Biomass production of trees and grasses in a silvopasture system on marginal lands of Doon Valley of north-west India. 1. Performance of tree species

Bouldery riverbed lands occupy about one-third of the geographical area in Doon Valley of north-west India. These marginal lands need rehabilitation through establishment of silvopasture system for meeting the biomass requirements of resource-poor farmers. A study was carried out for 14 years (1980 to 1993) at Dehradun, involving four multi-purpose tree species viz. Albizia lebbek (L.) Benth., Grewia optiva Drumm., Bauhinia purpurea L. and Leucaena leucocephala (Lamk.) de Wit.; and two grass species viz. Chrysopogon fulvus (Spreng.) Chiov. and Eulaliopsis binata (Retz.) C.E. Hubb. Fifty and 75% of the tree crown was lopped once a year after four years of age. Tree survival decreased with age, and was highest with L. leucocephala (86.9% in 1993 of initial density of 625 trees ha⁻¹) and lowest with B. purpurea (49.3%). Canopy parameters of trees progressed differently, presumably due to their varying growth habits. The trees of A. lebbek were taller with more crown spread, whereas G. optiva had more crown height with lesser crown spread. Stem diameter growth of A. lebbek was highest followed by B. purpurea. Lopping intensity did not affect tree survival and also canopy growth in most years, but collar diameter and diameter at breast height (dbh) decreased significantly at the higher intensity of lopping. Intercropped grass species E. binata caused a greater adverse effect on the tree survival and growth parameters than C. fulvus. The biomass production (leaf and branch wood weight of lopped material) was highest at eight years of age in 1987 but showed no definite trend over the study period. Bauhinia purpurea produced the highest biomass and L. leucocephala the lowest, despite opposite trends in their survival. The performance of A. lebbek was on par with B. purpurea in the later years (1990--1993), but G. optiva was inferior despite its superiority in the early years (1984--1985). The effects of lopping intensity were not appreciable, but intercropping with E. binata decreased leaf and branch wood production significantly compared with C. fulvus throughout. However, the adverse effect was comparatively less pronounced on B. purpurea and A. lebbek than on G. optiva and L. leucocephala. The fibre yield of G. optiva also decreased due to intercropped E. binata compared with C. fulvus. This revised version was published online in June 2006 with corrections to the Cover Date.     

吉林省西部苏打盐碱土区林草植被恢复技术研究

笔者阐述了在苏打盐碱土区种植不同的树、草植被进行试验研究,对比选出适宜盐碱土区种植生长的树、草品种,治理吉林省西部苏打盐碱土区的脆弱的生态现状,研究表明:西伯利亚白刺、吉生四号羊草抗盐碱能力强,模式(羊草0.5 m ×白刺0.5 m ×羊草)×1.0 m最适合迅速提高植被盖度.     

Acacia nilotica-based silvipastoral systems for resource conservation and improved productivity from degraded lands of the Lower Himalayas

High population density and unscientific land use practices have induced severe land degradation in the fragile Lower Himalayan ecosystem of India. Land cover management has been an effective strategy in managing land degradation through the reduction of water, soil and nutrient losses and improvement in soil fertility and quality. Acacia nilotica (Acacia)-based silvipastoral systems with five intercrops, viz., Eulaliopsis binata (bhabbar), Saccharum munja (munj), Vetiveria zizanioides (vetiver), natural grasses and no grass, were evaluated in a long-term study in degraded bouldery lands in Haryana state of India. All grasses resulted in a reduction of soil, water and nutrient losses and improved microbial properties. However, their association adversely affected the growth of Acacia, and the decline varied with grass species. After 11 years of establishment, sole Acacia plantation had the maximum height (7.58 m), diameter at breast height (dbh) (21.32 cm) and crown spread (7.41 m). Munj produced the highest biomass under Acacia, but most adversely affected its growth, resulting in minimum survival (48 %), height (7.07 m), dbh (16.23 cm) and crown spread (6.57 m). Yield of all the grasses increased during the initial 5–6 years, but declined sharply thereafter, with the maximum decline in bhabbar. Detailed investigations established that the sharp decline in survival and growth of bhabbar was due to shade and not because of nutrient competition or allelopathy. Acacia + V. zizanioides proved the most effective silvipastoral system for resource conservation and biomass production. It also provided the highest NPV (Rs 1.88 lakhs ha^?1), B:C (2.37) and IRR (24.70 %) as compared to Rs 6,998 ha^?1, 1.05 and 8.76 % under pure Acacia plantation, respectively.     

Agroforestry for soil and water conservation in the western Himalayan Valley Region of India 2. Crop and tree production

A ten-year-study (1983 to 1992) conducted on nine 15 × 90 m runoff plots at 4% slope compared production efficiency of Leucaena leucocephala and Eucalyptus hybrid based agroforestry as well as monocropping landuse systems in the warm, subhumid climate of the western Himalayan region of India. Treatments for the first sequence were: monocropping systems of leucaena, eucalyptus, Chrysopogon fulvus grass and maize – wheat rotation, and alley cropping systems of grass and crops at 4.5 and 10.5 m alley widths with paired contour tree rows of leucaena and eucalyptus. In the second sequence, alley width increased to 22.5 m in 1989, grass was replaced by turmeric Curcuma longa and paired contour rows of leucaena hedges were introduced in monocropping systems of grain crops and turmeric. Integration of leucaena and eucalyptus trees with crops caused severe reduction of crop yields ranging from 21 to 92% for wheat grain, 59 to 69% for maize grain, 60 to 67% for dry grass and about 50% for turmeric rhizome depending upon the age of trees and alley width. The grain yield of crops stabilized at about 50% reduction with 22.5 m alley width. Total crop biomass (grain + straw) also revealed a similar trend; however, its magnitude of reduction was less severe than for grain. Production of biomass was much lower near the tree rows than in mid alleys. Managing leucaena as contour hedgerows eliminated crop yield reduction in alleys. Performance of grass and turmeric in alleys was not found to be satisfactory. Biomass produced from trees adequately compensated the crop yield reduction. Land equivalent ratios of agroforestry landuses were comparable or even better than monocropping systems indicating suitability of these systems for the western Himalayan valley region. This revised version was published online in June 2006 with corrections to the Cover Date.     

金沙江流域龙须草的水土保持效应研究初报

对金沙江流域龙须草地水土保持效应的研究结果表明,与空旷地相比种植龙须草可降低土壤容重0.10 g/cm3,增加土壤总孔隙度9.6%,提高土壤贮水率22.03%,其改善土壤的物理性状,提高土壤蓄水能力的效果显著。     

Forage production and nitrogen nutrition in three grasses under coconut tree shades in the humid-tropics

Reduction in forage production (FP) under trees in the humid tropics is well known, but information on how different levels of nitrogen (N) fertilizer influence FP under trees is meager. The present study reports effects of four N fertilizer levels (0, 60, 80 and 120 kg ha⁻¹ N) on net soil N mineralization rate (NMR) and soil moisture (SM), FP, shoot biomass/root biomass ratio (SB/RB), N concentration in SB, N uptake and nitrogen use efficiency (NUE) of three grasses [guinea (Panicum maximum Jacq.), para (Brachiaria mutica (Forssk) Stapf) and hybrid-napier (Pennisetum purpureum Schumach.)] under three canopy positions [under canopy (UC, representing high shade), between canopy (BC, representing low shade) and open] of coconut trees (Cocos nucifera L.) in a coconut based silvopastoral system in the humid tropical climate of South Andaman Island of India. The study was performed for two annual cycles (2005–2006 and 2006–2007). The hypotheses tested were: (1) FP would decline under tree shades, both in N fertilized as well as no N fertilized conditions, when SM was not growth limiting in the open. However, amount of decline in the FP would depend on grass species and intensity of shades i.e., higher was the shade greater would be the decline; (2) N fertilizer would increase FP under tree shades, but the increase depended on grass species, intensity of shades and amount of N applied. Amount of N applied, however, would not annul the shades effects when SM was not growth limiting in the open. The study revealed that the tree reduced light 59% under UC and 32% under BC positions, but the N fertilizer levels increased NMR by 11–51% under UC and 3–44% under BC positions compared to the open. SM did not differ across the canopy positions. Under all situations, FP of all grasses declined under UC (47–78%) and BC (18–32%) positions compared to the open; the decline was greater in Hybrid-napier than Guinea and Para grasses. Forage production of all grasses increased with N fertilizer increments under all canopy positions reaching 32 t ha⁻¹ dry matters for hybrid-napier at 120 kg ha⁻¹ N in the open. Both guinea and para grasses outyielded hybrid-napier grass under UC but not under BC or in the open. N concentration in the forage (SB) also increased as N fertilizer level increased. These observations support our hypotheses and suggest that forage production under coconut palms can be increased by the application of N fertilizer with both guinea and para grasses being more productive than hybrid-napier grass under the high shade. Where light conditions are better, hybrid-napier would produce more forage than the other species.     

Morphological and physiological responses of beech (Fagus sylvatica) seedlings to grass-induced below ground competition

We examined morphological and physiological responses of beech (Fagus sylvatica L.) seedlings to grass-induced below ground competition in full-light conditions. Two-year-old beech seedlings were grown during two growing seasons in 160-l containers in bare soil or with a mixture of five grass species widely represented in semi-natural meadows of central France. At the end of the second growing season, beech seedlings in the presence of grass showed significant reductions in diameter and height growth, annual shoot elongation, and stem, root and leaf biomass, but an increase in root to shoot biomass ratio. Grasses greatly reduced soil water availability, which was positively correlated with daily seedling diameter increment. Beech seedlings seemed to respond to water deficit by anticipating stomatal closure. There was evidence of competition for nitrogen (N) by grasses, but its effect on seedling development could not be separated from that of competition for water. By labeling the plants with 15N, we showed that beech seedlings absorbed little N when grasses were present, whereas grasses took up more than 97% of the total N absorbed in the container. We conclude that, even if beech seedlings display morphological and physiological adaptation to below ground competition, their development in full-light conditions may be strongly restricted by competition from grass species.     

半干热石质山地墨西哥柏伴生树种选择研究

在云南省开远市浑水塘试验地设置了35个20 m×20 m的标准地,对标准地内4 a生的主要造林树种墨西哥柏及4个供选伴生树种（羽叶山黄麻、新银合欢、直干桉、赤桉）地径、树高、胸径、冠幅分别进行测量,选择平均木,对平均木生物量、生长量进行测定.经数据分析后认为,新银合欢生长速度慢于墨西哥柏且枝少叶疏,适宜作为墨西哥柏的伴生树种,羽叶山黄麻冠幅虽大,但枝叶稀疏,可作为墨西哥柏的次适宜伴生树种.     

Pb、Cd复合污染对龙须草光合作用的影响

通过九组不同水平的重金属Pb、Cd复合污染土壤种植龙须草试验,以检测对龙须草光合作用的影响,检测龙须草对重金属复合污染的耐性,从而探讨龙须草修复重金属污染环境的能力。光合分析的结果表明：随着Pb、Cd复合污染含量的升高,龙须草的光合综合指标下降幅度较小,说明龙须草对重金属复合污染具有较强的耐性,且Cd对龙须草的影响大于Pb。     

Agroforestry for soil and water conservation in the western Himalayan Valley Region of India 1. Runoff, soil and nutrient losses

Erosion losses were investigated in a field experiment, from 90 × 15 m erosion plots at 4% slope in the western Himalayan valley region of India. The plots were planted with Leucaena leucocephala and Eucalyptus hybrid, either as block plantation or in alley farming with maize (Zea mays), Chrysopogon fulvus grass or turmeric (Curcuma longa). The treatments also included contour planting of sole maize, sole Chrysopogon fulvus grass and a clean weeded cultivated fallow. During the nine year study period, the average annual monsoon rainfall was about 1000 mm and it caused 347 mm runoff and 39 Mg ha^-1 soil loss per year from fallow plots. The runoff and soil loss were reduced by 27% and 45% by contour cultivation of maize. Contour tree-rows or leucaena hedges reduced the runoff and soil loss by 40% and 48%, respectively, over the maize plot, reducing soil loss to about 12.5 Mg ha^-1. This reduction in erosion was primarily due to the barrier effect of tree or hedgerows and micro-terraces formed through sediment deposition along the contour barriers. Such vegetative measures, that are productive while being protective, offer viable alternative for erosion control in areas with gentile slopes of the valley region. High density block plantations of eucalyptus and leucaena almost completely controlled erosional losses and can be recommended for steeper slopes that are vulnerable to heavy erosion. This revised version was published online in June 2006 with corrections to the Cover Date.     

Early seedling establishment of two tropical montane cloud forest tree species: The role of native and exotic grasses

Tropical montane cloud forest has been undergoing a drastic reduction because of its widespread conversion to pastures. Once these forests have been cleared exotic grasses are deliberately introduced for forage production. Exotic grass species commonly form monodominant stands and produce more biomass than native grass species, resulting in the inhibition of secondary succession and tree regeneration. The purpose of this study was to assess the effect of native vs. exotic grass species on the early establishment of two native tree seedlings (Mexican alder, Alnus acuminata and Jalapa oak, Quercus xalapensis) on an abandoned farm in central Veracruz, Mexico. Seedling survival and growth were monitored (over 46 weeks) in relation to grass cover and height, and available photosynthetic active radiation (PAR). More seedlings survived in the presence of the native grass Panicum glutinosum than those growing with the exotic grass Cynodon plectostachyus (92% vs. 48%). The causes of seedling mortality varied between species; Q. xalapensis was affected by herbivory by voles but mainly in the exotic grass-dominated stands, whereas A. acuminata seedlings died due to competition with the exotic grass. A. acuminata seedlings increased more in height in the exotic grass-dominated stands (102 ± 7.8 cm) compared to native grass-dominated stands (51 ± 4.7 cm). Grass layer height, cover and available PAR were correlated (Pearson; p < 0.05). In the exotic grass dominated plots, grass layer height was correlated with the relative height growth rates of Q. xalapensis (Pearson; p < 0.05). These results indicate that the exotic grass may be affecting tree regeneration directly (grass competition) and indirectly (higher herbivory). Passive restoration may occur once P. glutinosum dominated pastures are abandoned. However, when C. plectostachyus dominates, introduction of early and mid successional tree seedlings protected against vole damage is needed.     

Modelling competition of paired columns of Eucalyptus on interplanted grass

Adequate evaluation of competition interfaces of tree growth on intercrops is an important consideration in the assessment of agroforestry systems. Spatial methods wherein positional effect of an observation with respect to trees or columns of trees can be accounted for were examined. Yield ofbhabbar grass (Eulaliopsis binata Ritz) planted under paired and staggered columns of Eucalyptus (Eucalyptus tereticornis Smith) was analysed for four years. Grass production at 7 positions recorded across staggered and paired tree columns was significantly different. Spatial correlations among contiguous observations across paired tree columns were significant. Cyclicity corresponding to seven columns of grass of 3.5 m distance between paired tree columns explained variation in grass production significantly during the first observation year. Additional competition interfaces at 1.75 m and 1.17 m appeared in the subsequent years. These competition interfaces were attributed to pairing of trees and staggering. Error distributions of fitting of cyclic patterns were more acceptable as compared to ANOVA. Possibilities of the use of this methodology in many other agroforestry situations are discussed.     

Growth Parameters and Nutrient Content in Four Multipurpose Tree Species with Potential Characteristics for Agroforestry Systems in a Cerrado Region in Botucatu,São Paulo State,Brazil

Abstract

In order to evaluate growth characteristics, adaptability, biomass production, nutrient recycling, nutrient distribution and the ability to regenerate degraded land, a trial using four multipurpose tree species (Leucaena leucocephala, Leucaena diversifolia, Acacia melanoxylon and Mimosa scabrella) was undertaken over two years in a distrophic red yellow latosol (oxisol) following a randomized block experimental design with four replications.

At the age of two years, A. melanoxylon and L. diversifolia were the tallest species (5.25 and 4.97 m, respectively) and A. melanoxylon and M. scabrella had the largest diameters at 20 cm from tree base.

Mimosa scabrella and A. melanoxylon had the highest dry matter production and quantity of nutrients in the above ground biomass. In all species, the highest nutrient contents were found in the leaves, followed by branches and stems. From all species, the highest Nutrient Utilization Efficiency Indexes were obtained for sulphur, phosphorous, and magnesium; L. diversifolia was the most efficient for nitrogen, potassium, calcium, sulphur, and manganese, while A. melanoxylon was the most efficient for phosphorus, magnesium, boron, iron, and zinc. Litter production levels over a three month period were as follows: M. scabrella > A. melanoxylon > L. diversifolia > L. leucocephala. Litter nutrient content was higher in M. scabrella than in the other species.     

Alley cropping of maize with calliandra and leucaena in the subhumid highlands of Kenya: Part 2. Soil-fertility changes and maize yield

Although N-rich leaf biomass of multipurpose trees is known to be a good source of N to crops, integrating such trees into crop production systems is a major challenge in the development of viable agroforestry systems. An approach to integrating calliandra (Calliandra calothyrsus Meissner) and leucaena (Leucaena leucocephala (Lam.) de Wit), two promising agroforestry tree species, into maize (Zea mays L.) production system was investigated in the subhumid highlands of central Kenya during four maize-growing seasons from 1994 to 1996. The experiment consisted of maize plots to which tree prunings obtained from hedgerows grown either in situ (alley cropping) or ex situ (biomass transfer from outside) were applied. When alley-cropped with leucaena, maize produced significantly higher yields compared to maize monoculture (both non-fertilized and fertilized) treatments, but when alley-cropped with calliandra, the yield of maize was less than that of the monocropped unfertilized control. Application of ex situ grown calliandra and leucaena prunings with or without fertilizer resulted in higher maize grain yield than in the nonfertilized and fertilized treatments. Yields of calliandra alley- cropped maize were 11% to 51% lower than those of nonalley-cropped treatments receiving calliandra prunings from ex situ grown trees; the decrease was 2% to 17% with leucaena, indicating that calliandra hedges were more competitive than leucaena hedges. The alley-cropped prunings-removed treatments produced the lowest maize yields. The study showed that, in the subhumid tropical highlands of Kenya, inclusion of calliandra hedges on cropland adversely affected maize yields. On the other hand, alley cropping with leucaena was advantageous. This revised version was published online in June 2006 with corrections to the Cover Date.     

Site factors associated with the performance of Leucaena leucocephala (Lam.) de Wit and Sesbania sesban (L.) Merill in pure and mixed stands in the northern highlands of Rwanda

Alley cropping with leguminous tree hedgerows planted on contours is an emerging practice in the northern highlands of Rwanda where field slopes range from 4 to 76% and loss of soil fertility due to erosion is the principal impediment to food production. Sesbania sesban and Leucaena Leucocephala, the two woody species recommended for alley cropping in the region, do not perform equally well across sites: Sesbania is faster growing, But is more sensitive to pruning than Leucaena. We tested the hypothesis that the two species could be grown together to make the best use of their relative advantages. Species were planted in pure and 1:1 mixed stands at 26 sites to give a range of altitudes, slopes, azimuths and soil fertility levels representative of the region. During two cropping seasons’ growth spanning 19 months, in pure stands, Sesbania produced more biomass, grew taller and had thicker stems than Leucaena: for the same species, stem diameter and height were not different in pure and mixed stands. Biomass and number of poles from 1:1 mixed stands of Sesbania and Leucaena were equivalent to biomass and pole numbers from Sesbania in pure stands. However, mixed stands produced more biomass and pole numbers than Leucaena in pure stands. Growth of S. sesban was positively correlated with soil pH, potassium and altitude; whereas, growth of L. leucocephala was positively correlated with soil phosphorus, but negatively correlated with field slope. All other soil and site variables tested were nonsignificant. The study suggests that farmers can plant S. sesban and L. leucocephala in 1:1 mixtures in an alley cropping setting and retain levels of biomass production and bean pole numbers equivalent to those from S. sesban alone without compromising future growth and production by Leucaena. As well, the impact of pH, K and P on growth and production of Sesbania and Leucaena as a function of field slope and altitude should be determined before extending their use to other regions. This revised version was published online in June 2006 with corrections to the Cover Date.     

Water harvesting in a ‘runoff-catchment’ agroforestry system in the dry lands of Ethiopia

A major production constraint in arid and semiarid areas of Ethiopia is a lack of water for crop growth. Run off water can be harvested by channeling it into micro- catchments (MC) where it slowly infiltrates into the soil. The increased moisture provides more plant growth in these dry lands. An experiment using MC was conducted in eastern Ethiopia to study the growth of four multipurpose tree species intercropped with grass. Trees and grass were grown in 25 m² and 100 m² MC. Plant height, root collar diameter, and mortality rate of trees were determined 12 months after planting. Dry matter yield of grasses and trees were measured and nutrient analysis of plant tissues was determined. In a separate experiment the biomass of trees was determined after 2.5 years of continuous growth without grass competition. Soil moisture, organic matter, texture and bulk density of the soils were also determined. Mean tree height was 10% greater in the 100 m² than in the 25 m² MC. Root collar diameter and survival rate showed similar increase with 13% and 7.8% respectively. Acacia saligna and Leucaena leucocephala showed better growth in both plot sizes than the indigenous Acacia seyal and Acacia tortilis. The dry matter yield of grass (Panicum maximum) was over 12 tons/ha and 8 tons/ha in the 100 m2 and 25 m2 plots respectively. Soil moisture content was greater in MC than in control plots with no water harvesting structures with 31% and 24% during the wet and dry seasons respectively. The use of water harvesting can improve fodder production and carrying capacity of the dry lands of Ethiopia.     

Growth, productivity and quality of Megathyrsus maximus under cover from Gliricidia sepium

The effect of tree canopy on the growth, productivity and forage quality of Megathyrsus maximus and changes in soil properties were evaluated over three seasonal periods. Four adjacent plots (15 m × 17 m each) in a tropical secondary deciduous forest having 12 years of growth and dominated by Gliricidia sepium were randomly assigned to two treatments: removal of trees (SCA) in two of the plots and leaving trees intact (COA) in the other two. In all plots, M. maximus was planted with 50 cm spacing among plants. Tree removal significantly increased the incident photosynthetically active radiation (PAR, P < 0.001) and grass size (12.5 % in height, P < 0.01, and 16.5 % in clump diameter, P < 0.05), but did not significantly affect any other variable. Season significantly affected grass height (P < 0.003), tiller number (P < 0.001), clump diameter (P < 0.001), net CO₂ assimilation rate (P < 0.001), forage biomass production (P < 0.003), and acid detergent fiber content (P = 0.033). Primary soil changes after 1 year of establishment of the grass were the decline by 3 % in organic carbon (P = 0.03), and qualitative changes in soil structure, regardless of tree presence. Results are consistent with the ability of M. maximus to tolerate shade. We conclude that under the conditions of the study there was no evidence for a negative effect of tree canopy on M. maximus mediated by a reduction in PAR.     

Studies on root distribution of five multipurpose tree species in Doon Valley,India

Observations on the growth performance, rooting behaviour and distribution of fine roots of five tree species viz., Bauhinia purpurea, Grewia optiva, Eucalyptus tereticornis, Leucaena leucocephala and Ougeinia oojeinensis (Family: Papilionaceae) are being presented here. Roots were exposed at the time of planting, 6 months, 16 months, and 28 months after planting. Total root weight and root volume were highest in Eucalyptus tereticornis and lowest in Bauhinia purpurea. Major part of the root system confined within 90–120 cm soil depth in case of Bauhinia purpurea, Grewia optiva and Leucaena leucocephala but Eucalyptus tereticornis and Ougeinia oojeinensis strike their roots to deeper depths. Bauhinia purpurea had its roots evenly distributed down to 120 cm. In general, the vertical distribution of fine roots (< 2 mm in diameter) of the five species indicate that more the depth, fewer the number of roots. The observations on soil binding capacity, indicated that Ougeinia oojeinensis had the maximum and Eucalyptus tereticornis had the minimum binding value. Due to leaf shading and other litter fall significant increase of nutrient components in soils under the tree canopies has been observed. The study indicates that bulk of the roots of the five tree species are found near the surface, but observation on soil moisture and nutrient content does not indicate variation under the tree canopies and in open, hence there may not be root competition in initial years of plantation.Authorised for publication by the Institute as contribution No.3311/239/89.     

Effect of lopping on water potential, transpiration, regrowth, (14)C-photosynthate distribution and biomass production in Alnus glutinosa

The effects of light, moderate and heavy branch pruning or lopping treatments (resulting in removal of 28-31, 56-60 and 80-82% of the foliage, respectively, unlopped control = 0%) were studied in 3-year-old black alder (Alnus glutinosa L. Gaertn.) plants. Within 24 h of lopping, transpiration rates decreased and water potentials increased. The effects of lopping continued for 48 days. The improved water status of the lopped plants enhanced water-use efficiency during the first 30 days. Regrowth was related to a combination of enhanced net assimilation rates during the first 30 days after lopping, use of current photoassimilates and stored carbohydrates in the roots (particularly in medium and heavily lopped plants) for the production of new leaf area, and delayed leaf senescence. The lightly and moderately lopped plants developed 62 and 57% more leaf area, respectively, than the unlopped plants, but total leaf area development was slower in the heavily lopped plants. Lightly lopped plants produced total biomass equivalent to that of unlopped plants, but the moderately and heavily lopped plants produced less biomass than the unlopped plants. During the first 30 days after lopping, there was a net loss of nodule dry matter and recovery to control values did not occur during the period of study. Immediately (24 h) after lopping, more (14)C-photosynthate was translocated to the roots of lopped plants than to the roots of unlopped plants. When a branch near the base of the main stem was fed (14)CO(2), (14)C-photosynthate translocation to the shoot increased with increasing intensity of lopping, at 50 days after lopping.     

Fine root biomass,distribution, and production in young pine-hardwood stands

Patterns of fine root biomass, production, and distribution were estimated for pure stands and mixtures of three-year-old loblolly pine (Pinus taeda L.) with red maple (Acer rubrum L.) or black locust (Robinia pseudoacacia L.) on the Virginia Piedmont to determine the role of fine roots in interference between pine and hardwood tree species. Estimates were based on amounts of live and dead fine roots separated from monthly core samples during the third growing season after planting. Live and dead fine root biomass and production varied by species, but mixtures of loblolly pine and black locust generally had greater fine root biomass and fine root production than pure stands or loblolly pine-red maple mixtures. Hardwood species had greater live fine root biomass per tree in mixtures with pine compared to pure stands. Greater live fine root biomass in pine-locust stands may be attributed to differential utilization of the soil volume by fine roots of these species. For all stands, approximately 50% of live five root biomass was located in the upper 10 cm of soil.