Effect of stand density and pruning on growth of ponderosa pines in NW Patagonia,Argentina

Due to the lack of knowledge about ponderosa pine performance under silvopastoral systems (SPS) conditions, the objective of this study was to determine the effect of stand density and pruning on the growth magnitude of ponderosa pines growing in NW Patagonia (SPS with 350 and 500 pines ha⁻¹ vs. commercial densities of 1,300 trees ha⁻¹, HPP). Individual growth rate was higher in SPS 350 trees than in SPS 500 trees, being both higher than in HPP plots, indicating a higher sensitivity of this drought resistance species to relative water availability. The higher individual growth compensated the lower amount of trees per land unit, being the whole stand growth similar or even higher in both SPS treatments than in the HPP stand. Pruning reduced diameter growth in both SPS treatments, at least until 2 years after pruning, with a more marked effect in the pruning treatment with the higher amount of extracted foliage. Carbon fixation reduction in addition to changes in carbon allocation within different plant parts after pruning could be the responsible of observed stem growth reductions. We suggest that higher growth rates in combination with frequent pruning in low density plantations can be applied to shorten the rotation period producing high quality timber in comparison with plantations managed under conventional conditions in Patagonia. Additional advantages could be associated to the lower environmental impact of low canopy cover plantations compared to high density stands.     

Effects of branch height on leaf gas exchange,branch hydraulic conductance and branch sap flux in open-grown ponderosa pine

Recent studies have shown that stomata respond to changes in hydraulic conductance of the flow path from soil to leaf. In open-grown tall trees, branches of different heights may have different hydraulic conductances because of differences in path length and growth. We determined if leaf gas exchange, branch sap flux, leaf specific hydraulic conductance, foliar carbon isotope composition (delta13C) and ratios of leaf area to sapwood area within branches were dependent on branch height (10 and 25 m) within the crowns of four open-grown ponderosa pine (Pinus ponderosa Laws.) trees. We found no difference in leaf gas exchange or leaf specific hydraulic conductance from soil to leaf between the upper and lower canopy of our study trees. Branch sap flux per unit leaf area and per unit sapwood area did not differ between the 10- and 25-m canopy positions; however, branch sap flux per unit sapwood area at the 25-m position had consistently lower values. Branches at the 25-m canopy position had lower leaf to sapwood area ratios (0.17 m2 cm-2) compared with branches at the 10-m position (0.27 m2 cm-2) (P = 0.03). Leaf specific conductance of branches in the upper crown did not differ from that in the lower crown. Other studies at our site indicate lower hydraulic conductance, sap flux, whole-tree canopy conductance and photosynthesis in old trees compared with young trees. This study suggests that height alone may not explain these differences.     

Effects of widely spaced trees and livestock grazing on understory environments in tropical savannas

The effects of widely spaced trees ofAcacia tortilis andAdansonia digitata on their understory environments were investigated in four savannas located along a gradient of increasing livestock utilization in Tsavo National Park (West), Kenya. Plant species composition and biomass and the physical and chemical properties of soils that occur below tree crowns were compared to open grasslands. The tree-crown zones at lightly and moderately grazed sites had a unique understory flora and higher plant biomass, lower temperatures and bulk densities, and higher levels of P, K, Ca and mineralizable N than their associated opengrassland zones. In the heavily grazed savanna, few differences between tree-crown and grassland zones were found. The beneficial effects of savanna trees on their understory environments appear to diminish with increasing livestock utilization.     

Changes in whole-tree water relations during ontogeny of Pinus flexilis and Pinus ponderosa in a high-elevation meadow

We measured sap flux in Pinus ponderosa Laws. and Pinus flexilis James trees in a high-elevation meadow in northern Arizona that has been invaded by conifers over the last 150 years. Sap flux and environmental data were collected from July 1 to September 1, 2000, and used to estimate leaf specific transpiration rate (El), canopy conductance (Gc) and whole-plant hydraulic conductance (Kh). Leaf area to sapwood area ratio (LA/SA) increased with increasing tree size in P. flexilis, but decreased with increasing tree size in P. ponderosa. Both Gc and Kh decreased with increasing tree size in P. flexilis, and showed no clear trends with tree size in P. ponderosa. For both species, Gc was lower in the summer dry season than in the summer rainy season, but El did not change between wet and dry summer seasons. Midday water potential (Psi(mid)) did not change across seasons for either species, whereas predawn water potential (Psi(pre)) tracked variation in soil water content across seasons. Pinus flexilis showed greater stomatal response to vapor pressure deficit (VPD) and maintained higher Psi(mid) than P. ponderosa. Both species showed greater sensitivity to VPD at high photosynthetically active radiation (PAR; > 2500 micromol m-2 s-1) than at low PAR (< 2500 micromol m-2 s-1). We conclude that the direction of change in Gc and Kh with increasing tree size differed between co-occurring Pinus species, and was influenced by changes in LA/SA. Whole-tree water use and El were similar between wet and dry summer seasons, possibly because of tight stomatal control over water loss.     

Influence of drought stress and low irradiance on plant water relations and structural constituents in needles of Pinus ponderosa seedlings

The influence of low light on tolerance to prolonged drought was tested on unshaded and shaded seedlings of ponderosa pine (Pinus ponderosa var. scopulorum Dougl. ex Laws.). Unshaded seedlings of P. ponderosa var. ponderosa were also drought stressed to compare varietal responses to drought. The maximum irradiance received by shaded seedlings was 10% of full light. Seedlings were progressively drought stressed until predawn water potentials (Psi(x)) were -5.0 MPa. Relative water content (RWC) and the reciprocal of Psi(x) were analyzed by means of an unusual application of the pressure-volume relationship for determination of RWC of the apoplast (RWC(a)), osmotic potential at full turgor (Psi(oft)), and ratio of fully turgid weight to dry weight. Major varietal differences in drought response were in RWC(a) and needle cellulose content. The shaded seedlings showed tissue damage at relative water contents < 60%, and were killed by water deficits from which unshaded seedlings recovered. Correspondingly, shaded plants had significantly higher cell volume/cell mass ratio, Psi(oft), less cellulose in needle tissue, and lower RWC(a) than unshaded plants. These differences suggest that low irradiance restricts drought adaptation in ponderosa pine.     

Restoration thinning and influence of tree size and leaf area to sapwood area ratio on water relations of Pinus ponderosa

Ponderosa pine (Pinus ponderosa Dougl. ex P. Laws) forest stand density has increased significantly over the last century (Covington et al. 1997). To understand the effect of increased intraspecific competition, tree size (height and diameter at breast height (DBH)) and leaf area to sapwood area ratio (A(L):A(S)) on water relations, we compared hydraulic conductance from soil to leaf (kl) and transpiration per unit leaf area (Q(L)) of ponderosa pine trees in an unthinned plot to trees in a thinned plot in the first and second years after thinning in a dense Arizona forest. We calculated kl and Q(L) based on whole- tree sap flux measured with heat dissipation sensors. Thinning increased tree predawn water potential within two weeks of treatment. Effects of thinning on kl and Q(L) depended on DBH, A(L):A(S) and drought severity. During severe drought in the first growing season after thinning, kl and Q(L) of trees with low A(L):A(S) (160-250 mm DBH; 9-11 m height) were lower in the thinned plot than the unthinned plot, suggesting a reduction in stomatal conductance (g(s)) or reduced sapwood specific conductivity (K(S)), or both, in response to thinning. In contrast kl and Q(L) were similar in the thinned plot and unthinned plot for trees with high A(L):A(S) (260-360 mm DBH; 13-16 m height). During non-drought periods, kl and Q(L) were greater in the thinned plot than in the unthinned plot for all but the largest trees. Contrary to previous studies of ponderosa pine, A(L):A(S) was positively correlated with tree height and DBH. Furthermore, kl and Q(L) showed a weak negative correlation with tree height and a strong negative correlation with A(S) and thus A(L):A(S) in both the thinned and unthinned plots, suggesting that trees with high A(L):A(S) had lower g(s). Our results highlight the important influence of stand competitive environment on tree-size-related variation in A(L):A(S) and the roles of A(L):A(S) and drought on whole-tree water relations in response to thinning.     

Growth of widely spaced trees. A case study from young agroforestry plantations in France

Pessimistic forecasts often suggest that widely spaced trees enjoying free growth (no competition with other trees) will fail to provide high quality timber. This challenges the temperate agroforestry practice of planting widely spaced trees to produce high quality timber. We analyse tree growth data from recent temperate agroforestry plantations aged three to eight years, featuring low tree plantation densities (50 to 400 stems ha−1), the association of trees with intercrops (silvoarable systems) or animals (silvopastoral systems), and the use of plastic shelter tubes to protect trees (1.2 to 2.5 m high) and avoid damage by cattle or sheep in pastures or facilitate maintenance in silvoarable systems. The plantations are located in climates ranging from Mediterranean, dry central temperate plains, cold and wet central uplands to mild oceanic areas. Some plantations included a forestry control (high density of plantation, no tree shelter, no intercropping nor grazing). Trees were evaluated for height and diameter growth and stem form (straightness and absence of knots). Trees in most agroforestry plots grew satisfactorily, often faster than in forestry control plots. In some experimental plots, average annual height increments as high as 1 m and diameter increments as high as 1.5 cm were observed. Few agroforestry plantations were unsuccessful, and the reasons for the failures are discussed: animal damage in silvopastoral plots, but also a wrong choice of tree species unsuitable for local soil and climate characteristics. From these early results we can formulate some guidelines for designing future agroforestry plantations in temperate climates, concerning tree density, use of tree shelters and care required for widely spaced trees. This revised version was published online in June 2006 with corrections to the Cover Date.     

Branch characteristics of widely spaced Douglas-fir in south-western Germany: Comparisons of modelling approaches and geographic regions

Models of Douglas-fir branch and whorl characteristics were developed from contrasting spacing experiments in southwest Germany. The dataset was based on 100 young (20–30 years old), unpruned and partially pruned trees from a 100, 200, and 1200 stems ha⁻¹ spacing experiment on Douglas-fir that was replicated 3 times across the region. The material was used to predict (1) the number of branches whorl⁻¹, (2) branch angle, (3) status (living/dead) of the branches within the living crown, (4) maximum branch diameter whorl⁻¹, and (5) relative diameter of branches within a whorl. For each of these models (except branch status), both a linear and nonlinear, generalised hierarchical mixed effects equation was developed. The comparison of the linear and nonlinear approaches showed that both had a relatively similar level of bias, but the nonlinear equations generally performed better (reduction in mean absolute error of 1.1–69.5%). Overall, individual branch and tree properties were sufficient to give logical and precise predictions of the branch characteristics for the models across the range of sampled stand densities. In addition, the models showed a similar behaviour compared to models on Douglas-fir crown structure from the Pacific Northwest, USA. This suggests that the allometric relationship between tree size and branch characteristics for a given species may be relatively consistent across regions, even ones with highly contrasting growing conditions like in this study. The models performed well across a range of stand conditions and now will be further integrated into an individual tree growth and yield simulations system.     

Effect of tree pruning and pruning application to trees on nitrogen fixation by Leucaena and Gliricidia

A microplot study was conducted with Leucaena leucocephala and Gliricidia sepium to determine the effect of pruning the trees and applying the prunings to the soil (as either surface mulch or soil-incorporated green manure) on N₂ fixation by the trees. N₂ fixation was estimated by the ¹⁵N dilution method, and the non-N₂-fixing species Senna siamea was used as a reference species. Number and dry weight of nodules were reduced with applications of prunings in Leucaena, and increased in Gliricidia. For both species, the lowest rate of N₂ fixation (47%) occurred under green manure incorporation after pruning, and the highest (66%) with mulching and no-pruning. Incorporation of green manure into soil significantly depressed N₂ fixation by 19.1% in Gliricidia and 20.6% in Leucaena compared to no-pruning treatment. The deleterious effect of incorporation of prunings into soil also affected the amount of N₂ fixed. During the nine months following pruning, green manure incorporation steadily decreased the percentage of N₂ fixed, whereas with no-pruning, pruning only and pruning with mulching, N₂ fixation increased with time. Reduction of N₂ fixation in pruned plants could imply that they would compete for N with interplanted food crops, causing lower N use efficiency by the latter. Despite lower dry-matter and N yields of Leucaena as compared with Gliricidia, its biological N₂ fixation amounted to 73% of its N requirement as opposed to the corresponding figures of 50% for Gliricidia. This revised version was published online in June 2006 with corrections to the Cover Date.     

Effect of pruning intensity and season on Diplodia pinea infection of Pinus radiata stem through pruning wounds

In a five-year-old Pinus radiata stand, trees were pruned to remove 50%, 40% or 25% of the crown and pruning stubs inoculated with Diplodia pinea. Infection occurred in 65–68% of the intensely pruned trees some of which also developed crown wilting and died, but in lightly pruned trees (25% crown removal) only 8% of the trees were infected and no mortality occurred. Pruning and inoculation every two weeks for a whole year cycle showed an infection peak in summer, but extremely low infection level during the rest of the year. A vertical gradient of resistance of infection was apparent along the stem with the basal part being extremely resistant, becoming more susceptible towards the top. Within limits infection also increased with increasing stub diameter.     

Bole girdling affects metabolic properties and root, trunk and branch hydraulics of young ponderosa pine trees

Effects of trunk girdling on seasonal patterns of xylem water status, water transport and woody tissue metabolic properties were investigated in ponderosa pine (Pinus ponderosa Dougl. ex P. Laws.) trees. At the onset of summer, there was a sharp decrease in stomatal conductance (g(s)) in girdled trees followed by a full recovery after the first major rainfall in September. Eliminating the root as a carbohydrate sink by girdling induced a rapid reversible reduction in g(s). Respiratory potential (a laboratory measure of tissue-level respiration) increased above the girdle (branches and upper trunk) and decreased below the girdle (lower trunk and roots) relative to control trees during the growing season, but the effect was reversed after the first major rainfall. The increase in branch respiratory potential induced by girdling suggests that the decrease in g(s) was caused by the accumulation of carbohydrates above the girdle, which is consistent with an observed increase in leaf mass per area in the girdled trees. Trunk girdling did not affect native xylem embolism or xylem conductivity. Both treated and control trunks experienced loss of xylem conductivity ranging from 10% in spring to 30% in summer. Girdling reduced xylem growth and sapwood to leaf area ratio, which in turn reduced branch leaf specific conductivity (LSC). The girdling-induced reductions in g(s) and transpiration were associated with a decrease in leaf hydraulic conductance. Two years after girdling, when root-to-shoot phloem continuity had been restored, girdled trees had a reduced density of new wood, which increased xylem conductivity and whole-tree LSC, but also vulnerability to embolism.     

Age-related changes in ecosystem structure and function and effects on water and carbon exchange in ponderosa pine

As forests age, their structure and productivity change, yet in some cases, annual rates of water loss remain unchanged. To identify mechanisms that might explain such observations, and to determine if widely different age classes of forests differ functionally, we examined young (Y, approximately 25 years), mature (M, approximately 90 years) and old (O, approximately 250 years) ponderosa pine (Pinus ponderosa Dougl. ex P. Laws.) stands growing in a drought-prone region of central Oregon. Although the stands differed in tree leaf area index (LAIT) (Y = 0.9, M = 2.8, O = 2.1), cumulative tree transpiration measured by sap flow did not differ substantially during the growing season (100-112 mm). Yet when water was readily available, transpiration per unit leaf area of the youngest trees was about three times that of M trees and five times that of O trees. These patterns resulted from a nearly sixfold difference in leaf specific conductance (KL) between the youngest and oldest trees. At the time of maximum transpiration in the Y stand in May-June, gross carbon uptake (gross ecosystem production, GEP) was similar for Y and O stands despite an almost twofold difference in stand leaf area index (LAIS). However, the higher rate of water use by Y trees was not sustainable in the drought-prone environment, and between spring and late summer, KL of Y trees declined fivefold compared with a nearly twofold decline for M trees and a < 30% reduction in O trees. Because the Y stand contained a significant shrub understory and more exposed soil, there was no appreciable difference in mean daily latent energy fluxes between the Y stand and the older stands as measured by the eddy-covariance technique. These patterns resulted in 60 to 85% higher seasonal GEP and 55 to 65% higher water-use efficiency at the M and O stands compared with the Y stand.     

修枝对茶条槭产叶量的影响

茶条槭 (Acer ginnala Marim)产自我国东北、内蒙及华北山地 ,为阳性树种 ,喜湿润土壤 ,耐寒抗风雪。其叶部含有大量的没食子酸 ,每公斤干叶可提取 70～ 80 g没食子酸。没食子酸在医药、化工、食品、轻工等方面用途广泛 ,经济价值极大 ,仅黑龙江省每年就计划生产 6 0 0 t,需要茶条槭干叶 70 0 0 t。随着对外贸易的不断扩大 ,对其需求将越来越大 ,因此 ,提高茶条槭的产量 ,尤其是提高其叶产量显得尤为重要和迫切 ,本试验的目的就是探讨不同的修枝强度对其叶产量的影响。1　试验地概况试验地设在黑龙江省苇河林业局冲河苗圃 ,地理位置为 12 8…     

Changes in understory production for three prescribed burns of different ages in ponderosa pine

The general lack of resource response information severely limits economic evaluation of prescribed burning in most forest types. This paper present changes in understory production at three sites on basalt soils following prescribed burning in ponderosa pine (Pinus ponderosa Laws.) in Arizona. The sites were burned during the fall 2, 5, and 7 years before sampling in 1981. Regression equations were developed to predict production from plant basal area for four common grass species; production of remaining herbaceous vegetation was measured by harvesting. Based on the results of our study and other studies of fall prescribed burning on volcanic soils in Arizona ponderosa pine, understory production response appears to be variable for 1–2 years following burning. Herbage production exhibits no change or an increase; forage production exhibits no change or a decrease. Studies of understory response more than 2 years following both wildfire and prescribed burning in Arizona ponderosa pine, however, show a general trend toward increased production. Thus, prescribed burning in southwestern pine on basalt soils produces long-term benefits in increased understory production, particularly in pole stands, which dominate much of the region.     

Pine needle consumption and bark stripping by sheep grazing annual pastures in young stands of widely spaced Pinus radiata and P. pinaster

The consumption of the pine needles and the stripping of bark by sheep grazing annual pastures in three open stands of P. radiata and one of P. pinaster were measured. The P. radiata were four years old and the densities were 250, 500 and 750 trees ha^-1. The P. pinaster included two, three and four year-old trees and the density was 440 trees ha^–1.Needles were eaten immediately after the sheep were admitted and this continued throughout the year. Within ten weeks of the start of winter grazing, sheep stocked at 7–10 ha^–1 had eaten about half of the accessible needles on both species where densities were less than500 trees ha^–1. The estimated consumption was about 260 g sheep^–1 day^–1 for the P. radiata. Bark stripping then commenced and continued throughout spring and summer on most plots. In some plots at low tree density, bark was stripped from more than 35% of the trees. In denser stands, needle consumption per tree was less and bark was stripped from less than 5% of the trees.Bark damage was most common on the smaller trees, and severity of stripping was negatively correlated with tree size. Of all the trees with bark damage, 22% of the P. pinaster and 14% of the P. radiata had more than half of the stem circumference stripped. A number of these will probably suffer both stem distortion and growth retardation, and some may die.Methods of reducing the incidence of bark damage are discussed.     

Effect of pruning frequency and pruning height on the biomass production of <Emphasis Type="Italic">Tithonia diversifolia</Emphasis> (Hemsl) A. Gray

Addition of tree or shrub prunings through alley cropping or biomass transfer systems have contributed to sustainable land-use systems in the tropics. Long term productivity of biomass transfer systems require shrub or tree species that coppice after cutting to provide sufficient plant nutrients. The effect of pruning frequency and cutting height on the biomass production of Tithonia diversifolia was studied to provide information for managing hedges. Results showed that height of cutting, pruning frequency, and their interactions significantly affected dry matter production of T. diversifolia. The results also showed that a significantly higher biomass production could be produced when Tithonia was pruned at long time intervals. Pruning height was also of importance in the harvesting of Tithonia biomass and it was evident that dry matter production was highest when Tithonia was pruned bimonthly at 50 cm height. With bi-monthly pruning frequency, dry matter production could be as high as 7.2 t ha⁻¹ yr⁻¹ which might be a sufficient biomass to improve soil productivity in biomass transfer systems.     

修剪强度对野生杜鹃生长开花的影响

对野生杜鹃采取不同修剪强度的对比试验,结果表明:重回缩使野生杜鹃枝条平均生长量能达到40 cm,有利于枝条更新复壮;摘花使野生杜鹃抽出枝条达603条/m2,摘花有利于枝条抽出;短截枝条生长量5 843 cm/m2,短截能有效促进枝条生长量增加;短截使杜鹃花芽72.8个/m2,中回缩和重回缩花芽为0;短截杜鹃开花68.5朵/m2,开花数量最多。     

Effect of pruning on nitrogen dynamics within crowns of Pinus radiata

Stem injection of (15)N-labeled ammonium sulfate was used to determine effects of pruning on canopy nitrogen dynamics in open-grown Pinus radiata D. Don in New Zealand. Trees were planted in July 1990 and the isotope introduced in December 1994. Tree crowns were divided into three zones: base section, from which branches of pruned trees were removed; mid section, between the pruned zone and the height of the trees at the start of the year in which they were pruned; and top section, which grew predominantly after the isotope was applied. Pruning removed 32% of the green crown length, representing 75% of foliage biomass. Needles were sampled from each region of the crown until July 1996. Branch growth was used to predict foliage biomass for each sampling occasion. Approximately 45% of the applied isotope was recovered from needles sampled in December 1994 (1 week after application and immediately before pruning), two-thirds of which occurred in needles in the base section. Thereafter, changes in isotope content of needles in the base section of unpruned trees largely reflected foliage biomass fluctuations and dilution of the isotope by continued uptake from the unlabeled soil nitrogen pool. Recovery of isotope in needles from the mid-crown section increased by 58 and 86% from December 1994 to July 1995 in control and pruned trees, respectively. Within this crown section, there was evidence of isotope translocation from old to new needles, with both isotope dilution and efflux observed in the needle cohorts that had been present at the time the isotope was applied. Therefore, isotope dynamics did not reflect the dynamics of the total nitrogen pool in the mid-crown section. By July 1996, a small proportion of the applied isotope was recovered from the new foliage formed in the top section of the crown. Within the top section, isotope dynamics closely matched total nitrogen fluxes. Pruning the lower crown did not affect nitrogen dynamics elsewhere in the crown for the following 18 months.     

Effect of pruning Acacia mangium on growth,form and heart rot

Small volumes of timber are now being produced from Acacia mangium plantations in Indonesia. These trees require pruning and thinning to increase the strength and appearance of the wood. However, cut surfaces from pruning are potential infection courts for the entry of decay-causing fungi like heart rot. This study investigated the effects of pruning on stem form and the incidence of heart rot in an 18-month-old plantation of Acacia mangium in South Sumatra. The objectives were to assess whether pruning is associated with an increase in the incidence of heart rot and whether form pruning compared to lift pruning reduced the incidence of heart rot and improved stem form. Form pruning removed 25% of leaf area by removing large branches and those subtending a narrow angle with the stem up to 3 m height, and lift pruning removed 25% of crown length from below. Trees in these treatments were singled before pruning. The third treatment, a control, was not singled and was used to assess base levels of heart rot.No significant difference in diameter increment between the two pruning treatments was found. There was strong evidence that form pruning was associated with better form 18 months after treatment. Trees in this treatment had a reduced number of branches >30 mm diameter and improved stem straightness (reduced kink). Lift pruning reduced average branch size but did not improve stem straightness. No heart rot was detected in any treatment.The results showed that form pruning is likely to have positive benefits on stem straightness and is likely to be effective to any selected pruning height. However a subsequent lift pruning is still considered a requirement. While wounds created from pruning and singling are assumed to have a large impact on the incidence of heart rot, this may not be an issue unless there is a sufficient source of fungi present in the environment to invade the wounds.     

城郊绿化整形树木修剪技术初探

整形树,作为城郊绿化美化的一种表现形式,随着人们欣赏水平的不断提高,在乡村特别是城郊绿化中越来越得到了广泛应用。实践证明,整形树在众多的园林植物中有着不可替代的独特功能和效果,深受人们的喜爱和普遍赞赏。