Effects of active pasture,teak (Tectona grandis) and mixed native plantations on soil chemistry in Costa Rica

In this study soil samples were taken from the O/A and B horizons of undisturbed forest, active pasture, and 8- to 12-year-old teak and mixed native plantations. Samples were analyzed for K, Ca, Mg, soil organic carbon, pH, exchangeable acidity, bulk density, and compared with a fertility equation. Bulk density was significantly lower in the undisturbed forest than other land uses, suggesting that after approximately 10 years of growth neither plantation lowered bulk density significantly from that of the active pasture. Teak plantations had significantly higher Mg and K (B horizon) and Ca (O/A horizon) concentrations than the undisturbed forest. This trend suggests that exchangeable base concentrations increase when land use changes from undisturbed forest to pasture, then pasture to plantation, with the most pronounced effect of this in teak plantations exhibiting more high fertility plots than other land uses. Soil organic carbon concentration was similar for all land uses except for a significantly lower concentration in teak plantations than in active pasture (O/A horizons). These results suggest that teak plantations may be advantageous for increasing soil fertility but, with respect to restoration of undisturbed forest conditions, present significant deviations in soil chemistry. Options for improved plantations soil management are discussed.     

Competitiveness, systems of innovation and the learning economy: the forest sector in Costa Rica

The different sectors of the economy, and the forest sector in particular, must create innovations in order to survive and to be competitive. Scientific knowledge should be introduced and institutionalized into the forest processes, but also knowledge derived from other sources such as user producer relationships and others should be incorporated. A new concept is emerging that considers that the economy should learn not only from the new scientific knowledge but also from the rest of the processes. This concept is National Systems of Innovation (created in 1985 in the Scandinavian countries), however, here the concept is broadened of a Sustainable System of Innovation. This approach of sustainable systems of innovation (SSI), considers that the sources for innovation towards development, particularly sustainable development, are formal organisations (universities, research centres, or so), the productive structure (the economy), the institutional set-up (patterns of behaviour), the economy–ecology relationships (potential and limitations of the ecosystem for production) and the policies of the sector. Each one of these factors interact and feed each other. They cannot be seen independently nor static; on the contrary, they are dynamic and evolutionary, without any doubt including the ecosystem. Then, the process of learning from all these sources, which is the base for SSI, may produce ‘greener’ innovations, greener technologies and greener institutions. Innovations related to forest services from Costa Rica will illustrate these ideas.     

Restoring abandoned pasture land with native tree species in Costa Rica: Effects of exotic grass competition and light

Understanding the early establishment requirements and performance of tropical tree seedlings is essential to ensuring the success of restoration plantings. This study was designed to characterize growth and light requirements of six common neotropical tree species Pseudosamanea guachapele (Fabaceae), Tabebuia impetiginosa (Bignoniaceae), Ceiba pentandra (Bombacaceae), Pachira quinata (Bombacaceae), Dalbergia retusa (Fabaceae), and Tabebuia rosea (Bignoniaceae) in an abandoned pasture under contrasting light environments and grass competition. Field studies were conducted in the pastures of the Santa Ana Conservation Center in Costa Rica. Two differing grass competition sites were selected, one dominated by a tall grass, Hyparrhenia rufa and another dominated by a short grass, Cynodon mlenfluensis. Three light treatments were created (2, 37 and 100% light) using either neutral shade cloth (2 and 37%) or no shade cloth (100%). Growth characteristics and biomass partitioning of the seedlings were measured. Species differed in their relative growth rates (RGRs). The light × species interaction was significant at both sites. While all species had similar performance under 100% light on both short grass and tall grass sites, species growth differences were evident under 37 and 2% light levels. For example, at the short grass site, T. rosea had higher RGR than D. retusa and P. quinata under 2% light whereas both Tabebuia species performed better than other species at the tall grass site. The general trend was to increase root mass ratio and decrease leaf mass ratio with increasing levels of light. As an initial step for restoring abandoned pasture lands we recommend using all these species in direct and moderate light conditions. Incorporating all species will create a more heterogeneous environment. Choosing light demanding species that can tolerate grass competition may help ensure success in the early stage of restoration.     

Growth of native forest species planted on abandoned pasture land in Costa Rica

Many forest plantations in the humid neo-tropics are established on degraded soils in abandoned pasture land and, with some exceptions, the species planted have not grown successfully. Studies of adaptability and growth under these conditions are scarce, particularly for native species. In this paper we present data on growth and tree form at 3 years of age for 11 species planted in abandoned pastures. The research plantation was established at La Selva Biological Station, in the Atlantic lowlands of Costa Rica (10° 26′N, 83° 59′W). Survival, diameter at breast height (DBH), total height, basal area, volume index and tree form were used to evaluate the species performance. After 3 years, Vochysia guatemalensis displayed a survival significantly higher (98%) than that of the other species. The lowest survival was 75% for Pinus tecunumanii. There were significant differences in growth measures among species. Acacia mangium exhibited significantly higher growth rates than the other species. Although this species grew rapidly, it did not develop straight single stems. About 14% of trees of A. mangium had bifurcation below DBH and more than 50% showed multiple axes or branches as large in diameter as the principal axis. The native species with the highest growth rate was Vochysia guatemalensis (DBH 12.7 cm, total height 7.5 m, volume index 55 m³ ha⁻¹ at 3 years of age). Vochysia ferruginea exhibited a slightly lower growth rate. Both Vochysia species were ranked highly with regard to tree form, with more than 80% straight single stemmed trees. The lowest growth rates were exhibited by Pentaclethra macroloba, Pithecellobium macradenium and Virola koschnyi; however, the latter two species formed straight single terms. Pentaclethra macroloba and Inga edulis failed to form straight single stems. In general, the growth rates of the species studied were high in comparison with results reported from other tree plantations in the tropics. In spite of this, it is not prudent to draw final conclusions on the basis of these 3 year measurements.     

A comparison of agrisilvicultural systems with plantation forestry in the Atlantic Lowlands of Costa Rica

Survival and growth data (ages 0–5 years) are presented for two timber species (Acacia mangium, Cordia alliodora) planted in monocultures or in association with a sequence of agricultural cropsZea mays, Zingiber officinale andEugenia stipitata (a fruit shrub) in the tropical humid Caribbean lowlands of Costa Rica. Average annual height and diameter growth rates were 3.2 m and 4.0 cm (C. alliodora), 3.5 m and 3.8 cm (A. mangium).C. alliodora associated with crops gave the greatest productivity with an average total stem volume increment of 19 m³ ha^–1 yr^–1. Root rot ofA. mangium (mainlyRosellinia sp.), leading to tree mortality, was greater in pure plots compared to associated plots.A. mangium can not be recommended for similar sites because of this problem.     

The ecological sustainability of tropical forest management: evaluation of the national forest management standards of Costa Rica and Nicaragua, with emphasis on the need for adaptive management

From a conceptual point of view, national forest management standards in Latin American countries have progressed significantly in recent years. Examples include the Costa Rican Standards and Procedures for Sustainable Forest Management and Certification, developed by the National Commission for Forest Certification and in Nicaragua, the National Institute of Forestry proposal of principles, criteria and indicators for sustainable forest management. In line with general approaches worldwide, these national standards primarily focus on the fulfillment of sound forest practice. There is comparatively little emphasis on the assessment of management outcomes or changes in key components of the eco- and social-systems that result from management impacts. Essentially, there is little emphasis on adaptive management, though arguments that management cannot be sustainable if it is not adaptive are persuasive. This study sought to contribute to the development of standards that include elements for adaptive management that define, communicate and evaluate sustainable forest management in Costa Rica and Nicaragua. Elements from the national standards and the CIFOR generic C&I template (predominantly focused on forest management outcomes) were used as a starting point. The basic research process consisted of three phases of evaluation (in-office, desk and field). The evaluations were carried out by multidisciplinary, international groups of experts in forest ecology, management and policy. This study demonstrated the value of forums and workshops that facilitate exchange between forest scientists and policymakers; the innovation and application of a practical, applicable and scientifically based methodology for developing national level C&I; and acceptance of this methodology by key players in the fields of forest management and policy. These experiences and the resulting proposals of C&I for the evaluation of ecologically sustainable forest management are expected to be used as points of reference for future development of forest policy in Costa Rica and Nicaragua, and to contribute to the overall understanding of C&I development processes in the region.     

Silvicultural and economic aspects of pure and mixed native tree species plantations on degraded pasturelands in humid Costa Rica

Reforestation of degraded land in tropical regions provides one means of restoring ecosystems and improving rural livelihoods. Most plantations in humid tropical regions are established in pure plots, including few species of high commercial value, generally exotics. The present study compares growth and economic viability of 15–16 year-old trees of native species in pure and mixed plantations on degraded pasturelands at La Selva Biological Station, Costa Rica. The species with the best growth were Vochysia guatemalensis, Virola koschnyi, Jacaranda copaia, Terminalia amazonia, and Hieronyma alchorneoides. The mixed plantations performed better than pure plantations for all growth variables considered, including height, diameter at breast height, volume, and aboveground biomass. Mixed plantations outperformed pure plantations economically, with Net Present Value (NPV) of 1,124 to1,124 to 8,155/ha and Internal Rate of Return (IRR) of 7.7–15.6% depending on the species mixture. The most profitable pure plantations were those of Vochysia guatemalensis, with NPV and IRR of $6,035/ha and 14.3%, respectively; Hieronyma alchorneoides ($6,035/ha and 14.3%, respectively; Hieronyma alchorneoides (2,654 and 10.8%); and Virola koschnyi ($1,906/ha and 9.22%). These are the estimated benefits that would be accrued at harvest. These projections are realistic since the decision to harvest is generally mandated not just by profit maximization but by economic necessity, thus many small- to medium-scale farmers in Costa Rica harvest plantations prior to the completion of the rotation length. Mixed plantation may be the preferred system for reforestation with native species designed for timber production or carbon sequestration because this system is more economically viable and productive than pure plantations.     

Survival, growth, timber productivity and site index of Cordia alliodora in forestry and agroforestry systems

Matching tree species to appropriate site conditions and stand management is crucial for sound agroforestry production. In this study, survival, growth and site index for laurel (Cordia alliodora (Ruiz and Pavón) Oken.) were measured between 1987–1999 in two forestry (line plantings and pure plantations) and four agroforestry systems (taungya and three laurel – cacao (Theobroma cacao L.) systems) in the lowland humid tropics of Costa Rica and Panama. Mortality ranged between 2 and 52% at age nine years. Poor drainage, flooding and high water tables resulted in low laurel survival on some sites. At age 5, laurel site index was 21 m in cacao – plantain (Musa AAB) – laurel associations (CLP), but only 15 m in line plantings. Diameter at breast height at that age was 28.5 cm in CLP and only 15.6 cm in pure plantations. Laurel growth and site index were high when planted in association with intensively managed crops (e.g., cacao, plantain or taungya sequences).     

Deforestation and forest restoration in Guanacaste,Costa Rica: Putting conservation policies in context

Traditionally Land Use Cover Change (LUCC) studies have focused on processes of negative land-cover change, primarily deforestation, partially because examples of positive land-cover change were not common. During the last two decades an outstanding tropical dry forest restoration process had taken place in the province of Guanacaste, Costa Rica, which has given us a unique opportunity to study how and why tropical dry forest regrowth occurs. The purpose of this paper is to undertake a retrospective analysis of the social dynamics of forest deforestation and restoration in Guanacaste from 1960 to 2005. Hence we investigate how structural drivers shape patterns of forest-cover change and examine how the role that Costa Rica's conservation policies had played in promoting forest restoration. Our study combined analyses of socioeconomic data and satellite images of forest cover. We showed that forest regrowth observed in Guanacaste after the 1980s was the result of multiple socioeconomic factors. Our results indicate that the degree of incentive provided by conservation policies such as Payment for Environmental Services are not enough to ensure that Guanacaste's forest will be protected against the potentially negative impacts of future socioeconomic changes. The findings from our analysis can assist decision-makers and managers in other regions to understand how social, economic and political dynamics impact the effectiveness of forest conservation efforts.     

Diversity of naturally-regenerated native woody species in forest plantations in the Ethiopian highlands

Woody species diversity and ground layervegetation cover were studied in plantations ofPinus patula, Cupressus lusitanica,Grevillea robusta, and Juniperusprocera, and in surrounding natural forests inWondo Genet, Ethiopia. In the understory of theplantations, a total of 53 naturallyregenerated tree and shrub species belonging to31 families were recorded. Important indigenoustimber species were also represented. Treesaccounted for 72% of all naturally-regeneratedwoody plant species. On the average, 78% ofthe woody species found in plantations werealso found in the natural forest. There was nosignificant variation in understory woodyspecies richness among plantations. C.lusitanica plantations had a significantlyhigher abundance of woody species than those ofP. patula, but the difference wasnot significant in relation to J. procera,G. robusta and the adjacent natural forest.The herbaceous ground cover percentage in G. robusta and P. patula stands wasconsiderably higher than in C. lusitanicaand J. procera stands. Relative toplantations of other species, plantations ofthe native tree species J. procera hadhigher understory species richness, mediumwoody species abundance and low ground cover.Although there was a marked variation inoverstory plantation density, its effect onunderstory species richness and abundance wasnot significant. The density ofnaturally-regenerated woody species inplantations was over three times the usualplanting density in Ethiopia, indicating a highpotential of forest plantations for restoringthe natural forest ecosystems on degraded landsat a comparatively low cost.     

Biomass and nutrient accumulation in pure and mixed plantations of indigenous tree species grown on poor soils in the humid tropics of Costa Rica

Aboveground biomass and nutrients and soil chemical characteristics were examined in young plantations of four indigenous tree species: Hieronyma alchorneoides, Vochysia ferruginea, Pithecellobium elegans, and Genipa americana, growing in mixed and pure stands at La Selva Biological Station, Costa Rica. Total tree biomass production rates ranged from about 5.2 Mg ha⁻¹ year⁻¹ for G. americana to 10.3 Mg ha⁻¹ year⁻¹ for H. alchorneoides pure stands, and for the species mixture it was about 8.9 Mg ha⁻¹ year⁻¹. Branches and foliage formed 25–35% of total tree biomass but they represented about 50% of total tree nutrients. H. alchorneoides, the four species mixture, and P. elegans had the greatest accumulations of total aboveground nutrients per hectare. The importance of the plantation floor as a nutrient compartment varied temporally. When forest floor litter biomass was at its peak, plantation floor litter N, Ca, and Mg were roughly equal to, or greater than stem nutrients for all species except for P. elegans. For P. elegans, the plantation floor consistently represented a very low proportion of total aboveground nutrients. G. americana and V. ferruginea trees showed 55–60% less biomass accumulation in mixed than in pure stands while H. alchorneoides and P. elegans trees grew 40–50% more rapidly in mixture. P. elegans foliage had 60% lower Ca but higher P concentrations in mixed than in pure stands, and G. americana had higher foliar Mg in mixed than in pure stands. V. ferruginea stands had the highest concentrations of soil Ca, Mg, and organic matter, particularly in the top layers. Relative to pure plantations, soil nutrient concentrations in mixed plantations were intermediate for N, P, and K, but lower for Ca and Mg. The results of this study can be used in the selection of tree species and harvest designs to favor productivity and nutrient conservation.     

Reducing financial risk in agroforestry planning: a case study in Costa Rica

We used fluctuations in net income from alternative cropping systems to assess the financial risk associated with an agroforestry system. Mean-variance analysis was used to derive a set of minimum-risk farm plans for a 15-hectare farm in Costa Rica. Monocultural coffee production provided the highest expected net income, but also had the greatest economic risk. As risk was reduced, the optimal agroforestry system diversified to include other cropping systems in addition to the coffee monoculture. Risk aversion was, however, accompanied by significant reductions in expected net income for the cropping systems studied. The inclusion of additional cropping systems whose net incomes are negatively correlated with the systems considered here could help reduce the economic risk facing rural agriculturalists in this region.This research was supported by the Utah Agricultural Experiment Station, Utah State University, Logan, Utah 84322-4845. Approved as journal paper no. 4211.     

Allelopathic effects of eucalyptus and the establishment of mixed stands of eucalyptus and native species

Allelopathic effects of eucalyptus are widely reported and are considered the major factor limiting the establishment of native species in eucalyptus forests. However, reports of the allelopathic effects of eucalyptus are mostly based on laboratory bioassay, not on field trials. In the present study, we conducted field trials to determine the allelopathic effects of eucalyptus and to develop strategies for establishment of mixed stands of native species and eucalyptus. In the field, seed germination and seedling survivorship of native species were determined in the presence and absence of eucalyptus leaf litter or living roots. In the seed germination experiment, seed germination rate of Delonix regia was higher than for Elaeocarpus sylvestris and Tsoongiodendron odorum. Seed germination rates of E. sylvestris and T. odorum were low in both the presence and absence of eucalyptus treatments. The germination rate of D. regia was significantly inhibited by all eucalyptus treatments, but only during the early period. In the seedling establishment experiment, seedling survivorship of E. sylvestris and Michelia macclurel was not inhibited by any treatments, but the seedling survivorship of Schima superba was significantly inhibited by eucalyptus litter addition alone. Seedling height of S. superba and M. macclurel was significantly suppressed when eucalyptus roots were present in treatments, but the seedling height of E. sylvestris was only significantly suppressed by the treatment of roots alone. We propose two strategies for establishment of mixed stands of native species and eucalyptus: (1) direct seed-sowing is effective only for those species (e.g. D. regia) with high germination rates in natural conditions; (2) seedling transplants are more efficient approach for establishing species (e.g. E. sylvestris) with low germination rates. We suggest that mixed plantations of eucalyptus and E. sylvestris can be established by transplanting seedlings of E. sylvestris into eucalyptus plantations.     

Carbon accumulation in the biomass and soil of different aged secondary forests in the humid tropics of Costa Rica

Efforts are needed in order to increase confidence for carbon accounts in the land use sector, especially in tropical forest ecosystems that often need to turn to default values given the lack of precise and reliable site specific data to quantify their carbon sequestration and storage capacity. The aim of this study was then to estimate biomass and carbon accumulation in young secondary forests, from 4 and up to 20 years of age, as well as its distribution among the different pools (tree including roots, herbaceous understory, dead wood, litter and soil), in humid tropical forests of Costa Rica. Carbon fraction for the different pools and tree components (stem, branches, leaves and roots) was estimated and varies between 37.3% (±3.3) and 50.3% (±2.9). Average carbon content in the soil was 4.1% (±2.1). Average forest plant biomass was 82.2 (±47.9) Mg ha⁻¹ and the mean annual increment for carbon in the biomass was 4.2 Mg ha⁻¹ yr⁻¹. Approximately 65.2% of total biomass was found in the aboveground tree components, while 14.2% was found in structural roots and the rest in the herbaceous vegetation and necromass. Carbon in the soil increased by 1.1 Mg ha⁻¹ yr⁻¹. Total stored carbon in the forest was 180.4 Mg ha⁻¹ at the age of 20 years. In these forests, most of the carbon (51-83%) was stored in the soil. Models selected to estimate biomass and carbon in trees as predicted by basal area had R² adjustments above 95%. Results from this study were then compared with those obtained for a variety of secondary and primary forests in different Latin-American tropical ecosystems and in tree plantations in the same study area.     

Woody understory plant diversity in pure and mixed native tree plantations at La Selva Biological Station,Costa Rica

Tree plantations can be an important tool for restoration of abandoned pasturelands in the tropics. Plantations can help speed up secondary forest succession by improving soil conditions, attracting seed-dispersal agents, and providing shade necessary for understory growth. In this study, abundance and richness of understory regeneration was measured in three native tree plantations 15–16 years of age at La Selva Biological Station in the Costa Rican Caribbean lowlands. Each plantation contained tree species in pure plots, a mixture of the species, and natural regeneration plots (no trees planted). The greatest abundance of regeneration was found in the understory of pure plots of Jacaranda copaia (Aubl.) D.Don., Vochysia guatemalensis Donn.Sm., Dipteryx panamensis Benth, Vochysia ferruginea Mart., and in two mixed stands, while the lowest was found in the natural regeneration treatments with about half the values as in the plantation stands. There was a significant negative correlation between percent canopy openness and abundance of regeneration in the understory. Two distinctive clusters separated the regeneration treatments from the mixed and pure plantations at a very low Bray–Curtis similarity value. The natural regeneration treatments are separated from mixed and pure plantations in the two-dimensional ordination. The lack of difference between the understory make-up of pure and mixed plantations in abundance, species richness, and seed-dispersal syndromes of understory species suggests that planting mixed stands is not necessarily superior to planting pure stands for promoting understory diversity of woody species. While regeneration of woody species can be faster under pure- or mixed-species plantations than in open pastures, the abundance, richness and species composition depends on each plantation species, or species assemblages in case of the mixtures.     

Growth characteristics of some native tree species used in silvopastoral systems in the humid lowlands of Costa Rica

Research for developing silvopastoral systems includes tree species selection and management, and interactions of trees with pastures. In the Caribbean humid lowlands of Costa Rica, total height, diameters at breast height (DBH), survival, basal area and bole volume were compared among three plantations with four species each, in pure stands as well as in the mixture of the four species. In Plantation 1, at 86 months old the pure plantations of Jacaranda copaia and Vochysia guatemalensis, and the mixed plantations (Jacaranda copaia + Vochysia guatemalensis + Calophyllum brasiliense + Stryphnodendron microstachyum) had the largest DBH. Vochysia guatemalensis is one of the most preferred species in the region for reforestation of degraded pastures, while Jacaranda copaia is not preferred because it has low local economic value. In Plantation 2, at 81 months, the greatest DBH was found in Virola koschnyi, Terminalia amazonia and the mixed plantations (Virola koschnyi + Terminalia amazonia + Dipteryx panamensis + Pseudosamanea guachapele). Dipteryx panamensis has good economic value and has a relatively open canopy that produces moderate shade. Virola koschnyi and Terminalia amazonia have denser canopies, are preferred by farmers, and they could be planted in low density in silvopastoral systems. In Plantation 3, at 70 months, the greatest DBH was found in Vochysia ferruginea, Hieronyma alchorneoides and the mixed plantation (Vochysia ferruginea + Hieronyma alchorneoides + Genipa americana + Balizia elegans). Most of these species are currently used in silvopastoral systems in the region. Mixed plantations may combine beneficial effects of the different species' characteristics, with the additional advantage of product diversification. This revised version was published online in June 2006 with corrections to the Cover Date.     

Carbon accumulation in aboveground and belowground biomass and soil of different age native forest plantations in the humid tropical lowlands of Costa Rica

Generic or default values to account for biomass and carbon accumulation in tropical forest ecosystems are generally recognized as a major source of errors, making site and species specific data the best way to achieve precise and reliable estimates. The objective of our study was to determine carbon in various components (leaves, branches, stems, structural roots and soil) of single-species plantations of Vochysia guatemalensis and Hieronyma alchorneoides from 0 to 16 years of age. Carbon fraction in the biomass, mean (±standard deviation), for the different pools varied between 38.5 and 49.7% (±3 and 3.8). Accumulated carbon in the biomass increased with the plantation age, with mean annual increments of 7.1 and 5.3 Mg ha⁻¹ year⁻¹ for forest plantations of V. guatemalensis and H. alchorneoides, respectively. At all ages, 66.3% (±10.6) of total biomass was found within the aboveground tree components, while 18.6% (±20.9) was found in structural roots. The soil (0–30 cm) contained 62.2 (±13) and 71.5% (±17.1) of the total carbon (biomass plus soil) under V. guatemalensis and H. alchorneoides, respectively. Mean annual increment for carbon in the soil was 1.7 and 1.3 Mg ha⁻¹ year⁻¹ in V. guatemalensis and H. alchorneoides. Allometric equations were constructed to estimate total biomass and carbon in the biomass which had an R ²aj (adjusted R square) greater than 94.5%. Finally, we compare our results to those that could have resulted from the use of default values, showing how site and species specific data contribute to the overall goal of improving carbon estimates and providing a more reliable account of the mitigation potential of forestry activities on climate change.     

Timber investment returns for selected plantations and native forests in South America and the Southern United States

Timber investment returns were estimated for the principal exotic and selected native species in the Southern Cone of Latin America and in the Southern United States. Exotic eucalyptus plantations in South America were most profitable, with internal rates of returns (IRRs) ranging from 13% to 23%, followed by exotic loblolly pine, with IRRs from 9% to 17%. Average loblolly pine plantation returns in the US South were less profitable, with an IRR of about 9.5%, and natural forest management in the South had IRRs of 4% to 8%. Subtropical native species plantations of the best araucaria and nothofagus species had reasonable financial returns, with IRRs ranging from 5% to 13%. Subtropical or tropical native forests had fewer commercial timber species, and had much lower growth rates and returns. Their IRRs were less than 4%, or even negative for unmanaged stands. State subsidy payments for forest plantations or for timber stand improvements increased IRRs somewhat and reserving areas for environmental protection reduced their IRRs slightly. Including land costs in the cash flows decreased these internal rates of return substantially. Natural stand returns in Latin America were much less than those of plantations, but management of those stands offered better rates of return than only holding the land.     

Growth,carbon sequestration,and management of native tree plantations in humid regions of Costa Rica

The Costa Rican government has provided incentives for reforestation programs since 1986 and initiated a Payment for Environmental Services program in 1996. These incentives yielded native species reforestation programs throughout the country. This research aims to provide information about growth, carbon sequestration, and management of seven native tree species (Vochysia guatemalensis, Vochysia ferruginea, Hyeronima alchorneoides, Calophyllum brasiliense, Terminalia amazonia, Virola koschnyi, and Dipteryx panamensis) growing in small and medium-sized plantations in the Caribbean and Northern lowlands of Costa Rica. A total of 179 plots were evaluated in 32 farms. Overall, I found that V. guatemalensis, V. ferruginea, H. alchorneoides, and T. amazonia were the species with the fastest diameter, total height, and volume growth; and T. amazonia and D. panamensis sequestered more carbon. Moreover, I found that the plantations that had been thinned before this assessment had the best growth. The results of the present research enhance the criteria elaborated in previous research findings to improve species choices for reforestation and silvicultural management in Costa Rica and in other regions with similar ecological features. Furthermore, they support the concept that tropical plantations can serve diverse economic, social, and ecological functions that may ultimately help reduce atmospheric CO₂ accumulation.     

Promoting intake of Cratylia argentea as a dry season supplement for cattle grazing Hyparrhenia rufa in the subhumid tropics

Cratylia argentea, a drought tolerant shrub, is known to produce significant amounts of fodder during the dry season but little is known of its nutritive value for feeding animals. Two studies of quality and intake of Cratylia argentea, as an arboreal protein supplement for crossbred dual purpose cattle grazing Hyparrenhia rufa in the dry season, were carried out in San Miguel de Barranca, Puntarenas, Costa Rica (January–May). The first experiment was a 2 × 2 factorial arrangement of two forms of C. argentea foliage presentation: 1) fresh or wilted; and 2) sprayed or not with sugarcane molasses diluted in water. Wilting did not affect crude protein content and in vitro dry matter digestibility (IVDMD) but treating fresh C. argentea with molasses increased IVDMD by from 55.6 to 60.8% (P < 0.01). Intake of fresh C. argentea increased by 32% with wilting and by 43% with molasses treatment. Pooling data across treatments, a quadratic relationship between dry matter intake of H. rufa (y) and the amount of N consumed in C. argentea (x) was found (y = 1.25 + 0.095x – 0.001x ²). In the second experiment, milk production from the following treatments was evaluated: 1) chicken litter and molasses (D₁); 2) chicken litter, sugar cane and wheat bran (D₂); and 3) C. argentea, sugar cane and wheat bran (D₃). Milk yields averaged 6.0 kg cow^–1 day^–1 and there were no significant differences between treatments. Milk fat was lower for D₂ (2.7%), which had a higher amount of sugar cane in the diet, but differences were not significant. On the other hand, non-fat solids and lactose concentration of milk was higher (P < 0.05) for D₂. Wilting or molasses treatment were effective management strategies for increasing dry matter intake of C. argentea. A higher dry matter intake of C. argentea was associated with increased intake of the fibrous H. rufa grass. Diets with C. argentea as the main protein source gave similar milk production levels and quality to those with chicken litter.