上层抚育对冠下红松生长影响的调查与分析

以吉林省黄泥河林业局金沟林场27林班24小班的林分为调查对象,对采取上层抚育和未采取抚育的林冠下人工更新红松的生长、健康状况和单位面积保存率进行调查与分析.结果表明:抚育林分红松幼树的高生长、地径生长较未抚育林分分别增加了40.0％和50.0％,冠幅增长量增加了15.5％;林分健康植株达到95.0％,增加了48.9％,...     

Effects of feathermoss removal, thinning and fertilization on lodgepole pine growth, soil microclimate and stand nitrogen dynamics

The influence of a continuous feathermoss layer (dominated by Pleurozium schreberi) on soil temperature, soil water potential and tree growth was assessed in a 5-year study. The feathermoss layer was manually removed from 900 m² plots in lodgepole pine (Pinus contorta) stands of northwestern Alberta. The interception and retention of nutrients by moss was estimated in N fertilization treatments with and without moss removal. As well, the potential for intraspecific competition to affect nutrient dynamics was assessed through a thinning treatment. Removal of the moss layer increased soil temperatures in summer and increased the period when soil was frost free, but the plots without moss had soil temperature as low as −13 °C in one winter period with little snow cover. Moss removal did not significantly affect N concentrations of the tree foliage but did reduce needle weight. Thinning had little effect on the rate of diameter growth after the first 3 years, but produced a significant increase in growth by year 4. Fertilization had a consistently positive effect on radial stem increments and N content of foliage, regardless of the presence of moss on the plot. Overall, the intraspecific competition between trees was apparently greater than interspecific competition between moss and trees.     

Timing and intensity of precommercial thinning in Pinus sylvestris stands

The effects of timing and intensity of precommercial thinning were studied in three Scots pine artificially regenerated stands on Vaccinium forest sites in southern Finland. A two-level factorial design (3×3) was used in each stand: thinning at dominant height of 3, 6 and 9 m to 1000, 1600 and 2200 stems ha^?1. The effects of the treatments were analysed after a period of 23–25 yrs when the dominant height was 14–15 m. Early thinning resulted in the highest standing volume and amount of merchantable wood, and also in slightly accelerated height development. Thinning to 1000 stems ha^?1 caused a considerable production loss, but there were no differences between the densities of 1600 and 2200 stems ha^?1. Branches became thicker after early thinning, but the differences between the treatments were negligible for crop trees. Crown ratio was lowest as the result of early or moderate thinning (2200 stems ha^?1).     

Effects of thinning on nutrient content pools in two Pinus sylvestris forests in the western Pyrenees

Abstract

Nutrient content [nitrogen (N), phosphorus (P), potassium (K), calcium (Ca) and magnesium (Mg)] in needles, branches and stems before (P0, not thinned) and after thinning (P20 and P30, 20% and 30% of basal area removed, respectively) was studied in 37- and 32-year-old Pinus sylvestris L. forests in the western Pyrenees: Garde and Aspurz. Thinning significantly decreased all nutrient pools in P20 and P30 relative to P0 on both sites, but no significant differences were found between P20 and P30 owing to low statistical power. Thinning increased the differences between the two forests for total above-ground content of P and Mg, and for the content of N, P and K in the 1999 needles cohort. The former result was due in part to the higher concentration of P (needles and branches) and Mg (needles) in Garde. Therefore, the importance of needles relative to the other nutrient pools increased for N and P after thinning. This may have resulted from the fact that the removal of dead trees with low needle biomass was greater in Garde than in Aspurz. Based on the percentage of basal area removed, nutrient removal in Garde for P30 was higher than expected, apparently owing to an increase in branch removal relative to P20. These results indicate that effects of thinning on nutrient pools were influenced by differences in stand structure and nutrient tissue concentrations between sites.     

Effects of growth regulators on germination of picea abies and Pinus sylvestris seeds

Nine seed lots each of Norway spruce (Picea abies L.), and Scots pine (Pinus sylvestris L.) were treated with combinations of the growth regulators: Indole‐3‐acetic acid, indole‐3‐butyric acid, etephone, kinetin, fusicoccin, and the gibberellins A₁ A₃ A_4/7, and A₉. GA₉ GA_4/7 and fusicoccin significantly promoted the germination of five Norway spruce, and three Scots pine, seed batches. Independent of species, the germinability of seeds treated with auxins, etephone, kinetin, GA₁ and GA₃ were unaffected or reduced. The rate of germination was stimulated for both species when treated with GA₉ or GA_4/7. The germination percentage for Scots pine seeds was enhanced by GA₉ but not by GA_4/7. Neither of the two GAs enhanced the germination percentage of Norway spruce seeds.     

Effects of temperature on shoot growth in northern provenances of Pinus sylvestris L

Effects of temperature on shoot growth in northern provenances of Pinus sylvestris L. were studied under natural long-day conditions (Lat. 69 degrees 39' N) at controlled temperatures. The optimum constant temperature for formation of stem unit primordia in the terminal resting bud was 18 to 21 degrees C. Stem unit number decreased linearly with temperature between 15 and 9 degrees C. Temperature during bud formation had a significant effect on final stem unit length achieved in the following year. Final shoot length was significantly affected by temperature during shoot elongation. The optimum constant temperature for elongation growth was 18 to 21 degrees C. The rate of shoot elongation was positively correlated with the number of stem units.     

Timber quality after thinning from above and below in stands of Pinus sylvestris

Timber quality and logging damage after different thinning types in Scots pine (Pinus sylvestris L.) stands were studied in a field experiment in north Sweden. Thinning operations were mechanized and carried out according to normal Swedish practise. The treatments were thinning from below and thinning from above. The quality of the residual stand was evaluated using variables such as diameter of the thickest branch, stem taper, annual ring width, straightness and lean of trees. The only variables showing differences between treatments were lean of tree and stem straightness. No difference in the frequency of trees with logging damage was found. The conclusion of this study is that differences between thinning types in first thinning may be evened out when carried out as in commercial forestry due to extraction of strip road and damaged trees. Consequently, differences in timber quality and logging damage in the residual stand may be small.     

Optimizing thinning and rotation in a stand of Pinus sylvestris on a drained peatland site

A non‐linear programming algorithm was combined with a stand simulator consisting of individual‐tree, distance‐dependent diameter growth, mortality and thinning models, and a distance‐independent height growth model. The Hooke and Jeeves’ direct search method was used for determining the optimum thinning schedule and rotation length. The optimum solutions for both deterministic and stochastic optimization, with stochastic growth predictions, were calculated. In the optimization, stand density was regarded as a function of the distance to ditch. On the rather infertile site, it was most profitable to keep the stand about twice as dense near ditches and the extraction road, which was placed midway between two adjacent ditches. Stochasticity did not affect the optimum regimes markedly nor did it increase the expected returns of the optimum regimes.     

Effects of pre-commercial thinning on growth and reproduction in post-fire regeneration of Pinus halepensis Mill

Context

Excessive recruitment in post-fire regeneration of Pinus halepensis can require pre-commercial thinning. The 1994 Moratalla fire (Spain) and the thinning applied there since 2004 provided good conditions for testing pre-commercial thinning effectiveness.

Aims

To analyse pre-commercial thinning effects on tree size, reproductive potential, stem biomass and annual growth in 15-year-old saplings.

Methods

Twenty nine circular plots (5 m radius) were established based on (1) years since thinning and (2) aspect. Non-thinned plots were included as controls. Sapling variables were measured and pine cones were counted and classified according to maturity. Three saplings per plot were felled, five sample disks were extracted from each and tree rings were analysed with Windendro® software.

Results

Sapling size, annual growth and stem biomass were higher in thinned plots than in control plots 2–4 years after thinning. Annual growth declined five years after thinning. Cone counts were higher 2–5 years after thinning, depending on the cone type. Aspect influenced some variables.

Conclusion

Thinning accelerated stand maturity but positive effects were indiscernible until 2–5 years later. Serotinous cones, which are fundamental to regeneration after fire, increased in number after 5 years. The effects of aspect were inconclusive due to interactions with years since thinning.     

Effects of environmental conditions on onset of xylem growth in Pinus sylvestris under drought

We determined the influence of environmental factors (air and soil temperature, precipitation, photoperiod) on onset of xylem growth in Scots pine (Pinus sylvestris L.) within a dry inner Alpine valley (750 m a.s.l., Tyrol, Austria) by repeatedly sampling micro-cores throughout 2007-10 at two sites (xeric and dry-mesic) at the start of the growing season. Temperature sums were calculated in degree-days (DD) ≥5 °C from 1 January and 20 March, i.e., spring equinox, to account for photoperiodic control of release from winter dormancy. Threshold temperatures at which xylogenesis had a 0.5 probability of being active were calculated by logistic regression. Onset of xylem growth, which was not significantly different between the xeric and dry-mesic sites, ranged from mid-April in 2007 to early May in 2008. Among most study years, statistically significant differences (P<0.05) in onset of xylem growth were detected. Mean air temperature sums calculated from 1 January until onset of xylem growth were 230 ± 44 DD (mean ± standard deviation) at the xeric site and 205 ± 36 DD at the dry-mesic site. Temperature sums calculated from spring equinox until onset of xylem growth showed somewhat less variability during the 4-year study period, amounting to 144 ± 10 and 137 ± 12 DD at the xeric and dry-mesic sites, respectively. At both sites, xylem growth was active when daily minimum, mean and maximum air temperatures were 5.3, 10.1 and 16.2 °C, respectively. Soil temperature thresholds and DD until onset of xylem growth differed significantly between sites, indicating minor importance of root-zone temperature for onset of xylem growth. Although spring precipitation is known to limit radial growth in P. sylvestris exposed to a dry inner Alpine climate, the results of this study revealed that (i) a daily minimum air temperature threshold for onset of xylem growth in the range 5-6 °C exists and (ii) air temperature sum rather than precipitation or soil temperature triggers start of xylem growth. Based on these findings, we suggest that drought stress forces P. sylvestris to draw upon water reserves in the stem for enlargement of first tracheids after cambial resumption in spring.     

Canopy density in stands of Picea abies and Pinus sylvestris after different thinning methods

Altogether 82 plots (261 estimations) of Picea abies (L.) Karst, and 193 plots (360 estimations) of Pinus sylvestris (L.) stands were estimated by a vertical tube. The “crown free projection”, CFP, of stands thinned in three methods with different thinning grades was measured: unthinned, heavily and very heavily thinned, heavily thinned delayed first thinning, extra heavily thinned and thinned from the top. Basal area (m²ha^?1) density (stems ha^?1) and diameter sum (m ha^?1) were plotted against CFP. Basal area was the best practical measure of stand in this study. Generally Scots pine stands have higher CFP and the curves are steeper than in Norway spruce stands. Depending on the grade of thinning, heavily and very heavily thinned spruce stands, delayed first thinning included, have CFP values of 10–15% and stands thinned from the top, 20–40%, compared with 30–80% and 30–60% respectively in pine stands. Extra heavily thinned stands have the highest CFP, 20–80% in spruce and 50–90% in pine stands. The CFP levels after thinning are too high in pine stands for avoidance of sucker and sprout production of aspen and birch. In dense Norway spruce stands thinned from the top or heavily and very heavily thinned, the CFP values are low enough (≤30%) to diminish the production of suckers.     

Effects of maternal environment on mortality and growth in young Pinus sylvestris in field trials

The same full-sib families of Scots pine (Pinus sylvestris L.) were created by artificial pollination of genetically identical grafts at three localities in Sweden at approximately 56, 59 and 64 degrees N. Two field trials were established with one-year-old plants in different years at latitude 64 degrees N. Height and survival were monitored for 4-5 years following planting. Maternal environment had significant aftereffects on the height of progeny. In both field trials, plants with the maternal parent at 59 degrees N were tallest and plants with the maternal parent at 64 degrees N were shortest. The aftereffects of maternal environment were as large in six-year-old plants as in one-year-old plants. In one field trial, maternal environment had a significant effect on mortality six years after germination but not after three years. Mortality increased with decreasing latitude of the maternal location. We conclude that the aftereffects of maternal environment are too large and too permanent to be regarded as unimportant.     

Influence of precommercial thinning grade on branch diameter and crown ratio in Pinus sylvestris in southern Sweden

Precommercial thinning is regarded as one of the most important measures for influencing timber quality in stands of Scots pine (Pinus sylvestris L.). The present study considers the influence of both thinning intensity and tree height at the time of thinning on diameter of the thickest branch and crown ratio. A total of 115 plots (either naturally regenerated, planted or sown, and either thinned down to 528–8000 stems?ha^?1 or untreated) included in 20 sites in southern Sweden were analysed. The average tree height after thinning varied from 1.2 to 8.3?m. An increasing number of remaining stems resulted in a reduction in branch diameter, although the reduction appeared to be only minor if the number of stems after thinning was more than 3000 stems?ha^?1. It was found that late thinning reduced the diameter of the thickest branch. The crown ratio decreased with stand height, number of stems after thinning and average height at thinning. The results were consistent for all trees and for the 500 thickest trees per hectare.     

不同间伐强度对火炬松林分生长的影响

对11年生火炬松人工林进行间伐强度试验,经3年观测表明:间伐促进了林分生长,间伐3年后与对照相比,胸径的年生长量增加了0.24～0.80㎝,树高的年生长量增加了0.19～0.33m,林分蓄积的年生长量增加了6.783 1～14.812 5m3/hm2,促进林分蓄积生长效果最好的处理是间伐强度为30%.采用30%强度进行初次间伐,可显著提高蓄积生长量,有利于发挥火炬松造林的经济和生态效益.     

环境因子对樟子松生长发育的影响

通过查阅文献资料和实地调查,从樟子松新技术造林试验和存在的问题以及环境因子等几个方面分析了樟子松的生长发育情况,以期为樟子松在榆林沙区大面积推广和进一步深入研究提供科学依据。     

The effects of timing of pre-commercial thinning and stand density on stem form and branch characteristics of Pinus sylvestris

The effects of pre-commercial thinning on tree and branch characteristicswere analysed at two experimental sites in northern Sweden,where pre-thinning of Scots pine stands to 600, 1000 and 1800stems per hectares at heights of 1.5, 3, 5 and 7 m had beencombined in a factorial design. A total of 90 trees were cutand branches were measured in whorls sampled at 1-m intervals,starting with the whorl closest to 0.5 m above ground. Effectsof the treatments on tree and branch characteristics were analysedby a fixed-effect analysis of variance model, which was appliedseparately for each experimental site and whorl height. Diameterat breast height (d.b.h.) and living crown to height ratio decreased,while the height/d.b.h. ratio (stem form) increased with increasingstand density, and generally with increased height at thinning.No treatment effect was found on the number of branches perwhorl or branch angle, but branch diameter (BD) was significantlyinfluenced by both stand density after thinning and height atthinning. BD decreased with increases in stand density and decreasedwith increases in height at pre-commercial thinning. Relativebranch size (RBS), defined as the ratio between the sum of thediameters of branches in a whorl and stem perimeter under barkat the location of that whorl, was significantly influencedonly by stand height at the time of treatment. RBS decreasedby 4–6 per cent in whorls located below the current baseof the living crown with each step increase in stand heightat pre-commercial thinning. The results indicate that the numberof stems after pre-commercial thinning has a greater impacton tree size and shape than timing of pre-commercial thinning.Stand density after thinning also has a greater impact on branchsize, in the sense that it will affect branch size further upthe tree than the timing of pre-commercial thinning. However,by delaying the time for pre-commercial thinning it is possibleto reduce the RBS in the lower part of the stem. This mightbe an important measure to improve timber quality later in therotation period, which cannot be done solely by regulating standdensity.     

Effects of overstorey canopy, plant-plant interactions and soil properties on Mediterranean maritime pine seedling dynamics

Seedling emergence, survival and early growth of the Mediterranean conifer P. pinaster were studied under closed canopy and open canopy (gaps) cover conditions in a Mediterranean forest of central Spain during two consecutive years (March 2008 to January 2010). Our main objective was to understand how overstorey structure, shrubs and soil properties influence recruitment in this species. Natural emergence and seedling survival were significantly better under closed canopy cover than under open canopy during the two consecutive years of the study; survival increased as radiation decreased. Proximity to shrubs under closed canopy cover was associated with lower mortality rates as well as a positive and apparently transitive effect on early growth. Amelioration of microclimatic conditions is hypothesized as the primary facilitation agent of those observed. Younger seedlings established during the spring of 2008 experienced higher mortality rates than older seedlings established before 2008, and soil properties affected seedling survival significantly. Water stress during summer appeared to be the main cause of seedling mortality in both years. Our study suggests that seedlings of shade-intolerant species may require overstorey cover to establish successfully, and that positive plant-plant interactions in forest communities may be more important than expected in the milder conditions provided by overstorey canopy cover.     

大强度抚育对油松人工林天然更新的影响

以油松人工林为研究对象,在大强度抚育条件下,通过林分层次结构、各层次林木胸径结构、树高结构及其林木分布格局等指标研究林下天然更新状况,结果表明:油松人工林在22a、32a时分别进行株数强度50%和40%的下层抚育,后经20a天然更新(林分年龄52a),林分形成明显层次结构,分为主林层、演替层及更新层,整个林分林木胸径、树高结构均呈多峰分布,其中主林层、演替层林木胸径和树高结构均呈单峰山状分布,更新层林木胸径和树高结构呈多峰分布,各林层平均胸径分别为23.2cm、12.6cm和3.1cm,平均树高分别为18.4m、14.5m和2.8m,而对照标准地胸径、树高结构均呈单峰分布,平均胸径、树高分别为18.6cm及17.9m;主林层、演替层各层林木空间分布呈均匀分布,密度分别为840株/hm~2和1 125株/hm~2,更新层林木空间分布呈聚集分布,密度为675株/hm~2,对照样地林木空间分布偏向均匀分布,密度为1 605株/hm~2。     

Comparing the intra-annual wood formation of three European species (Fagus sylvatica, Quercus petraea and Pinus sylvestris) as related to leaf phenology and non-structural carbohydrate dynamics

Monitoring cambial phenology and intra-annual growth dynamics is a useful approach for characterizing the tree growth response to climate change. However, there have been few reports concerning intra-annual wood formation in lowland temperate forests with high time resolution, especially for the comparison between deciduous and coniferous species. The main objective of this study was to determine how the timing, duration and rate of radial growth change between species as related to leaf phenology and the dynamics of non-structural carbohydrates (NSC) under the same climatic conditions. We studied two deciduous species, Fagus sylvatica L. and Quercus petraea (Matt.) Liebl., and an evergreen conifer, Pinus sylvestris L. During the 2009 growing season, we weekly monitored (i) the stem radial increment using dendrometers, (ii) the xylem growth using microcoring and (iii) the leaf phenology from direct observations of the tree crowns. The NSC content was also measured in the eight last rings of the stem cores in April, June and August 2009. The leaf phenology, NSC storage and intra-annual growth were clearly different between species, highlighting their contrasting carbon allocation. Beech growth began just after budburst, with a maximal growth rate when the leaves were mature and variations in the NSC content were low. Thus, beech radial growth seemed highly dependent on leaf photosynthesis. For oak, earlywood quickly developed before budburst, which probably led to the starch decrease quantified in the stem from April to June. For pine, growth began before the needles unfolding and the lack of NSC decrease during the growing season suggested that the substrates for radial growth were new assimilates of the needles from the previous year. Only for oak, the pattern determined from the intra-annual growth measured using microcoring differed from the pattern determined from dendrometer data. For all species, the ring width was significantly influenced by growth duration and not by growth rate, which differs from previous studies. The observed between-species difference at the intra-annual scale is key information for anticipating suitability of future species in temperate forests.     

红松人工林不同间伐强度对降雨再分配的影响

为研究间伐对红松人工林水源涵养功能的影响程度,该文以本溪县草河口地区70 年生不同间伐强度(对照区、中度区、极强区)的红松人工林为研究对象,研究不同间伐强度对红松人工林树干径流、穿透雨及林冠截留的影响,结果表明:红松人工林的穿透雨、树干径流以及林冠截留都会随降雨强度增加而增加,穿透雨量随间伐强度增加而增加,不同间伐强度的穿透雨率差异显著(p<0.05),树干径流率差异不显著。林冠截留量随间伐强度增加而减小,且不同间伐强度的林冠截留率差异显著(p<0.05)。