Effects of grass silage feed value on feed intake and performance of pregnant and lactating ewes and their lambs

Effects of grass silage feed value on intake, dietary sorting, body condition (BC) and body weight (BW) of pregnant and lactating ewes and on liveweight gain (LWG) of their suckling lambs were evaluated. Twin‐bearing ewes were fed ad libitum grass silage of (i) high feed value (HFV), (ii) medium feed value (MFV) and (iii) low feed value (LFV), plus 0·8 kg concentrate in late pregnancy and lactation. The HFV, MFV and LFV silages contained 11·7, 10·8 and 9·3 MJ metabolizable energy (ME) kg^?1 DM respectively. The HFV resulted in 8·2 and 15·2 MJ greater ME intake in pregnancy and 14·7 and 21·0 MJ greater ME intake from silage in lactation than MFV and LFV respectively (P < 0·01). Ewes fed the MFV and LFV silages sorted for more digestible particles during eating (P < 0·10). The HFV gave no loss in BC, whereas ewes fed MFV and LFV lost 0·7 and 0·8 units of BC, respectively, during lactation (P < 0·10). There were similar trends in BW. The nursing HFV lambs had 73 g higher daily LWG than the LFV lambs (P < 0·01). In conclusion, the HFV increased the ME intake of pregnant and lactating ewes, resulting in improved BC and BW of lactating ewes and LWG of suckling lambs compared to MFV and LFV.     

Diet selection of dry and lactating beef cows grazing extensive pastures in late autumn

The diet selection, made by twelve dry and twelve lactating spring‐calving Salers beef cows at the end of the grazing season on an extensively grazed natural mountain pasture in central France, was compared. Sward structure, selection bites, daily grazing time and digestibility of the diet were measured in mid‐October (P₁) and mid‐November (P₂). Live weights were measured twice monthly throughout the experiment. Daily milk production was assessed in the week before the grazing behaviour measurements. Lactating cows grazed more selectively than dry cows: they took more bites on green patches (0·20 vs. 0·13 of total bites, P < 0·01; Jacob's selectivity index: 0·13 vs. ?0·25, P < 0·001), which is consistent with the higher overall nitrogen concentration in faeces (18·1 vs. 17·3 g kg^?1 DM, P < 0·05). Lactating cows tended to slightly increase their daily grazing time compared with dry cows (on average 9·9 vs. 9·5 h, P = 0·07) and grazed faster in mid‐November (74 vs. 69 bites min^?1, P < 0·05). The lactating cows lost more live weight (?24 vs. ?12 kg cow^?1, P < 0·05) between P₁ and P₂. A positive correlation was found between consumption of green patches by lactating cows and their daily milk production in P₂ (r = 0·574, P = 0·05), whereas no correlation occurred between individual milk production and liveweight loss. Dry cows were less selective, which supports management practices that promote the use of cows with low requirements on extensively grazed pastures in late season.     

THE EFFECT OF FEEDING RED CLOVER CONSERVED BY DRYING OR ENSILING ON REPRODUCTION IN THE EWE

Forage diets were prepared from ryegrass (non oestrogenic control) and from red clover as pellets or as unwilted silage. Each diet was fed to separate groups of 20 Border Leicester × Cheviot ewes for three weeks prior to the introduction of the ram, and for two cycles after mating. Subsequently, all three groups of ewes were fed in a similar manner until lambing and the reproductive performance of the ewes was measured. The control diet was prepared by barn drying a primary growth crop of perennial ryegrass, Lolium perenne cv S23. Tetraploid red clover, Trifolium pratense cv. Hungarpoly, was harvested as a first regrowth crop and the pelleted diet prepared by high temperature drying, grinding (2 mm screen) and pelleting. The silage was prepared without wilting. The level of formononetin in the red-clover silage was three times that in the dried red-clover pellets. Compared with the control grass diet, the feeding of red clover in either form significantly reduced (P<0·001) the number of lambs born. The incidence of barrenness was higher (P<0·001) for ewes fed red-clover silage compared with red-clover pellets and the control diet. The rate of twinning was significantly (P<0·001) higher for the ewes fed the control grass diet than for either of the red-clover diets. The resultant lambing percentages were 190,165, and 100% for the control grass, red-clover pellet and red-clover silage treatments. In flock management, both the grazing of red clover and the feeding of red-clover pellets or silage, prior to and during mating, should be avoided.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 2. Animal production

Performance of continuously stocked Mule ewes nursing Suffolk-cross twin lambs over three grazing seasons, between April and August, was compared on swards of N-fertilized diploid perennial ryegrass (D), tetraploid perennial ryegrass (T) and tetraploid perennial ryegrass with white clover (TC), the latter receiving no fertilizer N. Sward height was maintained by variable stocking rate close to a target of 4–6 cm (constant treatment) from turnout and compared in July and August with a rising sward height treatment (target 6–8 cm). Lambs on TC swards had significantly higher (P <0·001) liveweight gains compared with lambs on T swards by 41 gd^-1 in April–June and by 68gd^-1 in July-August. Live weight and body condition score of ewes in August were significantly higher (P<0·001) on TC compared with T swards, by 11·3 kg and 0·75 respectively. Rising sward heights in July–August increased live-weight gain of lambs compared with constant sward heights by 102, 39 and 54gd^-1 in consecutive years, associated with sward height increases of 0·9, 0·5 and 0·6cm respectively. Rising sward height increased ewe live weight and body condition score by 5·1 kg and 0·3 respectively, compared with results from constant sward heights. Effects of sward height and sward type were additive. T swards had a significantly (P<0·01) 16% greater overall lamb output than the D swards due mainly to a 10% higher achieved stocking rate. Stocking rates of ewes on TC vs T swards were 40, 13 and 12% lower in April-August in successive years. The higher liveweight gain of lambs on the TC swards resulted in lamb outputs of 76, 105 and 101% of the T swards in successive years, showing that grass/clover swards containing over 20% clover could produce similar lamb output ha^-1 to grass swards given 150–180 kg N ha^-1.     

Seasonal patterns of diet composition, herbage intake and digestibility limit the performance of cattle grazing native pasture in the Falkland Islands

The botanical composition, intake and digestibility of the diet consumed by mature lactating and non‐lactating cows grazing a native white grass (Cortaderia pilosa) plant community in the Falkland Islands was measured in four periods between September 1998 and June 1999. Five lactating and five non‐lactating cows were used in the summer, autumn and winter; five non‐lactating cows were used in the spring. Different cows were used in each period. Plant cuticle patterns in the faeces of cattle were used, in conjunction with the patterns of concentrations of n‐alkanes in the faeces, to estimate the botanical composition of the diet and predicted concentrations of C₃₂‐ and C₃₃‐alkanes in the herbage allowed herbage intake and digestibility to be estimated using the n‐alkane technique. White grass, sedges and rushes comprised 0·78, 0·64 and 0·63 of the diet in autumn, winter and spring respectively. Fine grasses, smooth‐stalked meadow grass (Poa pratensis L.), annual meadow grass (Poa annua L.), bent grass (Agrostis capillaris L.), native fescue (Festuca magellanica Lam.), Yorkshire fog (Holcus lanatus L.), wavy hair grass [Deschampsia flexuosa (L) Trin.] and early hair grass (Aira praecox L.), were consumed at the expense of sedges and rushes in summer and this coincided with the period of greatest estimated metabolizable energy and crude protein intakes by cows. Lactating cows suffered substantial liveweight loss during autumn and winter and this reflected the low quality of the diet consumed. Phosphorus intakes were insufficient and there was an estimated deficiency of vitamin D in the winter and early spring in cattle that were grazed with the experimental cows. The nutrient restrictions imposed on cattle by the low quality of native pasture during autumn and winter are likely to impair the reproductive potential of breeding females and methods should be investigated to improve the diet consumed by cows during these critical periods if cattle systems are to become sustainable in the Falkland Islands.     

The effects of incorporating small quantities of straw in grass/grass silage-based diets for dairy cows

Two studies were conducted to examine the effects of incorporating small quantities of straw in the diets of dairy cows. In Experiment 1, forty Holstein Friesian dairy cows were used in a 2 × 4 factorial design experiment, with factors examined consisting of two parities (primiparous and multiparous animals) and four levels of straw inclusion in the diet (0, 0·08, 0·16 and 0·24 of forage dry matter). The basal forage offered in this study was grass silage, and the primiparous and multiparous animals were supplemented with 9·0 and 11·0 kg concentrate d^–1 respectively. In Experiment 2, forty‐eight Holstein Friesian dairy cows were used in a 2 × 3 factorial design experiment, with factors examined consisting of two basal forage types (grass silage and zero‐grazed grass) and three levels of straw inclusion (0, 1·0 and 2·0 kg d^–1). All animals were offered 7·0 kg d^–1 of a concentrate supplement. Both experiments were partially balanced changeover designs, consisting of two, 4‐week periods. In Experiment 1, the total dry‐matter intake followed a significant quadratic relationship (P < 0·05), increasing with low levels of straw inclusion and decreasing at higher levels of inclusion. With increasing levels of straw inclusion, there was a linear decline in milk yield (P < 0·001) and milk protein concentration (P < 0·05), but milk fat concentration was unaffected (P > 0·05). In Experiment 2, the effect of straw inclusion on total dry‐matter intake was quadratic (P < 0·001), with intakes being maximum at the 1·0‐kg level of straw inclusion. Milk yield exhibited a linear decrease (P < 0·001) with increasing level of straw inclusion. Milk fat concentration was lowest at the 1·0 kg rate of straw inclusion (P < 0·05), but milk protein concentration was unaffected by straw inclusion. There were no significant interactions between basal forage type and level of straw inclusion for any of the variables examined (P > 0·05). Despite small increases in total dry‐matter intake at a low level of straw inclusion, there was no evidence that straw inclusion improved either nutrient utilization or animal performance. The reduction in milk yield observed with straw inclusion reflects, to a large extent, a reduction in metabolizable energy intake.     

Urea supplementation effects on the utilization of low-quality forage and lamb production in patagonian rangelands

The effect of feeding a urea-based supplement for the last 8 weeks of pregnancy was examined on ewes grazing the grass-shrub steppes of NW Patagonia, Argentina. The productivity of ewes and lambs and the utilization rate of the major, less preferred, grass species (Stipa speciosa, 27% cover) was analysed. In a paddock-scale experiment, ewes grazed two areas of 450–500 ha from 15 August to 15 February at a stocking rate of 0·5 ewes ha^?1. From 15 August to 10 October, one group of ewes was given access to a urea-based supplement containing 300 g kg^?1 urea, 320 g kg^?1 salt, 30 g kg^?1 molasses and 300 g kg^?1 bone meal. Average supplement consumption was 3·5 g ewe^?1 d^?1. The performance of 100 ewes in each of the two areas was evaluated. Supplemented ewes gained 4·2 kg more live weight than the control ewes between 15 August and 10 October (prepartum) (liveweight gain was 10·3 vs. 6·1 kg ewe^?1, P < 0·05). The number of live lambs at 8 weeks of age, as a proportion of ewes mated, was 15% higher in the supplemented than in the control group (0·634 vs. 0·553, P < 0·05). A similar increase in the live weight of lambs at 8 weeks of age in the supplemented group compared with the controls was also observed (12·1 vs. 10·6 kg, P < 0·05). In the paddock with the supplemented ewes, there was a significant increase in the proportion of Stipa speciosa plants defoliated (0·62 vs. 0·38, P < 0·05) and in the intensity of defoliation of these plants. It was concluded that urea-based supplements increased productivity of ewes and that this was likely to be associated with an increase in the intake, and possibly digestibility, of the less preferred species such as Stipa speciosa.     

The effect of forage type and level of concentrate supplementation on the performance of spring-calving dairy cows in early lactation

Abstract In 1993 and 1994, 40 cows in early lactation in early spring were assigned randomly to four feeding treatments. One group of cows was kept indoors with access to grass silage ad libitum, plus 6 kg of concentrate daily. The other three groups had access to grass pasture (5–6 h per day in 1993 and 11–12 h per day in 1994) plus grass silage similar to that fed to the previous group while indoors plus 6, 4 or 2 kg of concentrate daily. The average daily allocations of herbage (> 3·5 cm) were 8·5 and 14·0 kg DM cow^?1 day^?1 in 1993 and 1994 respectively. The treatments were applied for 8 weeks (26 February to 23 April) in 1993, and 7 weeks (11 March to 29 April) in 1994. Cows with access to pasture had lower (P < 0·001) silage dry‐matter (DM) intakes and higher (P < 0·001) total forage DM intakes in both years than those kept indoors. This resulted in significantly higher yields of milk, fat, protein and lactose. Similarly, milk protein concentration was higher (P < 0·05 in 1993; P < 0·001 in 1994). There was a significant linear increase in total DM intake in both years with increased concentrate supplementation. In 1993, there was a linear increase in milk (P < 0·01), fat (P < 0·01), protein (P < 0·001) and lactose (P < 0·01) yields with increased concentrate supplementation. In 1994, only milk protein yield (P < 0·05) was increased. Concentrate supplementation had no effect on milk composition or liveweight change. Cows with access to grazed grass had higher liveweight gains (P < 0·05) than those kept indoors in both years. In 1993, increasing the energy intake increased the processing qualities of the milk produced. The results showed that access to grass pasture resulted in higher milk production, in reduced silage requirement and in reduced level of concentrate supplementation required for a given level of milk production with spring‐calving cows in early lactation compared with those kept indoors.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

Effect of early turnout to grass in spring on dairy cow performance

This experiment examined the effects of grazing severity and degree of silage restriction during early turnout of dairy cows to pasture in spring on animal performance. Forty late‐winter‐calving Holstein Friesian dairy cows were allocated to one of five treatments between 7 March and 17 April 1997. The treatments involved early turnout of cows to grass for 2 h per day at two residual sward heights and two silage allowances, plus a control treatment, in a randomized block design. Dairy cows on the control treatment remained indoors throughout the experiment and were offered grass silage ad libitum. Dairy cows on all treatments were also offered 6 kg d^–1 of a concentrate on a flat‐rate basis, split equally between the morning and afternoon milkings. Offering cows access to pasture in early spring for 2 h per day resulted in increases in both milk (P < 0·001) and protein yield (P < 0·01). On average, over all grazing treatments, cows produced an additional 2·6 kg milk per day compared with the control treatment (28·5 vs. 25·9 kg d^–1, s.e.m. 0·43). Furthermore, these increases in milk yield were obtained even when silage was restricted indoors (28·4 vs. 25·9 kg d^–1) and cows grazed down to a residual sward height of 40 mm (28·1 vs. 25·9 kg d^–1). Protein yield was higher (P < 0·01) with dairy cows grazing pasture compared with cows indoors (848 vs. 707 g d^–1, s.e.m. 28·9). Silage intake was significantly (P < 0·001) reduced when cows were turned out to pasture. In conclusion, early turnout of dairy cows to pasture in spring for 2 h per day reduced silage intake and increased milk yield and protein yield relative to those fully housed and offered grass silage with a low level of concentrates.     

Effect of defoliation management, based on leaf stage, on perennial ryegrass (Lolium perenne L.), prairie grass (Bromus willdenowii Kunth.) and cocksfoot (Dactylis glomerata L.) under dryland conditions. 2. Nutritive value

A field experiment was undertaken between April 2003 and May 2004 in southern Tasmania, Australia, to quantify and compare changes in the nutritive value of perennial ryegrass (Lolium perenne L.), prairie grass (Bromus willdenowii Kunth.) and cocksfoot (Dactylis glomerata L.) under a defoliation regime based on stage of leaf regrowth. Defoliation interval was based on the time taken for two, three or four leaves per tiller to fully expand. At every defoliation event, samples were collected and analysed for acid‐detergent fibre (ADF), neutral‐detergent fibre (NDF) and total nitrogen (N) concentrations and to estimate metabolizable energy (ME) and digestible dry matter (DDM) concentrations. Amounts of crude protein (CP) and metabolizable energy (MJ) per hectare values were subsequently calculated. There was a significantly lower (P < 0·001) NDF concentration for perennial ryegrass compared with prairie grass and cocksfoot, and a significantly lower (P < 0·001) ADF concentration for cocksfoot compared with prairie grass and perennial ryegrass, regardless of defoliation interval. The CP concentration of cocksfoot was significantly greater (P < 0·001) compared with the CP concentrations of prairie grass and perennial ryegrass. The estimated ME concentrations in cocksfoot were high enough to satisfy the requirements of a lactating dairy cow, with defoliation at or before the four‐leaf stage maintaining ME concentrations between 10·7 and 10·9 MJ kg^?1 DM, and minimizing reproductive plant development. The ME concentrations of prairie grass (10·2–10·4 MJ kg^?1 DM) were significantly lower (P < 0·001) than for cocksfoot (as above) and perennial ryegrass (11·4–11·6 MJ kg^?1 DM) but a higher DM production per hectare resulted in prairie grass providing the greatest amounts of ME ha^?1.     

Quantifying the influence of sward height, concentrate level and initial milk yield on the milk production and grazing behaviour of continuously stocked dairy cows

Two factorial design experiments were carried out in the spring of 1994 and 1995, each of 6 weeks, to quantify the effects of sward height (SH), concentrate level (CL) and initial milk yield (IMY) on milk production and grazing behaviour of continuously stocked dairy cows. In Experiment 1, forty‐five Holstein Friesian cows were in five groups with initial milk yields of 16·9, 21·1, 28·0, 31·5 and 35·5 kg d^–1, grazed sward heights were 3–5, 5–7 and 7–9 cm (LSH, MSH and HSH respectively), and concentrates were fed at rates of 0, 3 and 6 kg d^–1. In Experiment 2, 48 cows were in two groups with IMY of 21·3 and 35·5 kg d^–1, grazed sward heights were 3–5 and 7–9 cm (LSH and HSH), and concentrates were fed at 0 and 6 kg d^–1 and ad libitum. Multiple regression models were used to quantify the effects of the three variables on milk yield persistency (MYP), estimated herbage dry‐matter (DM) intake (HDMI), grazing time (GT) and rate of DM intake (RI). The partial regression coefficients showed that increased SH led to increased MYP (Experiment 1 P < 0·001, Experiment 2 P < 0·05), increased HDMI (P < 0·01, P < 0·01), increased GT (P < 0·001, P < 0·05) and increased RI (P < 0·001, P < 0·05). Increasing CL led to increased MYP (NS, P < 0·001), decreased HDMI (P < 0·001, P < 0·001), decreased GT (NS, P < 0·001) and decreased RI (P < 0·001, P < 0·001). Higher IMY level of cows decreased MYP (P < 0·001, P < 0·001), increased HDMI (P < 0·001, P < 0·001), increased GT (P < 0·001, P < 0·05) and increased RI (P < 0·05, P < 0·01). The models were highly significant (P < 0·001), and accounted for 0·48–0·87 of the total variance. The partial regression coefficients quantified the extent to which GT and RI by cows respond positively to higher IMY, and negatively to increased CL, but respond differently (GT declines in response to a higher RI) with increasing SH.     

Stocking rate effects on liveweight gain of ewes and their twin lambs when grazing subterranean clover–perennial grass pastures

Two grazing experiments were conducted on non‐irrigated tall fescue–subterranean clover and cocksfoot–subterranean pastures subject to summer‐dry conditions in Canterbury, New Zealand, to measure the effect of low (8·3–10 ewes and their twin lambs ha^?1) vs. high (13·9–20 ewes and their twin lambs ha^?1) stocking rates (SR) on lamb and ewe liveweight gain in spring. In tall fescue–subterranean clover pasture, lambs grew faster at low (374 g per head d^?1) than high (307 g per head d^?1) SR, but total liveweight gain per ha was greater at high (12·3 kg ha^?1 d^?1) than low (7·5 kg ha^?1 d^?1) SR. In successive years in spring in cocksfoot–subterranean clover pastures, lambs grew faster at low (327, 385 g per head d^?1) than high (253, 285 g per head d^?1) SR but total liveweight gain per ha was greater at high (7·26, 7·91 kg ha^?1) than low (5·43, 6·38 kg ha^?1) SR. These studies indicate that in summer‐dry areas, subterranean clover‐based pastures will support high twin lamb growth rates in spring, with lower SR leading to higher lamb growth rates and more lambs reaching slaughter weights before the onset of dry conditions.     

The effect of concentrate feed level on the response of lactating dairy cows to a constant proportion of fodder beet inclusion in a grass silage-based diet

Abstract The effects of level of concentrate supplementation on the response of dairy cows to grass silage‐based diets containing a constant proportion of fodder beet were examined. Forty Holstein‐Friesian dairy cows of mixed parity were used in a 2 × 5 factorial design experiment. Two basal diet types [grass silage alone or grass silage mixed with fodder beet in a 70:30 dry matter (DM) ratio] were offered ad libitum, and the effects of five levels of concentrate supplementation (mean = 3·0, 5·3, 7·5, 9·8 and 12·0 kg DM per cow d^?1) were examined. Concentrate supplements were offered via an out‐of‐parlour feeding system. These treatments were examined in a three‐period (period length = 4 weeks) partially balanced changeover design experiment. Fodder beet inclusion had no significant effect on the estimated metabolizable energy (ME) concentration of the ration (P > 0·001). Total DM intake, estimated ME intake, milk yield, milk protein content and milk energy output all showed significant linear increases with increasing level of concentrate inclusion (P < 0·001) while, in addition, milk yield and milk energy output exhibited a significant quadratic increase (P < 0·01). The inclusion of fodder beet in the diet reduced silage DM intake (P < 0·01) but resulted in an increase in total DM intake and estimated ME intake (P < 0·001). However, inclusion of fodder beet had no significant effect on milk yield (P > 0·05), while increasing milk protein content and milk energy output (P ≤ 0·05). Milk energy output, as a proportion of estimated ME intake, was significantly (P < 0·001) reduced by fodder beet inclusion (0·44 vs. 0·38). Despite large increases in estimated ME intake with the inclusion of fodder beet at all levels of concentrate supplementation, milk energy output responses were small, resulting in an overall reduction in the efficiency of conversion of ME intake into milk energy output. An increased partitioning of dietary ME intake to tissue gain is suggested as the most likely explanation for the observations made.     

Effects of the proportions of high or medium digestibility grass silage and concentrates in the diet of beef cattle on liveweight gain, carcass composition and fatty acid composition of muscle

Abstract An experiment was carried out over 2 years to evaluate the effects of increasing the proportion of cereal‐based concentrates in diets containing high‐digestibility and conventional medium‐digestibility grass silages on the dry‐matter (DM) intake, liveweight gain and carcass composition of beef cattle, and to examine the effects of grazed grass and the ratio of grass silage:concentrates in the diet on the fatty acid composition of selected muscle tissues. Late‐maturing steers (n = 231) were offered diets based on high‐digestibility (HD) (0·743 digestible organic matter (DOM) in DM) or medium‐digestibility (MD) (0·643 DOM in DM) grass silages supplemented with barley/soyabean meal‐based concentrates. The concentrates constituted 0·20, 0·40, 0·60 and 0·80 of total DM of the diets, which were offered ad libitum (AL). The two diets, which contained 0·80 concentrates, were also offered at 0·80 of AL intake. A further group of fourteen animals were given the medium‐digestibility silage only for 5 months and then grazed perennial ryegrass pastures for a further 5 months (silage/pasture treatment). For the diets containing HD silage and 0·20, 0·40, 0·60 and 0·80 concentrate, and 0·80 concentrate at 0·8 of AL intake, the DM intakes were 9·4, 10·2, 10·4, 10·2 and 8·1 (s.e. 0·16) kg d^?1, respectively, and daily carcass gains were 0·67, 0·78, 0·77, 0·79 and 0·62 (s.e. 0·029) kg d^?1, respectively; for those containing MD silage and 0·20, 0·40, 0·60 and 0·80 concentrate, and 0·80 concentrate at 0·8 of AL, the DM intakes were 8·2, 9·3, 10·1, 10·1 and 8·0 (s.e. 0·16) kg d^?1, respectively, and daily carcass gains were 0·38, 0·48, 0·64, 0·77 and 0·56 (s.e. 0·029) kg d^?1 respectively. Increasing the proportion of concentrates in silage‐based diets decreased the concentration of omega‐3 (ω‐3) polyunsaturated fatty acids (PUFA) (P < 0·001) and increased the concentration of ω‐6 PUFA (P < 0·001) in muscle. Cattle on the silage/pasture treatment had the highest concentration of ω‐3 PUFA in muscle (51 g kg^?1 lipid), this value being over three times that for animals given diets containing MD silage and 0·80 concentrate in the diet. These results demonstrate the potential of HD silage made from perennial ryegrass relative to high concentrate diets. The consumption of pasture‐finished beef could make a significant contribution towards increasing the intake of ω‐3 PUFA in the human diet.     

Nitrogen metabolism and milk production in dairy cows fed semi‐restricted amounts of ryegrass–legume silage with birdsfoot trefoil (Lotus corniculatus L.) or white clover (Trifolium repens L.)

In change‐over trials, mid‐lactation dairy cows were fed concentrate‐supplemented, isonitrogenous and isofibrous perennial ryegrass–legume silage diets that satisfied energy requirements but were suboptimal with respect to metabolizable protein supply. Legumes were either birdsfoot trefoil with low levels of condensed tannins (typical for hemiboreal conditions), or white clover. Averaged over two experimental years, birdsfoot trefoil–based silage resulted in lower digestibility (P < 0·001) of dry matter (50 g kg^?1), organic matter (52 g kg^?1), neutral detergent fibre (120 g kg^?1) and nitrogen (24 g kg^?1) and lower rumen total volatile fatty acid concentration (7 mm ; P = 0·009). Milk protein yield was 36 g d^?1 higher with birdsfoot trefoil silage (P = 0·002), while raw milk yield tended to be 0·8 kg d^?1 higher (P = 0·06). Rumen ammonia concentration was similar between diets, but milk urea concentration (P < 0·001), urinary urea excretion (P = 0·002) and faecal‐N proportion (P = 0·001) were higher with birdsfoot trefoil silage. The results suggest that grass–birdsfoot trefoil silage produced in hemiboreal areas exhibits a protein‐sparing effect in dairy rations, despite a low condensed tannin content that is further diluted by companion grasses and ration concentrate proportion.     

Effect of sward surface height on the performance of ewes and Iambs continuously grazed on grass/clover and nitrogen-fertilized grass swards

In three successive years, sward height was maintained at 3, 5, 7 or 9 cm on grass swards receiving a total of 300 kg N ha^?1 in six equal monthly dressings from April, and on grass/clover swards receiving 50 kg N ha^?1 as a single dressing in early spring. From turnout in April until weaning in July, 64 ewes and their lambs (mean litter size 1·5) were continuously grazed at the four sward surface heights on the two sward types. White clover content of grass/clover swards remained low throughout the experiment ranging from 0·2 to 7·4% of the herbage mass. During the first two years, lamb gains averaged over sward types were 204, 260, 285 and 308 g d^?1 up to weaning, while in the third year gains were 238, 296, 296 and 260 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Ewes lost live weight on 3 cm swards but apart from this sward height had little effect on performance. During the autumn, weaned lambs gained — 27, 87, 147 and 167 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Sward type had only a small effect on the performance of lambs up to weaning but in the autumn, mean gains of weaned lambs were lower on grass/N swards (73 g d^?1) than on grass/clover swards (115 g d^?1). Relative to 3 cm swards, carrying capacities of 5, 7 and 9 cm swards were 0·76, 0·57 and 0·52 respectively from turnout to weaning and 0·66, 0·52 and 0·44 respectively during autumn. Grass/clover swards carried 0·67 of the ewes carried by grass/N swards from turnout to weaning and 0·51 of the live weight carried by grass/N swards during autumn. The reaction of the two sward types to sward height did not appear to differ but in the third year there was evidence of a reduction in white clover content when swards were grazed at 9 cm. The data suggest that lamb growth rates will increase as sward height increases up to 9 cm and the evidence for this was stronger with weaned lambs in autumn than with suckling lambs in spring.     

The effect of grazing by cattle on animal performance and floristic composition in Nardus-dominated swards

Four groups of six spring-calving beef cows and their calves were allocated to two replicated grazing treatments on a Nardus stricta-dominated hill pasture in the UK during six consecutive summer grazing seasons. Treatments were applied by continuous variable stocking to maintain a sward height of either 4–5 cm (short) or 6–7 cm (tall) between tussocks of N. stricta. Cows on both treatments ingested a greater proportion of N. stricta than was present in the sward. The short treatment reduced the herbage intake of cows proportionally by 0·35 (P < 0·001) and reduced herbage digestibility by 0·03 (P < 0·01) compared with the tall treatment. Cows lost 0·02 kg d^?1 live weight on the short treatment but gained 0·27 kg d^?1 live weight on the tall treatment (s.e.d. 0·077; P < 0·001). Calf liveweight gains were 0·60 and 0·86 (s.e.d. 0·034; P < 0·001) kg d^?1 for the short and tall treatments respectively. The greatest liveweight gains for both cows and calves occurred during the first 5–6 weeks of the grazing season and differences between treatments increased over the season. Cattle on the short treatment consistently grazed a higher proportion of N. stricta tillers more closely than those on the tall treatment, and the percentage cover of N. stricta declined more rapidly on the short treatment. After 6 years of grazing there was a considerably greater (P < 0·001) proportion of very small tussocks, i.e. less than 140 cm² basal area, on the short treatment than on the tall treatment. The percentage cover of broad- and fine-leaved grasses also declined, in contrast to previous studies. This may have been due to lower soil fertility on the site of the current experiment. It is concluded that high levels of utilization of N. stricta are incompatible with high levels of individual animal performance in lactating cows, although short periods in early summer may give acceptable performance. Lower levels of utilization such as that achieved on the 6–7 cm treatment resulted in moderate levels of animal performance and also gradually reduced the cover of N. stricta, although it took longer for the effect to be established.     

Effect of application timing and grass height on the nitrogen fertilizer replacement value of cattle slurry applied with a trailing‐shoe application system

This study investigated the effect of using a trailing‐shoe system to apply cattle slurry, under different conditions of grass height (low [LG]: freshly cut sward [4–5 cm height] vs. high [HG]: application delayed by 7–19 d and applied to taller grass sward [4–11 cm] height) and month of application (June vs. April), on the nitrogen fertilizer replacement value (NFRV) and apparent N recovery (ANR_S) of cattle slurry applied to grassland. NFRV was calculated using two methods: (i) NFRV_N based on the apparent recovery of slurry‐N relative to that of mineral‐N fertilizer; and (ii) NFRV_DM based on DM yield. The effect of applying slurry into HG swards, relative to LG swards, decreased the DM yield by 0·47 t ha^?1 (P ≤ 0·001), N uptake by 5 kg ha^?1 (P = 0·05), ANR_S by 0·05 kg kg^?1 (P = 0·036), NFRV_N by 0·05 kg kg^?1 (P = 0·090) and NFRV_DM by 0·11 kg kg^?1 (P < 0·001). It was concluded that the main factor causing these decreases with HG, compared with LG applications, was wheel damage affecting subsequent N uptake and growth of the taller grass sward.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK: 7. Sustainability of white clover in grass/clover swards with reduced levels of fertilizer nitrogen

The sustainability of white clover in grass/clover swards of an upland sheep system, which included silage making, was studied over 5 years for four nitrogen fertilizer rates [0 (N₀), 50 (N₅₀), 100 (N₁₀₀) and 150 (N₁₅₀) kg N ha^?1]. A common stocking rate of 6 ewes ha^?1 was used at all rates of N fertilizer with additional stocking rates at the N₀ fertilizer rate of 4 ewes ha^?1 and at the N₁₅₀ fertilizer rate of 10 ewes ha^?1. Grazed sward height was controlled, for ewes with their lambs, from spring until weaning in late summer by adjusting the proportions of the total area to be grazed in response to changes in herbage growth; surplus pasture areas were harvested for silage. Thereafter sward height was controlled on separate areas for ewes and weaned lambs. Areas of pasture continuously grazed in one year were used to make silage in the next year. For treatments N₀ and N₁₅₀, white clover stolon densities (s.e.m.) were 7670 (205·4) and 2296 (99·8) cm m^?2, growing point densities were 4459 (148·9) and 1584 (76·0) m^?2 and growing point densities per unit length of stolon were 0·71 (0·015) and 0·67 (0·026) cm^?1 respectively, while grass tiller densities were 13 765 (209·1) and 18 825 (269·9) m^?2 for treatments N₀ and N₁₅₀ respectively. White clover stolon density increased over the first year from 780 (91·7) cm m^?2 and was maintained thereafter until year 5, reaching 8234 (814·3) and 2787 (570·8) cm m^?2 for treatments N₀ and N₁₅₀ respectively. Growing point density of white clover increased on treatment N₀ from 705 (123·1) m^?2 to 2734 (260·7) m^?2 in year 5 and it returned to the initial level on treatment N₁₅₀ having peaked in the intermediate years. Stolon density of white clover was maintained when the management involved the annual interchange of continuously grazed and ensiled areas. The non‐grazing period during ensiling reduced grass tiller density during the late spring and summer, when white clover has the most competitive advantage in relation to grass. The increase in stolon length of white clover in this period appears to compensate for the loss of stolon during periods when the sward is grazed and over winter when white clover is at a competitive disadvantage in relation to grass. The implications for the management of sheep systems and the sustainability of white clover are discussed.