Use of a model to analyse the effects of continuous grazing managements on seasonal patterns of grass production

The effects of management on the seasonal patterns of production of grass swards under continuous grazing are examined using a previously described mathematical model. For continuous set-stocking it is observed that, if the leaf area index of the sward at turn-out is too low, there may be a sustained reduction in animal intake throughout the season. Under continuous variable stocking, where the sward is maintained at a constant leaf area index, animal intake is observed to follow broadly a pattern determined by the environmental conditions. It is concluded that grass production, in terms of both gross tissue production and animal intake, is not independent of management. Furthermore, no evidence is found to suggest that the seasonal patterns of production undercutting and grazing are the same.     

Frequency and severity of defoliation of grass and clover by sheep at different stocking rates

The frequency and severity of defoliation of individual grass tillers and clover plant units was studied in Lolium perenne-Trifolium repens swards grazed by sheep at stocking rates ranging from 25 to 55 sheep ha^-1 and either receiving no N fertilizer or 200 kg N ha^-1. On average, sheep at the highest stocking rate defoliated individual tillers once every 4·2 d compared with once every 9·2 d at the lowest stocking rate with the removal of 58% and 47% of the leaf length of each tiller leaf at these stocking rates. Clover plant units were defoliated once every 4·2 d at the highest stocking rate and once every 7·2 d at the lowest stocking rate with the removal of 51% of its leaves and 12% of its stolon at the high stocking rate and 42% and 4% respectively at the low stocking rate. Differences in frequency and severity of defoliation between N fertilizer treatments were smaller than between stocking rates. Grass tillers and clover plant units were both defoliated less frequently and less severely in swards fertilized with N, though the difference in defoliation frequency between fertilizer treatments decreased as stocking rate increased. Defoliation frequency was related to the length of grass leaf per tiller or number of clover leaves per plant unit, and to the number of these tillers and the herbage on offer.     

The effect of frequency and height of defoliation on the production and persistence of 'Grasslands Matua' prairie grass

The performance of prairie grass ( Bromus willdenowii Kunth) cv. Grasslands Matua was examined under mowing at five defoliation intervals (10, 20, 30, 40 and 50 d) and two defoliation heights (3 cm and 8 cm). Annual and seasonal production of both the total herbage and of Matua, and the proportion of Matua in the sward, increased with less frequent defoliation. To a lesser extent, increasing the height of defoliation increased dry matter yield. Matua tiller numbers were increased by less frequent defoliation and were greater under the 8 cm than the 3 cm defoliation height. To increase prairie grass production and persistence, rest intervals between grazings of 40 to 50 d are recommended, or alternatively, a management involving silage or hay cutting offers scope.     

Interactions between leaf lifespan and defoliation frequency in temperate and tropical pastures: a review

In continuously stocked swards or pastures the frequency at which individual tillers and individual leaves are defoliated by ruminant livestock, relative to leaf lifespan of the grass species within the sward, determines the proportion of each leaf defoliated before senescence, and hence the efficiency of harvesting of herbage. In this paper, sets of data obtained in a range of climatic conditions and with a range of grass species are used in order to document this relationship. It is shown that the frequency of defoliation of individual tillers or individual leaves is closely linked to the average stocking density used within a period of time for maintaining a steady state sward or pasture height, herbage mass or leaf area index. Consequently, any decrease in herbage growth rate should lead to a decrease in the efficiency of harvesting of herbage and then to a more than proportional decrease in total herbage consumption by ruminant livestock. These effects will be more important for grass species having short leaf lifespan than for species with long lifespan. In rotational stocking, the link between herbage growth rate and frequency of defoliation of leaves can be broken by controlling the grazing interval, so any decrease in herbage growth would not be systematically associated with a decrease in efficiency of harvesting of herbage. Rotational stocking should be more efficient than continuous stocking in low herbage production conditions, while in high herbage production systems rotational and continuous stocking would have similar efficiency. The implications of these conclusions for the management of swards and pastures to meet different objectives are discussed briefly.     

Effects of grass defoliation on the establishment and growth of slot-seeded white clover

White clover was slot-seeded into a low-fertility permanent pasture in May 1979. The effects of cutting interval (1, 2 and 4 weeks) and cutting height (3 and 7 cm), in the presence and absence of above-ground partitions, were recorded in terms of clover establishment and growth up to 15 weeks. The partitions resulted in a considerable increase in growth, stolon production and survival of clover plants, especially from 6 weeks after sowing, indicating a large effect of shoot competition from the surrounding sward. However, the effectiveness of defoliation in reducing grass competition appeared limited; close cutting did give some benefit to establishing clover but cutting frequency had little effect. In view of the limited success in reducing grass competition by cutting, further attempts at alleviating root competition are needed, possibly involving fertilizer placement and the use of grass-suppressing herbicides.     

Silage and milk production: a comparison between unwilted and wilted grass silages

Three grass silages made in sunny weather in early July from second-harvest perennial ryegrass were compared in a 16-week feeding experiment with twelve Ayrshire cows. The silages were either unwilted or wilted with and without conditioning, and had mean dry matter (DM) concentrations of 201, 261 and 272 g kg⁻¹, and in vitro DOMD concentrations of 650, 669 and 672 g kg⁻¹ DM respectively. All the silages had formic acid ('Add-F') applied at a rate of 2.6 litres t⁻¹ and were offered ad libitum plus 6 kg concentrates per cow per d. The daily intakes of silage DM were 905 kg per cow on the unwilted treatment and 9.86 and 9.65 kg on the wilted treatments with and without conditioning respectively. Daily milk yields were 171, 17.6 and 17.4 kg per cow on the unwilted, and wilted with and without conditioning treatments respectively and were not significantly different. Fat concentrations in the milk were not affected significantly by treatment, whereas the crude protein and solids-not-fat concentrations were significantly higher on the wilted than on the unwilted treatment. The efficiency of utilization of metabolizable energy for lactation was 6–7% lower with the wilted than with the unwilted silages and it is concluded that the unwilted silage was superior to the wilted silages as a feed for dairy cows.     

Silage and milk production: a comparison between three grass silages of different digestibilities

Three first-harvest grass silages made from S23 perennial ryegrass cut on 25 May, 13 June and 25 June, and termed early, medium and late silages respectively, were compared in a 16-week feeding experiment with fourteen Ayrshire cows. The early, medium and late silages had D-values of 71·2, 65·0 and 62·5 respectively. The early silage was offered alone, whereas the medium and late silages were supplemented with a concentrate containing 208 g crude protein per kg DM at rates of 2, 3 and 4 kg per 10 kg milk. Silage DM intake was 12·8 kg per cow per d on the early silage treatment, and decreased progressively as concentrate intake increased on the other silage treatments. The mean daily milk yields were 16·0 kg per cow in the early silage treatment, 17·0, 18·4 and 20·4 kg per cow in the medium silage treatments, and 16·8, 18·1 and 20·2 kg per cow in the late silage treatments on the 2-, 3- and 4-kg concentrate treatments respectively. Fat concentration in the milk was not affected significantly by treatment, whereas the CP and SNF concentrations increased progressively and significantly as supplementary feeding increased. From the relationship between milk yield and concentrate intake it was calculated that the medium and late silages required a daily concentrate supplement of 2·1 kg DM per cow to give the same daily milk yields as the early silage.     

Leaf number as a criterion for determining defoliation time for Lolium perenne: 2. Effect of defoliation frequency and height

The object of this study was to determine the importance of frequency and height of defoliation on regrowth potential of Lolium perenne. Defoliation interval was based on stage of the regrowth cycle, as indicated by leaves per tiller.
Simulated swards of Lolium perenne cv Yatsyn were grown as individual plants in a glasshouse kept at a day/night temperature of 25°C/15°C.
Treatments imposed were defoliation at 2, 5 or 12 cm residual height, and low and high water soluble carbohydrate (WSC) level obtained by varying defoliation interval, i.e. defoliating at the 1-leaf or 3-leaf stage of the regrowth cycle. Regrowth after frequent short defoliations was only 65% of the less frequently defoliated plants taken over the full regrowth cycle. This was associated with a lower stubble WSC content (2·15 vs 17·5% in stubble) and a twenty-seven-fold difference in the amount of WSC in the stubble per plant. This difference in total WSC was a combined effect of more and heavier tillers and higher WSC content in stubble of plants defoliated less frequently at the end of the regrowth cycle. The regrowth of plants with WSC levels depleted by frequent defoliation when defoliated at 2 cm was significantly below that of those defoliated at 5 and 12 cm.
The results indicate the desirability of defoliating plants at the 3-leaf stage of the regrowth cycle. This not only allows the full regrowth potential to be expressed in that growth cycle, but also in the next cycle, by allowing the replenishment of WSC reserves and optimizing tiller status. The potential to regrow appears then to be based more on the total amount of WSC than the proportion of WSC in stubble.     

Silage and milk production: a comparison between grass silages of different chop lengths and digestibilities

Four grass silages of two chop lengths and two D-values were made from S23 perennial ryegrass, and compared in a feeding experiment with lactating cows and in an eating-behaviour study with non-lactating cows. The median chop lengths were 11·6 and 11·5 mm in the short silages and 18·2 and 19·5 mm in the medium silages. Chop length had negligible effects on D-value and on pH with values of 3·81 and 3·94 on the short and medium silages respectively.
Silage DM intakes were higher on the short than on the medium-chop silages in both experiments except where the high D-value silage was supplemented with concentrates. Milk yields were not affected significantly by chop length with daily values of 19·3 and 19·7 kg per cow on the short and medium treatments respectively with the high D-value silage, and 18·2 and 18·3 kg per cow with the low D-value silage. Chop length had only small and non-significant effects on milk composition.
Eating times expressed as min per kg DM were significantly lower on the short than on the medium-chop silages.
It is concluded that the small differences between the short- and the medium-chop silages were of no economic importance.     

Interaction between water-soluble carbohydrate reserves and defoliation severity on the regrowth of perennial ryegrass (Lolium perenne L.)-dominant swards

A field-study was undertaken in Hamilton, New Zealand to determine if there was an interaction between water-soluble carbohydrate (WSC) reserve content and defoliation severity on the regrowth of perennial ryegrass-dominant swards during winter. Perennial ryegrass plants with either low or high WSC content were obtained by varying the defoliation frequency. At the third defoliation at the one-leaf stage and at the first defoliation at the three-leaf stage (harvest H₁), swards were mown with a rotary lawnmower to residual stubble heights of 20, 40 or 60 mm. All swards were then allowed to regrow to the three-leaf stage before again defoliating to their treatment residual stubble heights (H₂). Frequently defoliated plants contained proportionately between 0·37 and 0·48 less WSC in the stubble after defoliation, depending on the severity of defoliation. There was no interaction between WSC content and defoliation severity for herbage regrowth between harvests H₁ and H₂. Herbage regrowth was lower from swards containing low WSC plants compared with high WSC plants (2279 vs. 2007 kg DM ha⁻¹). Furthermore, swards defoliated to 20 or 40 mm had greater herbage regrowth compared with those defoliated to 60 mm (2266, 2249 and 1914 kg DM ha⁻¹ for swards defoliated to residual stubble heights of 20, 40 and 60 mm, respectively). Regrowth of perennial ryegrass was positively correlated with post-defoliation stubble WSC content within defoliation severity treatment, implying that WSC contributed to the defoliation frequency-derived difference in herbage yield. However, the effect of defoliation severity on herbage regrowth was not associated with post-defoliation stubble WSC content.     

A comparison of grass/white clover and grass silages offered to dairy cows as the sole feed

Two diets were compared: perennial ryegrass ( Lolium perenne L.) silage and perennial ryegrass/white clover ( Trifolium repens L.) silage, in each case fed ad libitum , without supplementation, to lactating dairy cows. The comparison was made with silages cut on each of four dates. The crops were wilted to a dry matter content of at least 256g kg^-1 and no additives were used.
The grass/clover silages were well preserved at all four cuts. The grass was well preserved at three cuts, but, at a May cut, the grass silage was less well preserved and less digestible than the grass/clover silage. When the May silages were fed, milk yield was higher with grass/clover than with grass. Taking the experiment as a whole, however, milk yield and composition were similar on the two diets. The grass/clover silages had a lower proportion of cell wall and their intake by dairy cows was consistently higher than that of the grass silages.
It is concluded that perennial ryegrass/white clover crops can be ensiled successfully and fed successfully, with high intake, to lactating dairy cows, but it should not be assumed that cows will give more milk than when fed equivalent all-grass silage.     

Effects of an autumn defoliation on overwintering, spring growth and yield of a white clover/grass sward

An established sward of binary mixtures of meadow fescue (Festuca pratensis) and white clover (Trifolium repens) (either AberHerald, Grasslands Huia or Sandra) was subjected to (A) no further defoliation, (B) a defoliation in late September or (C) a defoliation in late October after four harvests had been taken during the grazing season. About a tonne of dry matter (DM) was removed by the autumn defoliations. There were two levels of nitrogen application in spring, either 0 or 90 kg ha^?1. The development of grass and clover morphology and population sizes from early autumn until the first harvest the following year was followed by regular sampling of the above-ground material. Stolons were analysed for total non-structural carbohydrates (TNCs), and the temperature at stolon level was continuously recorded. There were no interactions between autumn defoliation, clover cultivar or nitrogen treatments on any of the parameters studied. White clover growing-point numbers and stolon morphological characteristics were reduced in size during the winter and did not recover during the spring. A defoliation in late September resulted in the greatest reduction, whereas there were no differences between the other two treatments. The grass tiller population increased from early autumn until the last sampling occasion in May, but both autumn defoliations resulted in a smaller increase. Defoliation in late September had the greatest impact. The TNC content of white clover stolons fell from about 350 g kg^?1 to 150 g kg^?1 DM from late autumn until late April. There were small differences between the treatments, but a defoliation in late September resulted in a significantly lower level in late autumn. The temperature amplitude at stolon level was consistently greater in plots defoliated in late September. Total DM harvested in spring was 4367, 2564 and 3536 kg ha^?1, of which 388, 352 and 460 kg ha^?1 was white clover, from treatments A, B and C respectively. It is concluded that an autumn defoliation may affect the overwintering of white clover negatively, but that the effect on the grass may be even more detrimental.     

Some effects of fertilizer and frequency of defoliation on the botanical composition and yield of permanent grassland

Small plots of permanent pasture containing 50% Agrostis capillaris-Festuca rubra were defoliated frequently or infrequently during the growing seasons in 1978–80 and fertilized with 133, 125 and 125 kg N, P₂O₅ and K₂O respectively ha⁻¹ a ⁻¹ or not fertilized during 1978–81. Yields of herbage were assessed during 1979–80 and botanical composition assessed on three occasions in 1978, once in April 1979 and once in 1982.
Fertilizer application increased annual dry matter (DM) harvested from frequently cut plots by 40%, 53% and 65% in 1978, 1979 and 1980 respectively, and on infrequently cut plots it doubled hay DM harvested in 1978 and increased total DM harvested by 126% and 186% in 1979 and 1980 respectively. Infrequent cutting and fertilizer both decreased the proportion of A. capillaris but only fertilizer decreased the proportion of F. rubra. Fertilizer greatly increased the proportion of Holcus lanatus especially where plots had been cut frequently and of Alopecurus pratensis where they had been cut infrequently. It decreased the proportion of Luzula campestris. Infrequent cutting, especially with fertilizer, discouraged Cerastium fontanum ssp. glabrescens and Trifolium repens but encouraged Rumex acetosa.     

Effect of defoliation frequency and cutting height on growth,dry‐matter yield and nutritive value of Napier grass (Pennisetum purpureum (L.) Schumach)

Growth characteristics, dry‐matter (DM) yield, chemical components and in vitro dry‐matter digestibility (IVDMD) of Napier grass were studied in a randomized complete block design with three replications at Haramaya University, Ethiopia. The treatments were three defoliation frequencies (60, 90 and 120 d after Napier grass establishment) and five cutting heights (5, 10, 15, 20 and 25 cm above ground level). Except for leaf‐to‐stem ratio, all growth characteristics increased with decreasing frequency of defoliation. Similarly, DM yield of Napier grass increased as the frequency of defoliation decreased. Total ash, crude protein (CP), hemicellulose and IVDMD contents of Napier grass increased with increasing frequency of defoliation, whereas DM, acid detergent fibre, acid detergent lignin and cellulose contents increased with a decrease in defoliation frequency. In contrast, cutting heights had no significant effect on most chemical components and IVDMD contents, although total ash and CP decreased as the height of cutting increased. The CP yield per cut and per day was highest at an intermediate defoliation frequency. The results of this study indicated that Napier grass could be categorized under medium‐ to high‐quality herbage if defoliated at 90 d after establishment in the mid and highlands of Ethiopia.     

Simulation of allocation of accumulated biomass to leaf and stem in a grass growth model

To model variations in leaf/stem ratio in connection with studies of field-drying of grass, account was taken of the effects that the stage of morphological development of grass has on the allocation of biomass to plant components, as well as on duration of growth and peak yield. A comparison of experimental data and corresponding daily weather data suggested that it is more satisfactory to express the stages of grass development in terms of degree-days than time alone. On the basis of one set of weekly measured data, the ratio of green leaf weight to aboveground biomass was found to be a close fit to a positive linear relationship early in the growth period (durïng expansion of the first leaf on each tiller), followed by negative exponential relationship (during expansion of subsequent leaves), in relation to relative development expressed as growing degree-days. Similar trends, but with some quite large variations in absolute values because of unknown factors, were found in two other datasets consisting of a few measurements only.     

Silage and milk production: a comparison between three rates of high-protein concentrate supplementation of grass silages of two digestibilities

Grass silages made from first-harvest perennial ryegrass in mid-May and early-June and termed high-D and low-D respectively had mean DOMD values of 694 and 633 g kg⁻¹ and were offered ad libitum to twelve Ayrshire cows in a 16-week feeding experiment using a cyclic changeover design. The silages were supplemented with a concentrate containing 363g CP per kg DM at daily rates of 1.5, 3.0 and 4.5 kg per cow. The average daily intakes of silage DM were 12.6 and 11.1 kg per cow on the high- and low-D silage treatments respectively, and were not significantly different on the three concentrate treatments. The mean daily milk yields were 171, 18.6 and 21.0 kg per cow on the low-D silage treatments, and 19.6, 21.2 and 22.8 kg per cow on the high-D silage treatments at the 1.5, 3.0 and 4.5 kg rates of concentrate feeding respectively. Fat and lactose concentrations in the milk were not affected significantly by treatment, whereas the CP and SNF concentrations increased progressively and significantly as the rate of supplementary feeding increased. It is concluded that a high-protein concentrate supplement allows silage to make the maximum contribution in the diet, and gives a high response in terms of milk yield and composition.     

Analysis of adaptation of grass/legume mixtures to Italian alpine and subalpine zones through an additive main effects and multiplicative interaction model

Fourteen grass/legume mixtures, mostly including white clover or lucerne, in association with a variable number of other forage species were grown at six Italian alpine and subalpine locations under a mowing regime. The response of the mixtures in terms of dry-matter (DM) yield and visually estimated legume percentage over the second and third years of crop cycle was investigated through an additive main effects and multiplicative interaction (AMMI) analytical model. Locations and mixture-location interaction effects were significant whereas mixture was not. The interaction effect was very large, and the first two interaction principal component (PC) axes were significant at P<0·001. PC 1 accounted for 88% and 92% of the interaction variation for DM yield and legume percentage respectively. Both for sites and for mixtures, the ordination on PC 1 for these variables was very similar, indicating that the interaction effects for DM yield tended to parallel those for legume percentage. The adaptive response of the mixtures mainly depended on the response of their legume components. White clover was specifically adapted to locations with high soil clay content, probably through better tolerance to the associated severe waterlogging. Lucerne performed relatively better at high-yielding, favourable sites, probably because of its greater competitive ability than associated grasses under conditions that emphasize competitive effects. Minor interaction effects were related to the response of cocksfoot and timothy, the latter being indispensable for high DM yield of mixtures at high-elevation, cool-summer sites. No clear advantage of having a high number of components in the mixture was apparent, and it was concluded that the association of timothy and cocksfoot with one legume should be adequate for most situations in the given region.     

Plant growth and survival of five introduced and two native/naturalized perennial grass genotypes exposed to two defoliation managements in arid Argentina

The field performance of the native Pappophorum vaginatum, the naturalized Eragrostis curvula and various cultivars of the introduced Achnatherum hymenoides and Leymus cinereus was evaluated as potential forage resources in rangelands of arid Argentina during the warm seasons of 2007/2008 and 2008/2009. Plants of these grass species, obtained from seeds, were transplanted to the field in 2006, when they were 1 year old. During the study years, there were two defoliation managements: plants of all study genotypes either remained undefoliated (controls) or were defoliated twice a year during spring at 5 cm stubble height. Despite tiller number being lower (P < 0·05) on defoliated than on undefoliated plants, and total leaf length per unit basal area being similar (P > 0·05) between defoliation managements by mid‐spring, there were no differences (P > 0·05) in dry weight production between defoliated and undefoliated plants in all genotypes at the end of the second growing season. Plants of one or more of the introduced genotypes showed a similar (P > 0·05) or greater (P < 0·05), but not lower, tiller number per plant and per square centimetre, daughter tiller production, total leaf length and dry weight production per unit basal area than the native species at the end of the first and/or second growing seasons. These morphological variables were similar (P > 0·05) or greater (P < 0·05) in the native than in the naturalized genotype. Plant survival, however, was lower (P < 0·05, overall mean = 20%) in the introduced than in the native (>70%) or naturalized (>39%) genotypes at the end of the first or second growing seasons.     

二甲四氯钠与草甘膦混用防除棉田节节草的效果

节节草是棉田入侵性难除杂草,一般除草剂难以防治,作者采用草甘膦与二甲四氯钠混合施用,其防治效果可达95％以上,药效期可保持在30天以上,比人工除草或草甘膦、二甲四氯钠盐的单一防除效果要好,且防效期更长。     

Introduction of seasonal and spatial specification to grass production and grassland use in a dairy farm model

A linear programming model of a dairy farm, in which grass production and grassland use was originally modelled using one growing period and one area of land, is extended. To include seasonal and spatial elements of variation not allowed for in the original model, the model was modified to reflect three growing periods in the season and then to reflect management of two separate areas of land. The resulting three models were used to assess farm physical planning, economic results and nutrient balances for an average Dutch dairy farm in a situation without and with levies on nutrient surpluses. It is concluded that incorporating seasonal and spatial elements provides greater insight into grassland management. Seasonal model specification does not result in significant changes in farm physical planning, income and nutrient surpluses. Spatial specification has a larger impact if one of the two areas cannot be grazed by dairy cattle.