Maize and sesbania production in relay cropping at three landscape positions in Malawi

The relay cropping of sesbania (Sesbania sesban) — a N₂-fixing legume — with maize (Zea mays) has been proposed as a strategy to increase soil fertility and food production in densely populated areas in southern Africa. We determined the production of relay-cropped maize and sesbania at three landscape positions under researcher-designed and farmer-managed conditions in southern Malawi. Three landscape positions (dambo valley or bottomland, dambo margin with < 12% slope, and steep slopes with > 12% slope) were examined in factorial combination with N sources for maize (no added N, relay-cropped sesbania, and calcium ammonium nitrate fertilizer). Relay cropping of sesbania with maize increased maize grain yield, as compared to unfertilized sole maize in two of three years. Split application of 96 kg N ha^–1 as N fertilizer, however, was more effective than sesbania in increasing maize yields. Survival of sesbania seedlings and biomass production of sesbania were greater in the dambo valley and dambo margin than on steep slopes. Maize yields tended to be lower on steep slopes than in the dambo valley and dambo margin areas. Biomass production of sesbania and hence the potential benefits of intercropping sesbania with maize appear greater in the dambo valleys and dambo margins than on steep slopes.This revised version was published online in November 2005 with corrections to the Cover Date.     

Soil nitrate dynamics in relation to nitrogen source and landscape position in Malawi

Nitrogen is normally the nutrient most limiting production of maize (Zea mays) — the main staple food crop — in southern Africa. We conducted a field study to determine the effect of N sources on soil nitrate dynamics at three landscape positions in farmers' fields in southern Malawi. The landscape positions were dambo valley or bottomland, dambo margin, and steep slopes. The N sources were calcium ammonium nitrate fertilizer applied at 120 kg N ha^–1, biomass from Sesbania sesban, and no added N. Sesbania biomass was produced in situ in the previous season from sesbania relay cropped with maize. Nitrate in the topsoil (0 to 15 cm depth) increased to 85 days after maize planting (mean = 48 kg N ha^–1) and then decreased markedly. Application of N fertilizer and sesbania biomass increased soil nitrate, and nitrate-N in topsoil correlated positively with amount of incorporated sesbania biomass. The strongest correlation between sesbania biomass added before maize planting and topsoil nitrate was observed at 85 days after maize planting. This suggests that the sesbania biomass (mean N content = 2.3%) mineralized slowly. Inorganic N accumulated in the subsoil at the end of the maize cropping season when N fertilizer and sesbania were applied. This study demonstrated the challenges associated with moderate quality organic N sources produced in smallholder farmer's fields. Soil nitrate levels indicated that N was released by sesbania residues in the first year of incorporation, but relay cropping of sesbania with maize may need to be supplemented with appropriately timed application of N fertilizer.This revised version was published online in November 2005 with corrections to the Cover Date.     

Biomass production of tree fallows and their residual effect on maize in Zimbabwe

The rotation of maize (Zea mays) with fast-growing, N₂-fixing trees (improved fallows) can increase soil fertility and crop yields on N-deficient soils. There is little predictive understanding on the magnitude and duration of residual effects of improved fallows on maize yield. Our objectives were to determine the effect of fallow species and duration on biomass production and to relate biomass produced during the fallow to residual effects on maize. The study was conducted on an N-deficient, sandy loam (Alfisol) under unimodal rainfall conditions in Zimbabwe. Three fallow species — Acacia angustissima, pigeonpea (Cajanus cajan), and Sesbania sesban — of one-, two-, and three-year duration were followed by three seasons of maize. Pigeonpea and acacia produced more fallow biomass than sesbania. The regrowth of acacia during post-fallow maize cropping provided an annual input of biomass to maize. Grain yields for the first unfertilized maize crop after the fallows were higher following sesbania (mean = 4.2 Mg ha^–1) than acacia (mean = 2.6 Mg ha^–1). The increased yield of the first maize crop following sesbania was directly related to leaf biomass of sesbania at the end of the fallow. Nitrogen fertilizer did not increase yield of the first maize crop following one- and two-year sesbania fallows, but it increased yield following acacia fallows. Nitrogen fertilizer supplementation was not required for the first maize crop after sesbania, which produced high-quality biomass. For acacia, which produced low-quality biomass and regrew after cutting, N fertilizer increased yield of the first post-fallow maize crop, but it had little benefit on yield of the third post-fallow maize crop.This revised version was published online in November 2005 with corrections to the Cover Date.     

The root-knot nematode problem in sesbania fallows and scope for managing it in western Kenya

Sesbania (Sesbania sesban), which is promoted as a short-duration-fallow species in eastern and southern Africa, is an alternative host to root-knot nematode (Meloidogyne spp.). Therefore, it is important to know the extent of sesbania infection by the nematode on farms and its effects to crops grown in rotation with sesbania. A survey of 14 farms in western Kenya indicated that sesbania was infected by root-knot nematode in all the farms. The potential to build up the nematode population was much greater on farms where maize (Zea mays) was previously intercropped with a susceptible bean (Phaseolus vulgaris). Screening of 30 single plant accessions of four sesbania provenances (Kakamega, Siaya, Kisumu, and Kisii) from western Kenya in pots indicated limited scope for selecting material resistant to the nematode. All four provenances were infested by the nematode, and they increased its population in soil and root. A rangewide collection of sesbania germplasm needs to be screened to determine the scope of finding nematode-resistant material. Transplanted seedlings produced in heat-sterilized, nematode-free soil showed less nematode infection than direct-seeded plants. However, both direct seeding and transplanting seemed to increase nematode population similarly. A number of Crotalaria spp. that were found resistant to nematodes can be considered as alternatives to sesbania-planted fallows, but further field-scale testing is necessary to select appropriate species based on biomass production. Among other species suitable for short-duration fallows, only pigeonpea (Cajanus cajan) and senna (Senna siamea) showed promise. Species such as neem (Azadirachta indica), casuarina (Casuarina junghuhniana), and grevillea (Grevillea robusta) that are suitable for boundary plantings were free from the root-knot nematode.This revised version was published online in November 2005 with corrections to the Cover Date.     

Substitution of nitrogen requirement of maize through leaf biomass of Leucaena leucocephala: agronomic and economic considerations

In a 3-year field study, the effects of substitution of nitrogen requirement of maize through Leucaena leaves were studied on runoff, soil loss, maize and wheat yield and economic returns. The treatments were (1) 80 kg N ha^-;1 all through Leucaena leaves (80 L), (2) 40 kg N through Leucaena leaves + 40 kg N ha^-;1 through fertilizer (40 L + 40 F), (3) 20 kg N through Leucaena leaves + 60 kg N ha^-;1 through fertilizer (20 L + 60 F), (4) 80 kg N ha^-;1 all through fertilizer (80 F), and (5) control (No fertilizer). Green Leucaena leaf biomass (containing 3.3% N on dry basis) was incorporated every year in 15 cm top soil two weeks before sowing of summer maize.Other treatments being almost equal, runoff was reduced marginally in treatment 20 L + 60 F which was attributed to better crop growth in this treatment. Mean minimum soil loss (6.202 t ha^-;1) also occurred in treatment 20 L + 60 F. Soil loss in 80 L was 13% less than in 80 F. Maize yield was at par in 80 L and 80 F. However, mean maximum yield of maize was obtained with 20 L + 60 F.Residual effect of incorporation of Leucaena leaves to maize crop was observed on wheat yield. The mean yield differences were statistically at par in all the treatments except control. The total mean net returns were statistically at par in 80 L and 80 F. However, significantly higher mean net returns (Rs 6811 ha^-;1; one US$ = Rs30) were obtained with 20 L + 60 F. Substitution of N through Leucaena leaves even in small quantity may be helpful to small holders, particularly where chemical fertilizers are in short supply or too expensive.     

Leucaena + maize alley cropping in Malawi. Part 1: Effects of N, P, and leaf application on maize yields and soil properties

Yields under alley cropping might be improved if the most limiting nutrients not adequately supplied or cycled by the leaves could be added as an inorganic fertilizer supplement. Three historic leaf management strategies had been in effect for 3 years ina Leucaena leucocephala alley cropping trial on the Lilongwe Plain of central Malawi : 1) leaves returned; 2) leaves removed; and 3) leaves removed, with 100 kg inorganic N ha⁻¹ added. An initial soil analysis showed P status to be suboptimal under all strategies. A confounded 3⁴ factorial experiment was conducted with the following treatments: leaf management strategy (as above), N fertilizer rate (0, 30, and 60 kg N ha⁻¹), P fertilizer rate (0, 18, and 35 kg P ha⁻¹), and maize population (14,800, 29,600, and 44,400 plants ha⁻¹). Both N and P were yield limiting, and interacted positively to improve yields. The addition of 30 kg N and 18 kg P ha⁻¹ improved yields similarly under all leaf management strategies by an average of 2440 kg ha⁻¹. Increasing the rates to 60 kg N and 35 kg P ha⁻¹ improved yields an additional 1990 kg ha⁻¹ in the ‘leaves returned’ and leaves removed + N’ strategies, but did not improve yields under the ‘leaves removed’ strategy. Lower yields were related to lack of P response at the highest P rate in this treatment, which may have induced Zn deficiency. Plots receiving leaves had higher organic C, total N, pH, exchangeable Ca, Mg, K, and S, and lower C/N ratios in the 0–15 cm soil layer than did plots where leaves had been removed. Leaf removal with N addition was similar to leaf removal alone for all soil factors measured except for organic C and total N, which were higher where N had been added. The results show that N and P were the primary yield-limiting nutrients. Historic N application maintained the soil's ability to respond to N and P on par with leaf additions.     

Abundance of insect pests and their effects on biomass yields of single vs. multi-species planted fallows

Indigenous and exotic leguminous shrubs that are promising for planted fallow for soil fertility replenishment in east and southern Africa have been found to harbour many herbivorous insects, giving suspicion that widespread adoption of fallow systems may aggravate insect pests. Studies were conducted on farms in western Kenya from 1999 to 2001 to monitor the abundance of herbivorous insects and assess their effects on biomass yields of pure and mixed fallows. The treatments tested were single and two-species mixtures of Tephrosia vogelii, Sesbania sesban and Crotalaria grahamiana and a natural fallow in a split plot design, with the fallow systems in the main plots and protection vs. no protection against insects in sub-plots spread over six farms. Eighteen insect species belonging to seven orders and 14 families were identified as pests of␣the fallows with varied abundance and infestation level across the sites. While Hilda patruelis and Amphicallia pactolicus were most damaging to C. grahamiana, Mesoplatys ochroptera was detrimental to S.␣sesban. T. vogelii hosted fewer insects than others. Nevertheless the pest infestation did not cause significant biomass yield reduction during the study period. Pest attack was generally greater in villages that had been testing the planted fallows for some years compared with villages that took up the fallows recently. This indicates the potential for increased pest infestation with increased adoption of the system by farmers. Multi-species fallows did not indicate any advantage over single species fallows in terms of either reduced pest incidence or increased biomass production.     

Farmer assessment and economic evaluation of shrub fallows in the Humid Lowlands of Cameroon

Food crop production in highly populated areas around major cities of the humid lowlands of Cameroon is highly dependent on a fallow system (two–four years duration) mainly of Chromolaena odorata. Where such fallows have been in use for some time, problems of soil fertility with declining crop yields and higher incidence of weeds were reported. Although improved fallows have been widely adopted in sub-humid zones, there is no evidence of successful adoption of agroforestry-based technologies for soil fertility improvement in the humid forest areas. In response, ICRAF has developed a short fallow system with Cajanus cajan for soil fertility improvement in the humid lowlands of West Africa. Farmers' response to these cajanus fallows is positive. Benefits reported are higher crop yields after cajanus fallows compared to natural fallows, clearing of cajanus is easier and the shrubs shade out the weeds. Women particularly appreciate the technology for its low labour demand and for the fact that these shrubs can be planted on land with less secure tenure. Economic analysis of cajanus fallows compared to natural fallow over six years shows that cajanus fallows are profitable under most tested scenarios, both in terms of returns to land and to labour. It seems that improved fallows with Cajanus cajan are a good response to shortening natural fallows for households in the humid lowlands of Cameroon with land constraints. However, wider dissemination of the technology requires a targeted extension approach and adequate seed supply strategies, which should be based on joint efforts between farmers, extension services and research.This revised version was published online in November 2005 with corrections to the Cover Date.     

Effect on maize growth of the interaction between increased nitrogen availability and competition with trees in alley cropping

Maize growing next toErythrina hedgerows had 44% lower biomass (p<0.01) and 35% lower N content (p<0.1) than maize growing in the middle of the alleys. Maize growing next toGliricidia hedgerows had the same biomass but 56% higher N content (p<0.1) than maize growing in the middle of the alleys. However these differences did not develop until 2 months after sowing of the maize.Spatial variability in soil nitrogen mineralization and mulch nitrogen release did not explain any of the differences in growth or N uptake of the maize with respect to distance from the trees. It is hypothesized that the slower growth of the maize next to theErythrina trees after 2 months is due to increasing light and/or nutrient competition from the trees as the trees recover from pollarding. The apparent lack of competition fromGlirigidia may be due to different rates of regrowth or different shoot and root architecture.A theoretical model is described demonstrating that if a crop is to take advantage of the higher nutrient availability under alley cropping it must complete the major part of its growth before the trees recover significantly from pollarding, and start competing strongly with the crop.     

Soil water dynamics in cropping systems containing <Emphasis Type="Italic">Gliricidia sepium</Emphasis>, pigeonpea and maize in southern Malawi

The water dynamics of cropping systems containing mixtures of Gliricidia sepium (Jacq.) Walp trees with maize (Zea mays L.) and/or pigeonpea (Cajanus cajan L.) were examined during three consecutive cropping seasons. The trees were pruned before and during each cropping season, but were left unpruned after harvesting the maize; prunings were returned to the cropping area in all agroforestry systems to provide green leaf manure. The hypothesis was that regular severe pruning of the trees would minimise competition with crops for soil moisture and enhance their growth by providing additional nutrients. Neutron probe measurements were used to determine spatial and temporal changes in soil moisture content during the 1997/98, 1998/99 and 1999/00 cropping seasons for various cropping systems. These included gliricidia intercropped with maize, with and without pigeonpea, a maize + pigeonpea intercrop, sole maize, sole pigeonpea and sole gliricidia. Soil water content was measured to a depth of 150 cm in all treatments at 4–6 week intervals during the main cropping season and less frequently at other times. Competition for water was apparently not a critical factor in determining crop performance as rainfall exceeded potential evaporation during the cropping season in all years. The distribution of water in the soil profile was generally comparable in all cropping systems, implying there was no spatial complementarity in water abstraction by tree and crop roots. However, available soil water content at the beginning of the cropping season was generally lower in the tree-based systems, suggesting that the trees continued to deplete available soil water during the dry season. The results show that, under rainfall conditions typical of southern Malawi, the soil profile contains sufficient stored water during the dry season (ca. 75–125 mm) to support the growth of gliricidia and pigeonpea, and that gliricidia trees pruned before and during the cropping season did not deleteriously compete for water with associated crops. Water use efficiency also appeared to be higher in the tree-based systems than in the sole maize and maize + pigeonpea treatments, subject to the proviso that the calculations were based on changes in soil water content rather than absolute measurements of water uptake by the trees and crops.     

Tephrosia species and provenances for improved fallows in southern Africa

Seeking an alternative to Sesbania spp. tree fallows, a Tephrosia species and provenance trial was conducted at Msekera Research Station, Chipata (Zambia) to evaluate eleven Tephrosia vogelii and three Tephrosia candida provenances. They were tested for biomass production, quality of biomass, resistance to root-knot nematodes, nitrogen release, and for their effects on soil nitrogen dynamics. At the end of 1.5 years, the T. candida provenances 02970, 02971 and 02972 from Madagascar produced two times greater amount of aboveground biomass than the T. vogelii provenances. There was little variability among the T. vogelii provenances in terms of litter and biomass production. Weed growth was significantly greater under T. vogelii than T. candida provenances. While Tephrosia vogelii provenance 98/02 from Zambia and T. candida 02972 were highly tolerant to the Meloidogyne incognita nematodes, T. vogelii provenances 02977, 98/03, 02973 from Kenya, Zambia and Malawi, respectively, were highly susceptible to the nematodes. The Tephrosia species and provenances showed a wide variability in terms of N, lignin and polyphenol concentration in their foliage. Mineralization of N in the foliage of T. candida provenances 02970 and 02971 and T. vogelii provenances 98/04 and 02974 from Malawi occurred rapidly within 14 weeks of incubation. At the end of the 2-year growth period, there was significantly greater total inorganic N under T. candida provenance 02972 (12.5 mg kg⁻¹) than T. vogelii (5 mg kg⁻¹) provenance Mungwi 98/02. Maize (Zea mays L.) yields after T. candida provenances were greater than those after T. vogelii provenances. Further testing of the most promising provenances is needed for their effects on subsequent maize yields under a range of farm conditions. This revised version was published online in June 2006 with corrections to the Cover Date.     

Competition for light between hedgerows and maize in an alley cropping system in Hawaii, USA

Successful agroforestry systems depend on minimizing tree-cropcompetition. In this study, field experiments and a simulation model were usedto distinguish between tree-crop competition for light and belowgroundcompetition in an alley cropping system. Maize (Zea maysL.) was harvested periodically in three treatments: between vertical barriers ofshade cloth, hedgerows of Flemingia macrophylla (Willd.)Merr., and sole maize. Radiation intercepted by the maize was calculated using asimulation model based on measured values for direct and diffuse light, hedgerowdimensions and leaf area, and solar trajectory. Radiation use efficiency wascalculated as biomass production per unit of intercepted radiation. Maizebiomass and yield in both the alley crop and the shade cloth treatment weregreatest in the center of the alleys. Grain yield between hedgerows was 3.5Mg ha⁻¹ (averaged across the alley), significantlyless than in the shade cloth (7.4 Mg ha⁻¹) or thesole maize (7.7 Mg ha⁻¹) treatments. Lightintercepted by the maize in the alley crop was about half that intercepted bythe maize in the sole crop. The shade cloth intercepted less light than thehedgerows because it did not have an appreciable width. Radiation use efficiencyin the three treatments was 0.75 g mol⁻¹ PAR anddid not differ significantly among treatments. Tree-crop competition wasoverwhelmingly for light. This revised version was published online in June 2006 with corrections to the Cover Date.     

The effects of between-row (alley widths) and within-row spacings ofGliricidia sepium on alley-cropped maize in Sierra Leone

The effects of spacings between hedgerows (alley widths) and the spacings of trees within hedgerows ofGliricidia sepium on growth and grain yield of maize were investigated at Senehun in southern Sierra Leone. Four between-row spacings (2, 4, 6 and 8 m) were combined with three within-row spacings (0.25, 0.50 and 1.00 m) in a split block design. Maize, at densities of 20,000, 40,000 and 53,333 plants ha^–1, was established in the alleys and also as pure crops. N, P and K fertilizers were applied to all plots before pruning of the trees began. When pruning started, only the pure maize plots received fertilizer; prunings from the hedgerows were returned to the appropriate alleys in the other plots.Plots with the highest maize populations consistently gave the best yields before pruning started, but lower populations gave improved yields after pruning. Yields of maize increased with increasing alley widths before the start of pruning, after which the narrower alleys of 2 and 4 m outyielded the wider ones by almost double, probably because of the large amount of nutrients applied in prunings. Lack of light limited grain yields before the start of pruning, when there was some shading by the hedgerows. Alleys of 2–4 m wide, planted no closer than 0.50 m within rows, resulted in more than twice the yields of maize than in the 8-m alleys planted at 0.25 m within rows, once the hedgerows were well established and were being managed.     

Modelling trees' effect on maize in the Grevillea Robusta + maize system in Central Kenya

In the Central Highlands on Kenya, Grevillearobusta (A. Cunn.) trees are commonly planted on maize (Zeamays L.) fields. This type of production system covers about 750 000ha in the highland area around Mt Kenya. Knowing the influence oftrees on maize yield would help to regulate the density and size distribution ofthe tree cover. The spatial distribution of trees varies from rather uniform toextremely aggregated patterns, calling for a spatial modelling approach. Thisstudy employed three different spatial competition index types to model theeffect of trees on the maize yield at a given location. The maize yield wasexpressed as a function of competition index, which was calculated from thediameters and distances of trees. The data were collected from 14 maize fields,and included measurements on 857 trees and 2514 one-m² maize plots. The models indicated that high competition by treesdecreases maize yields considerably, but at the field level the decrease issmall with normal tree stockings (about 200 trees per hectare). Characteristicto the models was low degree of explained variance, which was partly due to highsampling error in maize yield measurement and low variation in competition(places with high competition by trees were seldom planted with maize). One ofthe maize yield models was combined with an earlier simulation program forG. robusta. The simulator allows one to predict thetemporal development of any G. robusta – maize fieldin Central Kenya. This revised version was published online in June 2006 with corrections to the Cover Date.     

Nitrate-N dynamics following improved fallows and maize root development in a Zimbabwean sandy clay loam

Improved or planted fallows using fast-growing leguminous trees are capable of accumulating large amounts of N through biological N₂-fixation and subsoil N capture. During the fallow phase, the cycling of nutrients is largely efficient. However, there are few estimates of the fate of added N during the cropping phase, after the 'safety net' of fallow-tree roots is removed. Nitrate-N at the end of the fallow phase, which is pre-season to the subsequent crop, was monitored in seven land use systems in successive 20-cm soil layers to 120 cm depth at Domboshawa, Zimbabwe in October 2000. Thereafter, nitrate-N dynamics was monitored during cropping phase until April 2001 at 2-week intervals in plots that had previously 2-year planted fallows of Acacia angustissima and Sesbania sesban, and in a continuous maize control. Pre-season nitrate concentrations below 60 cm soil depth were <3 kg N ha⁻¹ layer⁻¹ for S. sesban, A. angustissima, Cajanus cajan and natural woodland compared with the maize (Zea mays L.) control, which had >10 kg N ha⁻¹ layer⁻¹. There was a flush of nitrate in the S. sesbania and A. angustissima plots with the first rains. Topsoil nitrate had increased to >29 kg N ha⁻¹ by the time of establishing the maize crop. This increase in nitrate in the topsoil was not sustained as concentrations decreased rapidly due to leaching. Nitrate then accumulated below 40 cm, early in the season when maize root length density was still low (<0.1 cm cm⁻³) and inadequate to effectively intercept the nitrate. It is concluded that under light soil and high rainfall conditions, there is an inherent problem in managing nitrate originating from mineralization of organic materials as it accumulates at the beginning of the season, well ahead of peak demand by crops, and is susceptible to leaching before the crop root system develops. This revised version was published online in June 2006 with corrections to the Cover Date.     

Selective logging of lowland evergreen rainforests in Chiloé Island, Chile: Effects of changing tree species composition on soil nitrogen transformations

Lowland evergreen rainforests in southern Chile growing on highly productive soils and accessible sites have been subjected to traditional and industrial logging of valuable timber trees. Old-growth rain forests in this area are characterized by highly conservative N cycles, which results in an efficient N use of ecosystems. We hypothesize that different logging practices, by changing forest structure and species composition, can alter the quantity and quality (i.e. C/N ratio) of litterfall and soil organic matter and soil microbial processes that determine N storage and availability. To test this hypothesis we investigated chemical properties, microbial N transformations, N fluxes and N storage in soils of lowland evergreen rainforests of Chiloé Island after 10 years since industrial selective logging (ISL) and in stands subjected to traditional selective logging (TSL) by landowners in small properties. We compared them to reference unlogged old-growth stands (OG) in the same area. Tree basal area was more reduced in the stands subjected to ISL than to TSL. Litterfall inputs were similar in both logging treatments as in OG stands. This was due to greater biomass of understory species after logging. In TSL understory tree species determined a higher litterfall C/N ratio than ISL. We found higher soil N availability and content of base cations in surface soils of logged forests than in OG. The litter horizon of OG forest had significantly higher rates of non-symbiotic N fixation than logged forests. In the ISL treatment there was a trend toward increasing soil denitrification and significantly higher NO₃–N/N_t ratio in spring waters, which led to a stronger δ¹⁵N signal in surface and deep soils. We conclude that massive understory occupation by the shade-intolerant native bamboo Chusquea quila in ISL led to enhanced litter quality (lower C/N ratios) relaxing the tightness of the N cycle, which increased soil N availability leading to a higher proportion of nitrate in spring waters and higher gaseous N losses. In contrast, under TSL a higher litterfall C/N ratio slowed decomposition and net N mineralization rates thus reducing the chances for N losses, and enhancing C and N storage in soil. We suggest that sustainable logging practices in these rain forests should be based on lower rates of canopy removal to enhance colonization of the understory by shade-tolerant trees, which are associated with a more efficient N cycle.     

Soil nitrogen mineralization at various levels of canopy cover in red pine (Pinus resinosa) plantations

The objective of this study was to determine the rate of nitrogen (N) mineralization in response to various levels of canopy cover in red pine (Pinus resinosa Ait.) stands. Experimental plots consisted of various levels of canopy cover,i.e., clearcut, 25% (50% during first sampling year), 75%, and uncut in red pine plantations in northern Lower Michigan, USA. Net N mineralization and nitrification in the top 15 cm of mineral soil were examined during the first two growing seasons (1991–1992) following the canopy cover manipulations, using anin situ buried bag technique. Mean net N mineralization over the course of both growing seasons (May–October) ranged from 26.9 kg ha⁻¹ per growing season in the clearcut treatment to 13.4 kg ha⁻¹ per growing season in the uncut stand. Net N mineralization and nitrification increased significantly in the clearcut treatment compared to the uncut treatment during the second growing season only. However, net N mineralization and nitrification did not differ significantly between the partial canopy cover treatments and the uncut stand. Increased N mineralization and nitrification in the clearcut during the second growing season may be associated with increased soil temperature and changes of organic matter quality with time since canopy removal. This study was supported in part by the USDA Forest Service and Michigan Technological University.     

Nutrient stocks of short-term fallows on a high base status soil in the humid tropics of Papua New Guinea

In order to understand nutrient dynamics in tropical farming systems with fallows, it is necessary to assess changes in nutrient stocks in plants, litter and soils. Nutrient stocks (soil, above ground biomass, litter) were assessed of one-year old fallows with Piper aduncum, Gliricidia sepium and Imperata cylindrica in the humid lowlands of Papua New Guinea. The experiment was conducted on a high base status soil (Typic Eutropepts), and in Papua New Guinea such soils are intensively used for agriculture. Soil samples were taken prior to fallow establisment and after one year when the fallows were slashed and above ground biomass and nutrients measured. The above ground and litter biomass of piper was 13.7 Mg dry matter ha^-1, compared to 23.3 Mg ha^-1 of gliricidia and 14.9 Mg ha^-1 of imperata. Gliricidia produced almost 7 Mg ha^-1 wood. Total above ground biomass returned to the soil when the fallows were slashed was the same for piper and gliricidia (8 Mg ha^-1). Gliricidia accumulated the largest amounts of all major nutrients except for K, which was highest in the above ground piper biomass. Imperata biomass contained the lowest amount of nutrients. The largest stocks of C, N, Ca and Mg were found in the soil, whereas the majority of P was found in the above ground biomass and litter. Almost half of the total K stock of piper and gliricidia was in the biomass. During the fallow period, soil organic C significantly increased under gliricidia fallow whereas no net changes occurred in piper and imperata fallows. The study has shown large differences in biomass and nutrient stocks between the two woody fallows (piper, gliricidia) and between the woody fallows and the non-woody fallow (imperata). Short-term woody fallows are to be preferred above grass (imperata) fallows in the humid lowlands of Papua New Guinea because of higher nutrient stocks.     

Optimising the management of maize — Grevillea robusta fields in Kenya

The study optimised the management of a Grevillea robusta (A. Cunn.) stand growing in the central highlands of Kenya. The optimisations were conducted separately for even-aged and uneven-aged management system of trees. The management was also optimised with the requirement that maize production under the tree cover must be profitable every year. Technically, the optimisation problems were solved by linking a simulation program with the non-linear optimisation algorithm of Hooke and Jeeves. The simulation program calculated the tree growth, volumes of harvested trees, and maize yields with a given set of management parameters (decision variables). The maize yield predictions and simulated timber yields were converted into gross incomes of which the production costs were subtracted. In even-aged management the objective variable was the soil expectation value with 5% discounting rate. In uneven-aged management the mean annual net income was maximised. The optimal solutions indicated that with both management systems it is optimal to concentrate on wood production. The optimal stand densities were so high that profitable maize production was not possible under the tree cover. The mean annual wood production of the optimal management schedule was more than 50m³ ha^–1. Forcing profitable maize production in the solution decreased the wood production by 57% (even-aged forestry) or 27% (uneven-aged forestry) and net income by 45% (even-aged forestry) or 24% (uneven-aged forestry).This revised version was published online in November 2005 with corrections to the Cover Date.     

Nitrogen mineralization and maize yields following application of tree prunings to a sandy soil in Zimbabwe

Despite the promotion of prunings as sources of nitrogen for crops, lack of synchronization between N mineralization from prunings and plant uptake remains a major limitation to the impact of prunings on crop yields. A laboratory and a field experiment were therefore carried out to determine the mineralization patterns of selected prunings and assess the potential that exists to improve synchrony by mixing prunings of different quality. The laboratory incubation experiment was conducted for 84 days to determine the C and N release patterns of prunings of different quality and the manipulation of the C and N mineralization trends by mixing prunings of different quality. High quality prunings were considered to be those with high CO₂ evolution rates and mineralize N rapidly. The % C and % N released in 84 days were highest for the high quality prunings of Tithonia diversifolia (70% and 30% respectively) and least for the low quality prunings of Flemingia macrophylla (25% and –5% respectively). The medium quality prunings of Acacia angustissima and Calliandra calothyrsus had similar proportions of released C and N (about 40% C and 10% N). Different mixtures of T. diversifolia with other species showed contrasting influence on C release and N mineralization. Most mixtures released less C than that predicted, but in contrast most mixtures released N at a rate either matching or above the predicted. The % N released was strongly correlated with the polyphenol protein binding capacity (r² = 0.53) and also with the % C released (r² = 0.62). The field experiment was done for two cropping seasons to determine the effects of the prunings of these species and their mixtures on maize grain yield. The prunings were added at 5 t ha^–1 and incorporated into the top 15 cm by hand hoeing in the first season and their residual effects were monitored in the second season. Medium and low quality prunings produced significantly (p < 0.05) higher maize grain yields in the first season ranging between 2.4 t ha^–1 and 3.4 t ha^–1 compared with T. diversifolia which produced 1.7 t ha^–1. This suggested better synchrony in N release and uptake by maize with medium and low quality prunings compared with high quality prunings. The only mixture that indicated improved synchrony was the mixture of T. diversifolia and C. calothyrsus. This study showed that mixing prunings of different quality produce different patterns of N mineralization, some of which were unexpected and had a potential for improving N synchrony.This revised version was published online in November 2005 with corrections to the Cover Date.