0.457)的过渡斜率略高于优势木(0.443),表明优势木树干液流对环境因子的敏感性略低。该模型对不同分化等级林木液流密度的模拟精度均在90%以上,考虑EDI的时滞效应或引入土壤水热因子、降水因子对模型精度的影响较小。[结论]兴安落叶松树干液流对综合环境因子存在较强的响应性,且在不同分化等级间存在差异;利用"S"模型和综合环境因子可有效估算不同分化等级兴安落叶松的树干液流。