A research approach supporting domestication of Baobab (<Emphasis Type="Italic">Adansonia digitata</Emphasis> L.) in West Africa

The Baobab tree (Adansonia digitata L.) is a key multipurpose species for the African region. In the recent years there has been an extended commercial interest for different A. digitata products. As a spectacular African key species there has been a growing interest from NGO′s and various research groups. A research group, focussing on the following countries Burkina Faso, Mali and Niger, has participated in a concerted research action with cross counter disciplinary cooperation between plant physiology, population genetics, tree breeding, food science, and socioeconomics. This paper presents a review and the way knowledge gaps are being addressed using the above mentioned approach. The overall work was initiated in 2005–2006 when a large collection of A. digitata seeds was carried out in 15 African countries. Fourteen populations were selected in Burkina Faso, Mali, and Niger including more than 400 superior trees, and many of the trees have been selected along with farmers in a participatory process. Studies have been initiated in all countries on geographic variation of growth, adaptive and phenological traits. This includes studies in the nursery, establishment of international provenance series, seed orchards and studies of gene flow and phylogeographic variation with various markers. Specific nursery trials have been established in order to study provenance and progeny performance related to drought stress. Several stress related characteristics are being measured. Preliminary observations showed large morphological variation between African provenances of A. digitata. Food properties of plant parts have been studied within and between populations. This includes studies of total biomass production, vitamin A, B1, B2, and various minerals. The influence of stress will be related to these parameters. Fruits, which are rich in sugar and vitamin C, are used as an ingredient in juice and other foods. Seeds can be used directly as food ingredient or in fermented condition (Maari). The fermentation process was previously poorly described. Therefore, the microorganisms associated with Baobab seeds fermentation have been identified and a starter cultures for control production has been proposed. The efficiency of dissemination of superior A. digitata trees depends on vegetative propagation. Various grafting methods are currently being tested in cooperation with farmers. Micropropagation is tested as well as a method for clone propagation. The accumulated knowledge will be applied for a domestication strategy of A. digitata in Burkina Faso, Mali and Niger. The outcome from this study will be guidelines for protection and management of the germplasm of A. digitata resources.     

Early growth performance and water use of planted West African provenances of Vitellaria paradoxa C.F. Gaertn (karité) in Gonsé, Burkina Faso

A participatory selection trial of five provenances from Burkina Faso (3), Mali (1) and Senegal (1) was established in Burkina Faso in 1997 with the aim of addressing issues of the long juvenile phase and the large variability in annual fruit yields of Vitellaria paradoxa C.F. Gaertn in West Africa. The objectives of the present study were to evaluate survival rate and the growth performance of the five provenances, characterize the wetting profile under which the trees of these provenances are growing and quantify the variation in their seasonal transpiration. The design was a randomized complete block design (RCBD) with single tree as the experimental unit which was replicated 70 times. The results showed a mean survival rate of 50% for all provenances. Passoré (Burkina Faso), Djonon-Karaba (Mali) and Botou-Fada (Burkina Faso) provenances showed the highest height and collar diameter whereas the provenance of Gonsé (Burkina Faso) performed poorly. Djonon-Karaba provenance displayed the highest water use (2.70 l day⁻¹ tree⁻¹ in 2004 and 2.85 l day⁻¹ tree⁻¹ in 2005). Soil water content under Passoré provenance was the lowest (9.38%) whereas its content under Gonsé provenance (11%) was the highest with no clear pattern according to the distance from tree trunk. Samecouta and Djonon-Karaba provenances showed the highest transpiration values per sapwood unit area in 2004 (0.079 l cm⁻² day⁻¹) and in 2005 (0.069 l m⁻² day⁻¹), respectively. Based on growth performance and water use, it can be recommended at this early stage the selection of Djonon-Karaba and Passoré provenances as the most suitable for semi-arid regions of West Africa. However, there is a need for further data to model the long term effects of these provenances on soil water balance and their fruit production before reliable recommendations can be made to farmers.     

Evidence for important genetic differentiation between provenances of Parkia biglobosa from the Sudano-Sahelian zone of West Africa

Parkia biglobosa is a traditional economic tree legume of considerable multipurpose importance in the sudano-sahelian region in Africa. The species grows in multiple climatic zones with precipitation ranging from 600 to 2,500?mm a year and its natural distribution extends from Senegal and Guinea in West Africa to Uganda in Central Africa. In the present paper, a range wide sample of 25 provenances of P. biglobosa was tested in Burkina Faso, West Africa at two sites; Gonse (latitude 12°25′N; longitude 1°20′W; altitude 280?m) in the north-sudanian zone and at Dinderesso (latitude 11°18′N; longitude 4°35′W; altitude 425?m) in the south-sudanian zone. Based on analysis of survival and growth traits, we provide evidence of substantial genetic differentiation between P. biglobosa populations within West Africa. Height growth was best at Gonse, while the survival rate was higher at Dinderesso (61%) compared to Gonse (35%). Links between geographical parameters and the provenances performance were significant, and interesting geographic patterns were observed. Our results point towards superior fitness of the local Burkina Faso populations, and we speculate that presence of a continuum of locally adapted populations can be a part of the explanation for the species’ ability to thrive under quite different climatic conditions across West Africa. Based on the findings, we suggest recommendations for seed deployment and conservation strategies of the species in the West African Sahel. This is particularly important when considering the on-going climate change.     

Sesbania seed collections in Southern Africa

The International Centre for Research in Agroforestry (ICRAF) is a member of the Consultative Group on International Agricultural Research (CGIAR) system and has recently established a Multipurpose Tree Germplasm Resource Unit (MPT-GRU) in Nairobi, Kenya. The first major activity of the MPT-GRU was to undertake a regional germplasm collection ofSesbania in 5 countries in southern Africa in a collaborative mode with National Agricultural Research Services (NARS). The collections took place in Botswana, Malawi, Namibia, Zambia and Zimbabwe.Key elements of a generalised collaborative mode (the model) are indicated follwed by a summary of actual events. The initial phase involved joint planning and training sessions between NARS, three sister CGIAR centres and one regional institution, the Southern Africa Development Conference (SADC) Regional Gene Bank (SRGB). This was followed by actual collections during 1993 led by National scientists with different levels of CGIAR participation in each of the five countries. The International Board of Plant Genetic Resources (IBPGR) took the lead in pre-collection training activities, ICRAF for developing the collection programme and the International Livestock Centre for Africa (ILCA) for seed drying, testing and despatch for field trials. NARS gene banks guided the necessary administrative processes. The ICRAF lead scientist for the species and provenance evaluation phase was a crucial member of the planning process thus allowing integration of the collection with the eventual field evaluation. The article focuses on the collection strategy, administrative protocols for actual collections and summary results. Some advantages and difficulties associated with the collection programme and model are discussed. The collections were important in bringing together agriculturalists and foresters in a common germplasm collection exercise.     

ICRAF's multipurpose tree and shrub information system

Woody perennials are vital components of agroforestry systems. However, the state of knowledge about most of the large number of useful species is inferior to that about other components in such systems. In order to close this gap and expedite research, ICRAF has built up a multipurpose tree and shrub information system. The recommendations of an international workshop on multipurpose tree germplasm, organized by ICRAF in collaboration with CFI and IBPGR, have been instrumental in either encouraging the pursuance of activities already started or initiating new projects. In its endeavour to serve those institutions and individual scientists who require information on multipurpose trees, ICRAF has established a Multipurpose Tree and Shrub Data Base and compiled and published a Multipurpose Tree and Shrub Seed Directory. As by-products of these two core activities, a number of related projects have been carried out, including the development of a rapid appraisal methodology for selecting priority species for further development, the preparation of multipurpose tree and shrub crop sheets, and an inventory of MPT-related data bases. In a considerable number of publications the concepts and objectives of these activities have been explained. More than 140 information requests in multipurpose trees have been answered with the help of the Multipurpose Tree and Shrub Data Base and the information gap concerning germplasm availability has been closed by the publication of the Multipurpose Tree and Shrub Seed Directory.Senior Research Scientist, ICRAF     

Growth and survival of Prosopis africana provenances tested in Niger and related to rainfall gradients in the West African Sahel

Prosopis africana (Guill., Perrott. and Rich.) Taub. is important for farming and pastoralist communities in the West African Sahel. It provides a range of essential products (wood, fuel, food, fodder and medicines) and services (soil fertility improvement), and is one of many native tree species maintained in the traditional agroforesty system, known as the parkland. However, this and other valuable species are disappearing in many parts of the region due to over-exploitation and, perhaps, due to increasingly drier conditions in the region. A provenance/progeny test (275 families from 28 provenances in Burkina Faso and Niger) was established at one location in Niger to investigate geographic variation in the species. Mean annual rainfall decreases from south to north and from west to east in the seed collection area. Given these gradients, one would expect clinal variation in growth and survival of P. africana and other tree species that occur naturally in the region. Tree growth (height, number of primary branches, stem diameter at 20 cm and 1.3 m) and survival were analyzed at 11 years. Variation due to provenances was significant for growth variables and almost significant for survival. Variation due to families within provenances was not significant for any variable. Provenance means for growth and survival increased from the more humid to the drier parts of the sample region. The clines in growth and survival with regional rainfall patterns suggest that tree improvement and conservation programs should collect seeds from populations in the drier zones for planting and conservation in the West African Sahel. This may be particularly important when one considers the trend in global warming.     

Ethnicity, land use and woody vegetation: a case study from south-western Burkina Faso

Woody vegetation in cultivated landscapes in Burkina Faso is influenced by agricultural activities that are in turn influenced by institutional arrangements. Research was undertaken in a village in south-west Burkina Faso to investigate the relationship between species composition, diversity, density, species accumulation and land use category. Additionally the relationship between number of trees, size of farmed land and farmer ethnicity was investigated. Indigenous Tiéfo farmers had on average more than double the number of large (>4 m) trees of Vitellaria paradoxa Gaertn. f., Parkia biglobosa (Jacq.) R. Br. Ex G. Don and Anacardium occidentale L. than farmers belonging to other ethnic groups, but this was partly explained by a larger area being available. Differences between ethnic groups were not significant when expressed per ha. Botanical inventories of fallows more than 4 years old, cultivated parklands and plantations of Mangifera indica L. and A. occidentale showed that tree density and Simpson’s index of diversity for trees were highest in the fallow. Simpson’s index was not significantly different for regeneration, but sample-based species accumulation curves indicated that species accumulation in parklands was faster than in plantations when expressed per individual. The average regeneration density was 12,605, 1,995 and 6,772 seedlings ha^-1 for fallow, parklands and plantations, respectively. This low density for parklands means that species accumulation is slow compared to the other land uses when expressed per unit area. Fallow seems the most efficient way of keeping tree diversity in the agricultural landscape.     

Woody species composition,diversity and structure of riparian forests of four watercourses types in Burkina Faso

Riparian forests are classified as endangered ecosystems in general,particularly in sahelian countries like Burkina Faso because of human-induced alterations and civil engineering works.The modification of this important habitat is continuing,with little attention being paid to the ecological or human consequences of these changes.The objective of this study is to describe the variation of woody species diversity and dynamic in riparian forests on different type of watercourse banks along phytogeographical gradient in Burkina Faso.All woody species were systematically measured in 90 sample plots with sides of 50 m × 20 m.Density,dominance,frequency and species and family importance values were computed to characterize the species composition.Different diver-sity indices were calculated to examine the heterogeneity of riparian forests.A total of 196 species representing 139 genera and 51 families were recorded in the overall riparian forests.The species richness of individuals with dbh ≥ 5cm increased significantly from the North to the South along the phytogeographical gradient and varied significantly between the different types of riparian forests.Similarity in tree species composition between riparian forests was low,which indicates high beta diversity and reflects differences in habitat conditions and topography.The structural characteristics varied significantly along the phyto-geographical gradient and between the different types of riparian forests.The diameter class distribution of trees in all riparian forests showed a reverse "J" shaped curve except riparian forest of stream indicating vegetation dominated by juvenile individuals.Considering the ecological importance of riparian forest,there is a need to delineate and classify them along watercourses throughout the country.     

Farmers' contributions to the conservation of tree diversity in the Groundnut Basin,Senegal

Tree diversity in West Africa is threatened by intensified land uses and salinization,and farmers' role in conservation of tree species is unclear.We hypothesized that farmers contribute to conservation of tree diversity through protection of trees in their agroforestry landscapes and compared the diversity and structure of the tree vegetation across landscape classes.Inventories were carried out in three villages in the Groundnut Basin in Senegal,assessing tree diversity,density and crown cover.Tree diversity as assessed by species accumulation curves was high in forests,but cultivated landscapes had comparable or almost comparable diversity,especially in the cases where the forest was planted or was affected by charcoal production.However,the occurrence of exotic species was higher in cultivated parts of the landscape,and although many species were in common,ordination plots indicated that forests and cultivated landscapes to some degree had different species composition.Salinity had a strong influence on vegetation,not only in the tans(salt marshes) but also across the other landscape classes.In conclusion,agroforestry landscapes in the three villages harbor considerable tree diversity,but insufficient to fully conserve the tree species.We argue that informing and including farmers in tree management in the region will contribute to overall conservation of tree genetic resources.     

Soil amelioration and root symbioses ofParkia biglobosa (Jacq.) Benth. in West Africa

Parkia biglobosa is an important multipurpose tree from the savanna zone of West Africa. It has been reported to increase soil fertility and crop yields beneath its crown. However, no work has been conducted to determine the role of root symbioses in soil amelioration by this species. The existing reports of nodulation inParkia biglobosa are contradictory and the presence of mycorrhizae is not documented in the literature.Fieldwork was conducted at six sites in Burkina Faso and Nigeria, to investigate the amelioratory effect ofParkia biglobosa on soil fertility and to ascertain the occurence of root symbioses. Soil samples were collected from points close to the trunk, the mid-crown, crown edge and the open areas away from individual trees. The samples were analysed chemically to assess pH, organic matter, nitrogen, phosphorus and potassium content. Root samples were also collected from the field and examined for nodules and for evidence of mycorrhizal infection.The results of the soil analysis showed a significant amelioration for total nitrogen and available potassium with proximity to the tree, but organic matter, available phosphorus and soil pH showed no significant trend. This pattern was similar for all sites. The results also indicated that amelioration of nitrogen and potassium with proximity to the tree increased with tree size.Root nodules were absent from the roots of trees examined in the field, both in Burkina Faso and Nigeria. A pot experiment was set up using the soil collected from beneath the trees and a laboratory grown cowpea miscellany culture as rhizobial inoculum forParkia biglobosa seedlings. The roots of these seedlings also showed no evidence of nodulation. Ectomycorrhizae were not present in roots collected from the field, but endomycorrhizal infection was prolific.A study of root distribution in Burkina Faso suggested that one mechanism for increased soil fertility beneathParkia biglobosa is the widespread lateral root system, which redistributes nutrients from an extensive radius to a concentrated area beneath the crown. The root system is aided in the efficient uptake of nutrients by endomycorrhizae. Other possible mechanisms include interception of dust by the crown and deposition by throughfall and stemflow, defacation by animals beneath the tree and decomposition of leaf litter.     

The state of the art of agroforestry diagnosis and design

Seven years ago the International Council for Research in Agroforestry set out to develop a methodology for the diagnosis of land management problems and design of agroforestry systems. Since then over 60 documents (articles, conference papers and manuals) relating to the D & D methodology have been published by ICRAF and the methodology has been used to develop agroforestry plans and identify research priorities for a wide range of ecological and cultural conditions throughout the tropical regions of Africa, Latin America and Asia. This article reviews the evolution of the D & D methodology and suggests directions for its future development.Ecological Anthropologist, ICRAF     

Variability of Baobab (Adansonia digitata L.) fruits’ physical characteristics and nutrient content in the West African Sahel

The present study was carried out to evaluate variability in fruit characteristics and nutritional quality of Baobab fruits with the aim of providing the background to select trees bearing fruit with desirable characteristics for further utilisation. Vitamin C, total sugar and ash contents were assessed in 178 Baobab fruit samples from 11 sites in Burkina Faso, Mali and Niger. Furthermore the following tree and fruit physical characteristics were recorded: tree height, bark colour, fruit size, pulp weight, seed weight, seed size and pulp colour. The content (mean?±?SD) of vitamin C was 4.78?±?1.02?g?kg^?1, sugar 514?±?72?g?kg^?1 and fruit weight 293?±?96?g. There was a significant correlation between annual precipitation of the tree population site and vitamin C content but not with sugar content. For sugar, there were significant positive correlations with latitude and longitude. Negative correlations were found between fruit size and both longitude and latitude with smaller fruits generally being found to the north/east. No relation was found between pulp or bark colour and the sugar or vitamin C content. The contents of protein, lipid, carbohydrates, ash and moisture in the seeds ranged from 156 to 159, 143 to 150, 641 to 652, 44 to 49 and 50 to 55.7?g?kg^?1 respectively. The variation for vitamin C and sugar found within populations is a first indication that valuable gains could be made by selection of superior trees.     

Mistletoes of the agroforestry parklands of Burkina Faso

In sahelian countries, parklands are very important for human populations from an economic point of view. Unfortunately, they are currently being threatened by pests such as Loranthaceae which are stem parasites. In Burkina Faso, six species of Loranthaceae have been identified and located geographically: Agelanthus dodoneifolius (DC.) Polh. & Wiens, Englerina leucardii (Engl.) Balle, Globimetula cupulata (DC.) Van Tieghem, Tapinanthus bangwensis (Engl. & K. Krause) Danser, T. globiferus (A. Rich.) Van Tieghem and T. ophiodes (Sprague) Danser. Agelanthus dodoneifolius is by far the most widespread in the country whereas Globimetula cupulata is strictly located in the southeast. A list of their hosts has been established showing: 154 species dispatched in 96 genera and 42 families. Among them, the most frequently parasitized families are the Caesalpiniaceae, the Combretaceae and the Mimosaceae. A biological cycle scheme showing the main phenological stages common to the six Loranthaceae observed in Burkina Faso is reported. Based on a review of the damage caused by Loranthaceae, pruning, chemical treatments, genetic selection of resistant individuals and biological control are suggested as possible control methods.This revised version was published online in November 2005 with corrections to the Cover Date.     

Differentiation of Prosopis juliflora (Sw.) DC. and P. pallida (H. & B. ex. Willd.) H.B.K. using foliar characters and ploidy

Two species of Prosopis, Prosopis juliflora and Prosopis pallida, native to Central America and northern South America, have been widely introduced and are now found throughout the world’s hot arid and semi-arid zones. However, these species are frequently misidentified, especially where introduced, hampering genetic improvement and effective management. Samples from trees previously identified as either P. juliflora or P. pallida were divided into two groups by multivariate analysis using selected foliar characters. One group contained known tetraploids and the other contained known diploids. It was concluded that the tetraploid and diploid groups correspond to P. juliflora and P. pallida, respectively. Triploid individuals were also identified, the first report of these in the genus Prosopis. A dichotomous key was developed to differentiate the species, using number of interfoliar glands per rachis, number of leaflet pairs per pinna and leaflet length. P. juliflora and P. pallida grown from seed collected within their respective native ranges were correctly identified using the key. Where introduced, samples from Burkina Faso, Jordan and India were confirmed as P. juliflora, but those from Brazil, Cape Verde and Senegal were identified as P. pallida, rather than P. juliflora, the species assumed to be naturalized in these countries. The correct identification of these two species will facilitate the genetic improvement and management of P. juliflora and P. pallida. There is a strong case for basing future selection and genetic improvement programs on P. pallida rather than P. juliflora. Further collection and evaluation of P. pallida is therefore required, together with conservation programs in the native range.     

Human impact on population structure and fruit production of the socio-economically important tree Lannea microcarpa in Burkina Faso

Non-timber forest products (NTFPs) are of high socio-economic value for rural people in West Africa. Main factors determining the status of the populations of socio-economically important tree species providing those NTFPs are human activities. This study assesses the impact of human population density, land use, and NTFP-harvesting (pruning and debarking) on population structure and fruit production of the socio-economically important tree Lannea microcarpa that is normally conserved by farmers on fields. We compared L. microcarpa stands of protected sites with those of their surrounding communal sites in two differently populated areas in Burkina Faso. Our results reveal an opposed land use impact on the population structure of L. microcarpa in the two areas. In the highly populated area, the species population was more stable in the protected site than in the communal site, while the opposite was observed for the less populated area. Trees of the communal sites bore more fruits than trees of the protected sites. Debarking and pruning had a negative impact on fruit production of the species. We conclude that low intensity of human impact is beneficial for the species and that indirect human impact facilitates fruit production of L. microcarpa. In contrast, in the densely populated area, human impact has reached an intensity that negatively affects the populations of L. microcarpa. While the extent of protecting L. microcarpa on fields still seems to be enough to guarantee the persistence of this important species in the less populated area, it is no longer sufficient in the densely populated area.     

Genetic variation in wood density and correlations with tree growth in Prosopis africana from Burkina Faso and Niger

? Variation in wood basic density and its correlation with tree growth were investigated at 13 years in a provenance/progeny test of Prosopis africana in Niger. The test included progeny from 256 trees sampled from 24 provenances in the Sahelian ecozone of Burkina Faso and Niger.

? Variation in wood density was significant due to provenances and families within provenances. Individual tree heritability was higher for wood density than for growth traits. Provenance means for wood density increased from the more humid to the drier parts of the sample region. Phenotypic correlations indicated that larger trees tended to have denser wood.

? Clines suggest that natural populations of P. africana from the drier parts of the sample region have the genetic capacity to produce denser wood, compared with populations from the more humid parts of the region. Correlations and heritability estimates suggest that selection of faster-growing trees may produce a small gain in wood volume but little (if any) gain in density in the subsequent generation. Multi-location provenance/tests are needed to confirm this tentative conclusion.

     

Live fences in Ségou, Mali : an evaluation by their early users

Market-oriented production of gardening crops and cassava (Manihot utilissima) in the dry season is an increasingly frequent practice in Ségou, Mali. Traditionally, these crops are protected from roaming livestock with the help of dead fences. In order to provide a sustainable alternative to dead fences, the International Centre for Research in Agroforestry (ICRAF) began promoting the use of live fences, living trees planted closely together around a field plot. This study was conducted with the first farmers to use these live fences. These farmers expressed satisfaction with the protection offered by the trees and their ability to provide a variety of medicinal, economic, and food products. The form of land tenure, the social status of farmers within their families, and the availability of labour seem to be important factors in the decision to test the live fence. This raises questions about the accessibility of this technique and its possible contribution to the social and economic differentiation of its users. This revised version was published online in June 2006 with corrections to the Cover Date.     

A national level forest resource assessment for Burkina Faso - A field based forest inventory in a semiarid environment combining small sample size with large observation plots

Even though considerable parts of the global tropical forests are located in Africa, reliable data on African forest resources is limited. While this is widely recognized for tropical moist forests, it also holds for tropical dry forests. To partially fill the gap a forest inventory was carried out in Burkina Faso, West Africa. In this paper we present a methodological approach and sample based estimates of the tree and forest resources including estimates of (1) land cover classes, (2) species composition, and (3) above ground tree carbon stocks. Following the land classification of the Food and Agriculture Organization of the United Nations (FAO), the forest cover of Burkina Faso was estimated as 42.6% (116,847 km²). For the classes “other wooded land”, “other land” and “other land with tree cover” the estimates were 1.6%, 53.6%, and 9.1%, respectively. We found notable differences to the estimates published by FAO, in particular when considering the classes “forest” and “other wooded land” separately, but lesser so when the two classes are combined. That points to a major issue in applying these class definitions in semiarid environments. Given the relatively small sample size (n = 46 field observed plots), relative standard errors (SE%) of area estimates are high (around 9% for the larger area classes). Aboveground tree carbon stocks were estimated to be 6.640, 5.580 and 7.222 Mg ha⁻¹ for “forest”, “other wooded land” and “other land with tree cover”, respectively (SE% around 18% for all three estimates). Availability of biomass models is very limited for all classes, in particular when it comes to shrubs. Furthermore, it was estimated that the most abundant tree species in Burkina Faso is Vittelaria paradoxa, the “shea butter tree” which is a multi-use tree species of high relevance for rural livelihoods.To our knowledge this study is the first field-based forest inventory on national level in Burkina Faso where the estimation of errors was possible on statistical grounds, and done. The results of this study revealed major issues that should be taken into account when doing similar studies, including carbon monitoring and accounting: increasing the sample size will lead to smaller standard errors (at a higher costs, of course), but will not solve the crucial points (1) of non-availability of suitable biomass models, in particular for shrub lands and (2) of implementation issues regarding the definition of land cover types.     

Inter- and intra-provenances variations in seed size and seedling characteristics of Khaya senegalensis A. Juss in Burkina Faso

Within- and between-provenance variations in seed and seedling traits of Khaya senegalensis A. Juss were studied at INERA in Burkina Faso. Nursery grown seedlings from four provenances in Burkina Faso were used for the study. The studies revealed significant variability in all traits evaluated. Seed length and weight significantly varied among provenances and families within provenances, where Bopiel and Koyenga had the highest mean values. Height and root collar diameter of 1 year old seedlings significantly varied among families within provenances. Except leaf biomass ratio and carbon isotope ratio which varied significantly among provenances but not among families within provenances, all other seedling biomass traits—total plant biomass, stem biomass ratio, leaf biomass ratio, root biomass ratio, root shoot ratio, specific leaf area, and leaf area ratio were significantly affected by provenances and families within provenances. The magnitude of variation due to family effect ranged from 65 to 93 % for seed size traits, and from 4.5 to 17.8 % for seedling characters. Estimates of family heritability were moderate to high (0.67–0.95) for seed traits, but low to moderate (0.19–0.59) for seedling characters, indicating that much of the total variation in seed traits is due to the genetic effect. The two most prominent provenances, Koyenga and Bopiel, with higher seed size and greater seedling growth could be considered for an eventual K. senegalensis improvement program in Burkina Faso.     

西双版纳现有混农林系统的主要模式

本文在对西双版纳现有混农林系统进行广泛调查的基础上,根据国际混农林研究委员会(ICRAF)P.K.R.Nair的分类方法,将其进行了分类和评价。结果表明,该地区现存混农林系统的模式极为丰富,并以经济植物为主体的模式占较大优势。这些模式,有的已经发挥出较好的生态、经济和社会效益,有的仍处于实验阶段,有的则对合理利用当地自然资源造成不良的影响。提出了必须合理开发利用当地自然资源,积极创立和引进以粮食作物为主体的混农林系统模式的见解。