Microclimatic differences between young longleaf-pine silvopasture and open-pasture

Information is needed relative to changes that occur in microclimatic conditions in the early stages of open-pasture to pine silvopasture conversion in the Southeastern USA. This research tested the hypothesis that microclimatic conditions would vary between young-pine silvopasture and open-pasture. The research was conducted in a longleaf-pine (Pinus palustris Mill.)-bahiagrass (Paspalum notatum Flugge) silvopasture and adjacent open bahiagrass pasture at Americus, Georgia. Longleaf-pine was planted in the bahiagrass pasture in December 2000; the trees were not pruned or thinned until the study was completed. Microclimatic parameters (air temperature, soil temperature at 5 and 10 cm, wind speed, gust speed, wind direction, humidity, dew point, rainfall, soil-moisture content, total solar radiation, and photosynthetically active radiation) were collected from November 2005 to October 2008 using HOBO^© weather stations located in each pasture type. Data were analyzed to determine overall, as well as seasonal, monthly, and diurnal differences between pasture types. Evapotranspiration was calculated from air temperature, wind speed, relative humidity, and total solar radiation. With the exception of rainfall, overall average values for all microclimatic parameters were different between pasture types. Silvopasture had higher overall average values for air temperature, soil temperature at both depths, and soil water content, but lower values for all remaining microclimatic parameters. Seasonal, monthly, and diurnal differences between pasture types were also detected for several parameters. The results suggested that water loss can be significantly reduced in the early stages of open-pasture to longleaf-pine silvopasture conversion in the Southeastern USA.     

Cattle distribution and behavior in southern-pine silvopasture versus open-pasture

Shade present in silvopasture systems could reduce heat stress associated with microclimatic conditions that characterize warm-weather portions of the year on the Coastal Plain of the Southeast USA. Objectives of this research were to: (1) quantify diurnal distribution patterns of landscape use and behavior of cattle in loblolly-pine (Pinus taeda) silvopasture versus open-pasture landscapes, and (2) relate observed differences in landscape use and cattle behavior patterns between the two pastures to differences in microclimatic conditions, and forage quantity and quality. The research was conducted in Chipley, Florida USA within a 5-ha cell of a loblolly-pine-bahiagrass (Paspalum notatum) silvopasture (tree age 20 year), and a 5-ha open bahiagrass pasture with unlimited access to an adjacent 1-ha wooded area. One-day observations of diurnal distribution and behavior of cattle were conducted in March, June, and September 2007; microclimatic conditions were measured, and forage quantity and quality were estimated within each landscape. Cattle utilized the landscape more evenly in the silvopasture versus the open-pasture and this difference was mainly attributed to reduced solar radiation recorded in the silvopasture. Grazing was the dominant behavior in the silvopasture while loafing was dominant in the open-pasture. Shade present in silvopasture appeared to reduce heat stress for cattle grazing during warm-weather portions of the year on the Coastal Plain of the Southeast USA. Further study is needed to determine how this reduction in heat stress influences cattle performance in southern-pine silvopasture, and the nature and extent of the interactions among animal distribution and behavior, microclimatic conditions, and forage characteristics in these systems.     

Conversion of deciduous forest to silvopasture produces soil properties indicative of rapid transition to improved pasture

Differences in soil properties between forests and pastures have been well documented in the literature, especially under coniferous forests. However, since nearly all of these reports have been time-point comparisons, utilizing long-term paired-sites, properties of transitional states and time of their appearance can only be inferred at present. In this study, a deciduous forest ecosystem was converted to a silvopasture ecosystem by tree thinning, fertilization, and sheep incorporation of seed and forest litter. After 2 years, topsoil (0–15 cm) physico-chemical properties, particularly P fractions, and phosphatases were monitored over the growing season in these ecosystems, and a nearby pasture ecosystem. Initially, before spring vegetative growth, differences were found for pH, exchangeable cations and soil moisture, most of which could be explained by management history. Compared to forest, organic-C (C_o) and organic-N (N_o) concentrations were reduced in silvopasture by 17 and 9%, respectively, indicative of substantial litter decomposition. Most values for all these physico-chemical properties for silvopasture were intermediate between forest and pasture, and generally remained so throughout the growing season. Initial total P (TP_t), organic-P (TP_o) and inorganic-P (TP_i) concentrations were generally as anticipated for the forest and pasture. Silvopasture, however, had 36 and 23% greater TP_o than forest and pasture, respectively, presumably due to fertilizer-P immobilization induced by incorporation of forest litter. Total P components remained essentially constant over the growing season in all ecosystems, with the exception of pasture, likely due to high forage TP_i uptake. Bray I-extractable-organic-P (BrP_o) and bicarbonate-extractable-organic-P (BiP_o) concentrations, although consistently highest in the forest and silvopasture soils, were not reflective of the increase in TP_o under silvopasture. Acid phosphatase (PMEac) activities were highest in spring in all the ecosystems, then gradually declined to typically 25–50% initial activities. Alkaline phosphatase (PMEal) activities showed a broadly-similar pattern, with exception of forest and silvopasture soils, which exhibited low activities throughout the season. For the entire data set, PMEac and PMEal activities were poorly correlated with TP_i, BrP_i, and BiP_i. These results demonstrate that conversion of forest to silvopasture results in soil changes indicative of its rapid transition to pasture and that an increased P_o reservoir results that should be taken into account in fertilizer-P recommendations for temperate silvopastures.     

Modeling forage growth in a Midwest USA silvopastoral system

Tree effects on understory pasture growth in a silvopastoral system were modeled by explicit simulation of tree canopy light and rainfall interception, evapotranspiration, and nutrient uptake. The algorithms to model these effects were incorporated into a multispecies grazing simulation model, GRASIM, to form the Silvopasture GRASIM model (SGRASIM). The new model was evaluated using forage biomass data and soil moisture data collected from a silvopasture field experiment with black walnut (Juglans nigra L.). The SGRASIM model performed well in simulating the growth of three competing dominant forage species (orchardgrass [Dactylis glomerata L.], Kentucky bluegrass [Poa pratensis L.], and tall fescue [Festuca arundinacea (Schreb.)] in the pasture both under tree canopy and in open pasture (linear regression of observed on simulated biomass for the species gave r ² values above 0.97). Model growth parameters for forage under tree canopy, compared with those for an open pasture, bear testament to the shading effects from the forest canopy in terms of reduced photosynthetic efficiency, increased leaf area ratio, and photosynthate partitioned to aboveground biomass. The new model reasonably followed the soil moisture time series in the upper soil layer (0–30 cm), where the bulk of the forage roots reside.     

A financial analysis of a silvopasture system in southern Mississippi

Silvopasture is reemerging as a land use in the southern US. Alternate land use treatments based on field trials for timber and pasture for beef cattle production were financially evaluated. Multiple-use management aspects of these systems were further illustrated by the addition of fee hunting. Land Expectation Values (LEVs) were lower when silvopasture treatments were compared to steer grazing only. However, silvopasture treatments compared favorably to some grazing treatments. LEVs were higher when silvopasture treatments were compared to commercial forest plantation applications on similar sites. Silvopasture systems promote multiple-use management of the land under an environmentally friendly cropping system whereby certain types of wildlife can thrive. This is particularly relevant in light of recent increases in fee hunting in the South. LEVs were reduced by $289, $200, and $151 ha⁻¹ at discount rates of 5%, 7%, and 9% respectively, when fee hunting was excluded in the recommended silvopasture treatment. The monetary value of a wildlife component in this system can be interpreted as the expected value gained per hectare per rotation in perpetuity when fee hunting is part of the management plan. On average, it represented an 8.6% gain in LEVs for this treatment. Results from this study support the potential for silvopasture applications in the South for private landowners. Cattle grazing of improved forage in commercially productive loblolly pine (Pinus taeda) stands can relieve annual cash flow problems inherent in tree production. This revised version was published online in June 2006 with corrections to the Cover Date.     

Light Measurement Methods Related to Forage Yield in a Grazed Northern Conifer Silvopasture in the Appalachian Region of Eastern USA

The Appalachian region is characterized by hilly topography and a humid temperate climate. In most areas agriculture is limited to pasture although the native climax vegetation is a species-diverse forest. Silvopasture systems can help diversify and increase farm income. Information is needed on the yield response of forage grown as an understory crop among trees. The light environment of a conifer silvopasture was characterized by three methods, a hand-held photosynthetically active radiation (PAR) meter for quick measurements over a large area, fixed PAR meters recorded using data loggers for a large number of measurements over time, and hemispherical photography with software to calculate seasonal direct beam radiation. Light data were considered in terms of forage yield. Plots were harvested when forage reached 20–25 cm in height after which the entire area was grazed by sheep. There were limitations to all methods of quantifying radiation environment for predicting yield. Yield decreased linearly with decreased PAR; however, data variability was high and correlations, while statistically significant, were weak. Grazed silvopastures are dynamic with shifting yield patterns in response to the interactions between the spatially variable soil, changing seasonal environment, and spatially variable animal impacts for each grazing event.     

Mediterranean cork oak wooded grasslands: synergies and trade-offs between plant diversity,pasture production and soil carbon

Mediterranean wooded grasslands that emerge from silvopastoral activities are multifunctional systems that result in high biodiversity and offer ecosystem services such as forage production and soil carbon sequestration. During 3 years, ten grazed wooded grassland fields were studied in the Berchidda–Monti long-term observatory, located in NE Sardinia, Italy, with the aim of exploring the synergies and trade-offs between biodiversity and selected ecosystem services. Positions below and outside the canopy of three cork oak trees in each field were randomly selected to compare seasonal pasture production, pasture utilization rate by animals, botanical composition, biodiversity indicators (Shannon index and plant species richness) and soil organic carbon. In autumn, dry matter production of pasture was similar in the two positions; in two winters out of three it was greater below the trees than outside, and in spring it was greater outside than below the trees. While plant species richness and Shannon index were not significantly influenced by the position, the overall wooded grassland plant species richness was 31% higher than that outside of the tree crown. The soil organic carbon content in the 0–40-cm soil layer was also higher below the trees. Our findings highlight that if the main purpose of the wooded grasslands is to provide forage for grazing animals rather than conserving and/or enhancing plant diversity and soil fertility, the presence of trees constrains the overall forage productivity, although the greater forage availability in winter under the trees can contribute to improve the seasonal distribution of forage production.     

Comparing silvopastoral systems and prospects in eight regions of the world

Silvopasture systems combine trees, forage, and livestock in a variety of different species and management regimes, depending on the biophysical, economic, cultural, and market factors in a region. We describe and compare actual farm practices and current research trials of silvopastoral systems in eight regions within seven countries of the world: Misiones and Corrientes provinces, Argentina; La Pampa province, Argentina; northwestern Minas Gerais, Brazil; the Aysén region of Patagonia, Chile; the North Island of New Zealand; the Southeast United States; Paraguay; and Uruguay. Some countries use native trees and existing forests; some use plantations, particularly of exotic species. Natural forest silvopasture systems generally add livestock in extensive systems, to capture the benefits of shade, forage, and income diversification without much added inputs. Plantation forest systems are more purposive and intensive, with more focus on joint production and profits, for small owners, large ranches, and timber companies. Trends suggest that more active management of both natural and planted silvopastoral systems will be required to enhance joint production of timber and livestock, achieve income diversification and reduce financial risk, make more profit, improve environmental benefits, and realize more resilience to adapt to climate change.     

Cumulative Forage Production, Forage Quality and Livestock Performance from an Annual Ryegrass and Cereal Rye Mixture in a Pine Walnut Silvopasture

Annual ryegrass (Lolium multiflorum Lam.) and cereal rye (Secale cereale L.) pastures can extend the grazing season in the lower Midwest. There is little data from this region on the productivity of these forages in a silvopasture system. Our objective was to determine the forage and livestock production from annual ryegrass/cereal rye in a silvopasture versus an open (non-forested) system. This study was conducted near New Franklin, MO, USA. The treatments were (1) annual ryegrass/cereal rye planted into a 6- to 7-year-old stand of pitch pine (Pinus rigida Mill.) × loblolly pine (Pinus taeda L.) hybrids and black walnut (Juglans nigra L.) (TREE treatment) and (2) the same forages planted into pastures without trees (OPEN treatment). Marshall annual ryegrass and cereal rye were no-till seeded at 22 and 67 kg ha^-1 of pure live seed, respectively, on 18 September 2000 and 30 August 2001. Treatments were replicated three times in a randomized complete block. In March 2001 and 2002, three beef heifers were assigned to each pasture and rotationally grazed until early June. Cumulative forage production and quality were measured for each treatment. Cumulative forage production in the TREE treatment was reduced by approximately 20% compared to the OPEN treatment. However, beef heifer average daily gain and gain ha^-1 were equal for both treatments. Beef producers using a annual ryegrass/cereal rye in a silvopasture system likely would not sacrifice livestock production when hybrid pine and black walnut trees are 6- to 7-year-old when compared to an open pasture. Contribution of the Missouri Agricultural Experiment Station and the University of Missouri Center for Agroforestry. This work was funded through the University of Missouri Center for Agroforestry under cooperative agreements AG-02100251 with the ARS and CR 826704-01-0 with the US EPA. The results presented are the sole responsibility of the authors and/or the University of Missouri and may not represent the policies or positions of the ARS or EPA.     

Establishment and production from thinned mature deciduous-forest silvopastures in Appalachia

Small Appalachian hill farms may benefit economically by expanding grazing lands into some of their under-utilized forested acreages. Our objective was to study the forage production potential of forest to silvopasture conversion. We thinned a white oak dominated mature second growth forested area establishing two orchardgrass-perennial ryegrass-white clover silvopasture replications for comparison with two nearby open pasture replications. After thinning trees, silvopastures were limed, fertilized and seeded. Sheep were fed hay and corn scattered across the area to facilitate removal of residual understory vegetation, disruption of litter layer and incorporation of applied materials into surface soil. Each area was divided into multiple paddocks and rotationally grazed by sheep. Two 1 m² herbage mass samples were taken from each paddock prior to animal grazing. There was no significant difference in soil moisture between silvopastures and open pastures however, there was adequate rainfall to prevent drought all 3 years. The two silvopastures received 42 and 51% of total daily incident PAR compared to the open field. Total dry forage mass yield from open pasture over the 3 years averaged 11,200 kg ha⁻¹ y⁻¹ and from silvopasture 6,640 kg ha⁻¹ y⁻¹. Silvopastures, however, had a higher PAR use efficiency (PARUE) than open pasture. Hill farms could increase grazing acreages without sacrificing all benefits from trees on the landscape by converting some areas to silvopasture.     

Potential of several native and introduced warm season grasses as components of Silvopastures in the Southeastern United States

The goal of this study was to evaluate growth and nutritional characteristics of seven warm season grasses, including several natives, produced under simulated partial shading (50%) typical of loblolly pine silvopastoral systems in the southeastern United States. Forages included ‘Tifton 9’ bahiagrass (Paspalum notatum), ‘Tifton 85’ bermudagrass (Cynodon dactylon), ‘Alamo’ switchgrass (Panicum virgatum), ‘Kaw’ Big Bluestem (Andropogon gerardii), ‘Americus’ Indiangrass (Sorghastrum nutans), ‘Harrison’ Florida paspalum (Paspalum floridanum), and Nacogdoches Eastern gamagrass (Tripsacum dactyloides). Shade affected several quality parameters, including crude protein (CP) (p?<?0.0001), acid detergent fiber (ADF) (p?=?0.0413), in vitro true digestibility (IVTD) (p?<?0.0001), and total digestible nutrients (TDN) (p?=?0.0132). Shade affected the parameters differently depending on forage type, but generally improved quality by increasing CP, IVTD, and TDN; however, shade significantly increased ADF (p?=?0.0413), though the magnitude was small (344.2 vs. 351.1 g kg^?1), and increases were isolated to big bluestem and bahiagrass. Shade reduced dry matter yield (DMY) (p?<?0.0001), and there were differences among forage species (p?<?0.0001). Bahiagrass and Florida paspalum showed the highest yields, regardless of shade treatment. Gamagrass and Florida paspalum would likely have performed better if harvested by days of rest, instead of by height. These could be viable forage species to a silvopasture system, but further studies should be conducted. Based on overall quality and yield, potential beef cattle gains, and persistence under intensive defoliation, the best forage was bahiagrass (introduced), and the best selections for native grasses were switchgrass and Indiangrass. These results indicate that there is potential for several warm season forages, including native grasses, to maintain productivity and quality under shade, which would increase the site-specific options for forage selections in this system.

     

Parasitic helminth infection in young cattle raised on silvopasture and open-pasture in Southeastern Brazil

More than 95 % of Brazil’s beef production occurs in pastures where parasitic helminths are a serious problem, potentially inhibiting an average annual weight gain of 8 kg per young animal. Current pharmaceutical treatments for parasitic infestation are expensive (estimated to cost Brazil 370 million U.S. dollars per year) and sometimes inefficient due to rapid adaptation of worms to new drugs. The presence of trees in pastures (silvopasture) has proven to be environmentally beneficial, but may favor multiplication of parasites and cause higher infestation rates compared to traditional pastures. This study in Southeastern Brazil compared the weight gains of crossbred Holstein and Gir (Zebu) bovines, 8–24 months of age, under two different grazing systems: open (tree-less) pasture, and silvopasture (primarily “Sucupiras Brancas” at 156 trees ha^?1). Despite the favorable environment for helminths, silvopasture did not increase the worm egg count per gram of animal feces and did not create overall weight or weight gain patterns different from those of animals grazing on the open pasture. The experiment took place from August 2008 to February 2009, encapsulating the transition from the dry to rainy season; a period when worm infection rates are highest, causing the greatest impact in animal development. The significance of this period aside, the results must be taken as indicative due to the short time frame and the extraneous challenges of climate variation, individual immune system resistance development, and natural helminth predator growth conditions.     

Soil compaction by grazing livestock in silvopastures as evidenced by changes in soil physical properties

Livestock, pasture, and timber trees are intimately interrelated in silvopastures. Most silvopasture research to date has focused on forage/animal/tree interactions, with less attention paid to animal/soil interactions in silvopastures. While a considerable body of work has been devoted to understanding the effects of livestock trampling on plants and soils in pastures, less has been done for livestock grazing in forests, and even less is available for silvopastures. Three replications of Douglas-fir forest, Douglas-fir/subclover pasture/sheep silvopasture, and subclover/sheep pasture were established in 1989 near Corvallis, Oregon USA. Pastures and silvopastures were grazed each spring during 1990–2001. These plots were sampled in 2002, after 11 years of grazing, and again in 2004 following 2 years without grazing. Soil in the silvopastures had 13% higher bulk density and 7% lower total porosity than those in adjacent forests in 2002. Most of the difference in total porosity was air-filled pores. Average water infiltration rate was 38% less in silvopastures than in forests, however total water stored in the top 6 cm of soil at field capacity was similar. Soil bulk density, total porosity, and air-filled pore space was similar for forests, pastures, and silvopastures after 2 years without livestock grazing. The infiltration rate of silvopasture soils in 2004 had increased to be similar to those of forests in 2002, however, forest soil infiltration rates also increased and continued to be higher than those of silvopastures. Plant production was not sensitive to changes in any of the soil parameters measured. Although livestock grazing did change soil infiltration rates, soil bulk density, and soil porosity, the effects were quickly reversed following cessation of grazing and had little detrimental effect on silvopasture forage or tree production.     

Stockpiled tall fescue and livestock performance in an early stage midwest silvopasture system

Using stockpiled forage can substantially reduce livestock feed costs over the winter. However, little is known about utilizing stockpiled forage in an early-stage silvopasture system. This study was conducted to determine if silvopasture production practices utilizing stockpiled forage influence stocker steer performance. The treatments were: (1) stockpiled forage in a non-forested pasture (OPEN) and (2) stockpiled forage in a silvopasture (TREE). Grazing began early December and ended in late February in each of 2 years. Each treatment was replicated three times in a completely randomized design. Forage nutritive value, production, and steer average daily gain (ADG) for the OPEN and TREE treatments were not significantly different as long as the areas occupied by trees was excluded from analyses. When the area occupied by trees was included, the OPEN treatment produced more (P < 0.01) forage than the TREE treatment, with the OPEN producing 3510 kg ha⁻¹ and the TREE producing 2812 kg ha⁻¹. Average daily gain (P = 0.21) was 0.41 kg for the steers in the OPEN treatment and 0.37 kg for steers in the TREE treatment. Gain per ha was significantly different (P < 0.01); the OPEN treatment produced 193 kg of animal gain and the TREE treatment produced 125 kg of animal gain. Exclusion of the area under the tree row from the analysis changed the total gain per ha for the TREE treatment to 148 kg, but was still less (P = 0.01) than the OPEN treatment.     

Foliage re-establishment of <Emphasis Type="Italic">Pinus palustris</Emphasis> Mill. saplings after spring or fall prescribed fire

Repeated fire is key to the viability of longleaf pine (Pinus palustris Mill.) ecosystems, but its acceptance as a management tool may depend on satisfactory longleaf pine growth. This is because longleaf pine establishment often has the dual-purpose of ecosystem restoration and stemwood production. Timely recovery of scorched foliage among longleaf pine seedlings and saplings supports maximum juvenile growth. We identified two means of foliage re-establishment in the growing season after prescribed fire regardless of the season of fire application. New foliage growth after spring or fall fire was correlated with un-scorched foliage biomass and the presence of lateral branches. After prescribed fire in spring, foliage biomass recovery also appeared to benefit from the mobilization of starch. The high carbon demand of foliage recovery after fall prescribed fire was associated with interruption of seasonal starch accumulation in the stem and taproot. The implication of low starch accumulation in stem and taproot tissues during the growing season after fall prescribed fire is unknown and warrants further investigation. Our results demonstrate a positive influence of residual foliage, lateral branches, and stored starch on timely foliage recovery of young longleaf pines after fire. Together with knowledge of longleaf pine development and fuel and climate conditions at the time of prescribed fire, this information will aid prescribed fire practitioners charged with maintaining longleaf pine stands of high vigor.     

Restoring longleaf pine (Pinus palustris Mill.) in loblolly pine (Pinus taeda L.) stands: Effects of restoration treatments on natural loblolly pine regeneration

Historical land use and management practices in the southeastern United States have resulted in the dominance of loblolly pine (Pinus taeda L.) on many upland sites that historically were occupied by longleaf pine (Pinus palustris Mill.). There is currently much interest in restoring high quality longleaf pine habitats to such areas, but managers may also desire the retention of some existing canopy trees to meet current conservation objectives. However, fast-growing natural loblolly pine regeneration may threaten the success of artificially regenerated longleaf pine seedlings. We evaluated the establishment and growth of natural loblolly pine regeneration following different levels of timber harvest using single-tree selection (Control (uncut, residual basal area ∼16 m²/ha), MedBA (residual basal area of ∼9 m²/ha), LowBA (residual basal area of ∼6 m²/ha), and Clearcut (complete canopy removal)) and to different positions within canopy gaps (approximately 2800 m²) created by patch cutting at two ecologically distinct sites within the longleaf pine range: Fort Benning, GA in the Middle Coastal Plain and Camp Lejeune, NC in the Lower Coastal Plain. The density of loblolly pine seedlings was much higher at Camp Lejeune than at Fort Benning at the end of the first growing season after harvesting. Following two growing seasons, there were no significant effects of canopy density or gap position on the density of loblolly pine seedlings at either site, but loblolly pine seedlings were taller on treatments with greater canopy removal. Prescribed fires applied following the second growing season killed 70.6% of loblolly pine seedlings at Fort Benning and 64.3% of seedlings at Camp Lejeune. Loblolly pine seedlings were generally less than 2 m tall, and completeness of the prescribed burns appeared more important for determining seedling survival than seedling size. Silvicultural treatments that include canopy removal, such as patch cutting or clearcuts, will increase loblolly pine seedling growth and shorten the window of opportunity for control with prescribed fire. Therefore, application of prescribed fire every 2-3 years will be critical for control of loblolly pine regeneration during restoration of longleaf pine in existing loblolly pine stands.     

Effects of shelter tubes on hardwood tree establishment in western Oregon silvopastures

Honey locust (Gleditsia triacanthos), black locust (Robinia pseudoacacia), and honey mesquite (Prosopis glandulosa) are warm season forage trees with potential to efficiently share site resources with cool season pasture plants in Pacific Northwest silvopastures. Establishment of hardwood trees can be difficult, however, because of feeding damage from wildlife and livestock. This study compared establishment and growth of trees planted in 88 cm tall solid plastic shelter tubes to 88 cm tall plastic mesh tubes, used to protect trees from animal damage. Three replications were established in May 1995 for each of the three tree species on a hill pasture near Corvallis, Oregon. Initial tree survival during the first summer and winter following planting was higher in shelter tubes than in mesh tubes. At the end of the third growing season, 58% of black locust and 94% of honey locust trees in shelter tubes were still alive compared to only 14% of black locust and 47% of honey locust in mesh tubes. Few honey mesquite trees survived regardless of tube type used. Average three-year total height growth for black locust was increased by 650% and basal diameter growth by 380% within shelter tubes, while honey locust height growth was increased by 300% and diameter growth was increased by 150% compared to trees in mesh tubes. However, shelter tube trees tended to be taller relative to their diameter and had difficulty standing upright if tubes were removed. This revised version was published online in June 2006 with corrections to the Cover Date.     

Productivity of two Douglas fir/subclover/sheep agroforests compared to pasture and forest monocultures

Resource sharing between tree and forage plant components in silvopastoral systems includes a complex set of facilitative and competitive interactions. To the extent that facilitation exceeds competition, agroforests are expected to outyield monocultures of their components. Pasture and Douglas fir (Pseudotsuga menziesii) tree production of young agroforests was compared to pasture and forest monocultures under both grid and cluster patterns of tree planting near Corvallis, Oregon, USA, during 1983–1987. The height and diameter growth of forest and agroforest trees was similar, regardless of tree planting pattern. Five-year average annual forage production was 6500, 5800, and 2800 kg ha^–1 on pasture, agroforest, and forest plots, respectively. The total cumulative 1982–1987 above-ground phytomass yield of forage plus trees was similar for pasture and conventional grid forest monocultures. The total productivity of agroforests, however, was over 30% greater than either pasture or forest components grown in monoculture. Approximately 1.6 ha (0.96 ha forest + 0.64 ha pasture) of monocultures would be needed to equal the productivity of 1 ha of agroforest.Oregon Agricultural Experiment Station Technical Paper No. 10, 825.     

Carbon, nitrogen, and phosphorus dynamics in a loblolly pine-goat silvopasture system in the Southeast USA

Agroforestry presents an opportunity to increase land productivity and improve cash flow by combining income from crop or animal production and forestry on the same land. In addition, agroforestry offers numerous environmental benefits such as increased diversity of plants and animals, nutrient recycling, erosion control, and carbon (C) sequestration. We investigated the effect of grazing and forage enhancement on total soil C (TSC), soil nitrogen (N), and phosphorus (P) dynamics in a goat (Capra aegagrus hircus)??loblolly pine (Pinus taeda L.) silvopasture system on a Kipling silt loam soil (fine, smectitic, thermic, Typic Paleudalfs) at Epes, Alabama from 2006 to 2010. In 2007, the soil in the silvopasture plots was characterized by low pH, low TSC, and was deficient in N and P. Four years after tree thinning and after 3?years of grazing in June 2010, the silvopasture plots still depicted low soil pH (<6) and TSC levels less than 20?g?kg^?1. TSC content in all the silvopasture plots in June 2010 were generally similar to levels obtained after thinning and prior to grazing in May 2007. However, soil disturbance for liming, fertilizer incorporation, and forage planting increased soil N and P levels, but resulted in lower TSC in the enhanced forage plots compared to the other treatments. Grazing increased N and P levels during the study period. Our study suggests that in the long-term, grazing without additional soil management practices can improve soil fertility through nutrient recycling and C sequestration and thereby making the goat-loblolly silvopasture system both environmentally and economically sustainable.     

Seasonal change and distribution of grass nutritive values and minerals in an open pasture surrounded by forest

The forage quality of pasture species is spatially heterogeneous. In this study, we evaluated the dynamics and spatial structure of forage quality and attempted to relate the observed variation to differences in light availability and soil properties. We compared grass forage quality in open grasslands with forest edge environments and established plots in open Japanese grasslands surrounded by secondary forests. We sampled Anthoxanthum odoratum L. and soil from each plot in May and September. These samples were analyzed to determine the crude protein (CP), neutral detergent fiber, and mineral concentrations. We calculated Moran’s I coefficients across discrete lag distance classes to construct autocorrelograms. The CP model showed a non-random pattern in May at a lag distance of 40 m, but a clustered pattern in September at a small scale. The mineral models produced similar clustered spatial patterns. We found no strong correlation between aboveground (plant) and belowground (soil) mineral contents. CP and some mineral element concentrations in the plants collected in September were significantly greater at the forest edge than in open grasslands. The forest edge therefore appears to be a valuable place to provide grazing animals with adequate protein and mineral intake during the later part of the growing season.