共查询到20条相似文献,搜索用时 26 毫秒
1.
Seedlings of seven temperate tree species (Acer pseudoplatanus L., Betula pendula Roth, Fagus sylvatica L., Fraxinus excelsior L., Juglans regia L., Quercus petraea Matt. Liebl. and Quercus robur L.) were grown in a nursery under neutral filters transmitting 45% of incident global irradiance. During the second or third year of growth, leaf photosynthetic capacity (i.e., maximal carboxylation rate, Vcmax, maximal photosynthetic electron transport rate, Jmax, and dark respiration, Rd) was estimated for five leaves from each species at five or six leaf temperatures (10, 18, 25, 32, 36 and 40 degrees C). Values of Vcmax and Jmax were obtained by fitting the equations of the Farquhar model on response curves of net CO2 assimilation (A) to sub-stomatal CO2 mole fraction (ci), at high irradiance. Primary parameters describing the kinetic properties of Rubisco (specificity factor, affinity for CO2 and for O2, and their temperature responses) were taken from published data obtained with spinach and tobacco, and were used for all species. The temperature responses of Vcmax and Jmax, which were fitted to a thermodynamic model, differed. Mean values of Vcmax and Jmax at a reference temperature of 25 degrees C were 77.3 and 139 micromol m(-2) s(-1), respectively. The activation energy was higher for Vcmax than for Jmax (mean values of 73.1 versus 57.9 kJ mol(-1)) resulting in a decrease in Jmax/Vcmax ratio with increasing temperature. The mean optimal temperature was higher for Vcmax than for Jmax (38.9 versus 35.9 degrees C). In addition, differences in these temperature responses were observed among species. Temperature optima ranged between 35.9 and above 45 degrees C for Vcmax and between 31.7 and 43.3 degrees C for Jmax, but because of data scatter and the limited range of temperatures tested (10 to 40 degrees C), there were few statistically significant differences among species. The optimal temperature for Jmax was highest in Q. robur, Q. petraea and J. regia, and lowest in A. pseudoplatanus and F. excelsior. Measurements of chlorophyll a fluorescence revealed that the critical temperature at which basal fluorescence begins to increase was close to 47 degrees C, with no difference among species. These results should improve the parameterization of photosynthesis models, and be of particular interest when adapted to heterogeneous forests comprising mixtures of species with diverse ecological requirements. 相似文献
2.
Subcanopy tree species are an important component of temperate secondary forests.However,their biomass equations are rarely reported,which forms a vertical gap between canopy tree species and understory shrub species.In this study,we destructively sampled six common subcanopy species(Syringa reticulate var.amurensis(Rupr.) Pringle,Padus racemosa(Lam.) Gilib.,Acer ginnala Maxim.,Malus baccata(Linn.) Borkh.,Rhamnus davurica Pall.,and Maackia amurensis Rupr.et Maxim.) to establish biomass equatio... 相似文献
3.
David Sánchez-Gomez Miguel A. Zavala Daniël B. Van Schalkwijk Itziar R. Urbieta Fernando Valladares 《Annals of Forest Science》2008,65(6):605-605
4.
D. Craven D. DentD. Braden M.S. Ashton G.P. BerlynJ.S. Hall 《Forest Ecology and Management》2011,261(10):1643-1653
Relative to closed-canopy tropical forests, tree seedlings planted in open grown areas are exposed to higher light intensity, air temperatures, vapor pressure deficit, and greater seasonal fluxes of plant available water than mature tropical forests. The species-specific adaptive capacity to respond to variable precipitation and seasonality in open grown conditions, therefore, is likely to affect species performance in large-scale reforestation efforts. In the present study, we compared the photosynthetic characteristics of eight tropical tree species within and between seasons at two study sites with contrasting dry season intensities. All species except Pseudosamanea guachapele reduced leaf physiological function between the wet and dry seasons. The contrasting severity of seasonal drought stress at the study sites constrained growth rates and photosynthetic characteristics differently. Variation of photosynthetic characteristics at the species level was high, particularly in the dry season. Faster growing species at the less seasonal site, Terminalia amazonia, Inga punctata, Colubrina glandulosa, and Acacia mangium, exhibited a greater adaptive capacity than the other species to down-regulate leaf photosynthesis between seasons. As the dry season was more severe at the more seasonal site, most species strongly reduced physiological function regardless of relative growth rates, except two species (Tectona grandis and P. guachapele) with widespread distributions and relatively high drought tolerance. Our results underscore the need to consider seasonal drought tolerance when selecting tree species for specific reforestation sites. 相似文献
5.
We examined the effects of structural and physiological acclimation on the photosynthetic efficiency of Scots pine (Pinus sylvestris L.) shoots. We estimated daily light interception (DLI) and photosynthesis (DPHOT) of a number of sample shoots situated at different positions in the canopy. Photosynthetic efficiency (epsilon) was defined as the ratio of DPHOT to the potential daily light interception (DLI(ref)) defined as the photosynthetically active radiation (PAR) intercepted per unit area of a sphere at the shoot location. To calculate DLI(ref), DLI and DPHOT, the radiation field surrounding a shoot in the canopy was first modeled using simulated directional distributions of incoming PAR on a clear and an overcast day, and estimates of canopy gap fraction in different directions provided by hemispherical photographs. A model of shoot geometry and measured data on shoot structure and photosynthetic parameters were used to simulate the distribution of PAR irradiance on the needle surface area of the shoot. Photosynthetic efficiency (epsilon) was separated into light-interception efficiency (epsilon(I) = DLI/DLI(ref)) and conversion efficiency (epsilon(PHOT) = DPHOT/DLI). This allowed us to quantify separately the effect of structural acclimation on the efficiency of photosynthetic light capture (epsilon(l)), and the effect of physiological acclimation on conversion efficiency (epsilon(PHOT)). The value of epsilon increased from the top to the bottom of the canopy. The increase was largely explained by structural acclimation (higher epsilon(I)) of the shade shoots. The value of epsilon(PHOT) of shade foliage was similar to that of sun foliage. Given these efficiencies, the clear-day value of DPHOT for a sun shoot transferred to shade was only half that of a shade shoot at its original position. The method presented here provides a tool for quantitatively estimating the role of acclimation in total canopy photosynthesis. 相似文献
6.
Development of leaf chloroplast ultrastructure at five levels in a willow (Salix cv. Aquatica gigantea) canopy was followed during one growing season in the field. Changes in chloroplast ultrastructure were compared with the rate of CO(2) uptake of the same leaves. The highest rates of CO(2) uptake were recorded in young leaves exposed to full available sunlight. In these leaves, the area of the grana stacks was less than 20% of the total chloroplast area and the degree of thylakoid stacking was less than 1.5. The chloroplasts of these leaves contained large amounts of starch and small amounts of plastoglobuli. As the canopy grew and the leaves in the lower parts of the canopy became shaded, the structure of the chloroplast thylakoids gradually changed. In leaves at the two lowest levels of the canopy, the degree of stacking at the end of the growing season was close to 2 and correspondingly the rate of CO(2) uptake was low. The areas of grana stacks and plastoglobuli in these chloroplasts increased and were about 30 and 10% of the chloroplast area, respectively, by the end of the growing season. The increase in the degree of thylakoid stacking was caused by increased biosynthesis of grana lamellae, which in general were thinner than the lamellae of young leaves. The length of the stroma lamellae did not change with leaf age. Morphometric measurements showed that the structure of the chloroplasts in leaves 160 cm above ground was dynamic and responsive to environmental conditions so that photosynthetic capacity remained high for 7 weeks despite an increase in leaf shading. 相似文献
7.
A well-recognized phenomenon during forest development is that stand level forest growth rate begins to decrease after canopy closure. Binkley [Binkley, D., 2004. A hypothesis about the interaction of tree dominance and stand production through stand development. Forest Ecology and Management 190, 265–271] proposed an interesting hypothesis relating individual contribution of trees to forest growth patterns, considering changes in dominance levels and resource use efficiency (ReUE, biomass growth per unit of resource used) of dominant vs suppressed trees. He stated that “the decline in standlevel growth near canopy closure is driven by increasing dominance of larger trees, leading to declining efficiency of resource use by smaller trees”. This decrease in ReUE of suppressed trees once canopy closes would lead to a general decline in standlevel ReUE and thus, in stand growth. This author with other colleagues found evidence according to this hypothesis, but much more research is needed to prove its generality in forests of different species and environmental conditions. The goal of this study was to test Binkley's hypothesis using information of growth patterns, water use efficiency (WUE) and growth efficiency (GE) in different stands of Pinus ponderosa in N.W. Patagonia. According to the hypothesis, we found no dominance in open stands, and some degree of dominance in closed canopy stands. However, in contrast to predictions, WUE differed between the smallest and the largest trees of each stand both with and without dominance. No differences in GE were observed between different tree size groups, and considering individual trees, the more suppressed ones were those with the highest GE. Based on our results, we proposed another hypothesis relating individual and stand ReUE: a decrease in ReUE is not determined by the establishment of dominance and subsequent decrease in suppressed trees efficiencies, but by the availability of resources for individual trees. In this regard, we postulate that differences in ReUE between trees of a stand can appear before dominance, and that these differences in ReUE lead to differences in size of the trees and later, in its social position. When trees grow, they have access to more resources, increasing at the same time, their efficiency using the resources. This efficiency is going to decrease when availability of resources decreases (for competition or environmental stress). The stand ReUE will be maintained, decreased or increased based on the balance of available resources for the individual trees, which in turn will depend on their growth rates, competitive ability and environmental site conditions. 相似文献
8.
在东江中上游,对6年生木荷、红锥、火力楠、藜蒴、灰木莲和枫香的净光合速率(Pn)、蒸腾速率(Tr)、水分利用效率(WUE)等生理生态指标开展观测研究,结果表明:8月,6种树种均表现较强的生活力,保持了较高的净光合速率与蒸腾速率;10月,6树种均通过降低光合、蒸腾和提高水分利用效率来适应逆境。8月平均Pn和Tr要比10月均值分别高71.86%和129.22%,10月WUE却要比8月均值高25.78%。6树种Pn、WUE的平均值高低排序,均表现为木荷〉红锥、藜蒴和火力楠〉枫香和灰木莲,说明木荷有较强的环境适应能力和抗旱能力,其次为红锥、火力楠和藜蒴,而灰木莲和枫香较低。 相似文献
9.
《林业研究》2019,(6)
In 2014, a 40-ha undisturbed plot was established in the Beijing Songshan Nature Reserve. The spatial distribution and spatial associations of five dominant tree species of different height classes were examined using spatial point pattern analysis and the effects of these species on local community diversity assemblages were examined using the ISAR method. The dominant species were characterized by an aggregated distribution at spatial scales of 0–50 m. At smaller spatial scales, trees in larger height classes had significant positive and negative effects on the diversity of the lower height classes. However, at larger spatial scales this effect was diminished. At small scales(0–10 m), accumulator species had positive effects on species diversity, maintained an over-representative proportion of diversity in their proximity, and supported the ecological niche theory. At the larger scale(10–50 m),neutral species were dominant and had positive effects on species diversity, though repeller species, which had negative effects on species diversity, also contributed to diversity. Neutral and accumulator species together determined local species diversity, but the relative importance of the two was closely related to spatial scale. A combination of the ecological niche theory and neutral processes together determines species coexistence and biodiversity of an undisturbed pine forest. 相似文献
10.
11.
Relationships between CO(2) assimilation at light saturation (A(max)), nitrogen (N) content and weight per unit area (W(A)) were studied in leaves grown with contrasting irradiances (outer canopy versus inner canopy) and N supply rates in field-grown nectarine trees Prunus persica L. Batsch. cv. Fantasia. Both A(max) and N content per unit leaf area (N(A)) were linearly correlated to W(A), but leaves in the high-N treatment had higher N(A) and A(max) for the same value of W(A) than leaves in the low-N treatment. The curvilinear relationship between photosynthesis and total leaf N was independent of treatments, both when expressed per unit leaf area A(maxA) and N(A)) and per unit leaf weight (A(maxW) and N(W)), but the relationship was stronger when data were expressed on a leaf area basis. Both A(maxA) and N(A) were higher for outer canopy leaves than for inner canopy leaves and A(maxW) and N(W) were higher for leaves in the high-N treatment than for leaves in the low-N treatment. The relationship between A(max) and N resulted in a similar photosynthetic nitrogen-use efficiency at light saturation (A(max)NUE) for both N and light treatments. Photosynthetic nitrogen-use efficiency was similar among treatments throughout the whole light response curve of photosynthesis. Leaves developed in shade conditions did not show higher N-use efficiency at low irradiance. At any intercellular CO(2) partial pressure (C(i)), photosynthetic CO(2) response curves were higher for outer canopy leaves and, within each light treatment, were higher for the high-N treatments than for the low-N treatments. Consequently, most of the differences among treatments disappeared when photosynthesis was expressed per unit N. However, slightly higher assimilation rates per unit N were found for outer canopy leaves compared with inner canopy leaves, in both N treatments. Because higher daily irradiance within the canopies of the low-N trees more than compensated for the lower photosynthetic performances of these leaves compared to the leaves of high-N trees, daily carbon gain (and N-use efficiency on a daily assimilation basis) per leaf was higher for the low-N treatment than for the high-N treatment in both outer and inner canopy leaves. 相似文献
12.
Naba R. Devkota Peter D. Kemp John Hodgson Ian Valentine I. Komang D. Jaya 《Agroforestry Systems》2009,76(2):363-374
Silvopastoral systems in New Zealand that incorporate trees planted to control soil erosion on hills largely rely on the productivity
of the pastoral system for financial returns. The effect on pasture productivity of increasing the tree canopy height by pruning
Italian gray alder (Alnus
cordata) was investigated by measuring the response of light, soil moisture, soil temperature, pasture production of major pasture
species, and grazing behaviour of sheep. A split-plot design with four replicates was used. The main plot treatments were
three levels of shade (81, 23, and 12% of available photosynthetic photon flux (PPF)), created by pruning 11 year old alder
grown at the same density. The sub-plot treatments were four pasture mixes: perennial ryegrass (Lolium perenne), Yorkshire fog (Holcus lanatus), and cocksfoot (Dactylis
glomerata), each sown with white clover (Trifolium repens), and cocksfoot sown with lotus (Lotus
pedunculatus). Soil temperature was highest under light shade. Total herbage yield at 50 mm stubble height from October to May under heavy
and medium shade was 60 and 80%, respectively, of the total herbage harvested under light shade. Cocksfoot had the greatest
herbage yield, either with lotus or white clover. The tillering of perennial ryegrass was suppressed by shade more than for
the other grass species making ryegrass unsuitable for use in this silvopastoral system. More sheep grazed in the light shade
than in the heavy shade, but there was no difference in sheep preference for cocksfoot or Yorkshire fog. Lotus was grazed
more frequently than white clover. Pruning of alder to increase canopy height has the potential to improve the productivity
of the understorey pasture and its acceptability to sheep. 相似文献
13.
《Scandinavian Journal of Forest Research》2012,27(1-4):75-84
Seedlings of Pinus sylvestris were cold‐stored for three or six months at ‐4°C or 2°C or overwintered outdoors. Dry weight development and net photosynthetic capacity were then measured during a 60‐day cultivation period in a controlled environment. In all storage regimes the longer storage period gave a faster growth initiation. Photosynthetic recovery was faster for seedlings stored at 2°C than at ‐4°C, due to better recovery of stomatal conductance. The results suggest that there is little difference in seedling development between storage temperatures of ‐4°C and 2°C whereas outdoor storage might cause certain negative effects on subsequent growth of seedlings. This result is discussed with regard to the present winter climate. 相似文献
14.
通过调查不同树种新造林树高生长量、保存率及其土壤细菌、真菌、放线菌数量等,研究其生长、适应性及对土壤微生物数量的影响。结果表明:光皮桦、桤木、楸树、栾树、湿地松5年生树高分别为7.4、6.5、6.3、4.8、4.2m,保存率分别为92%、65%、82%、78%、90%;不同树种造林后对土壤微生物数量的提高有明显差异,5个树种造林后林分土壤微生物数量提高的效果排序为桤木〉光皮桦〉栾木〉楸树〉湿地松;新造林均表现为造林当年土壤微生物数量明显下降,一般下降35%~40%,第2年底回升并超过造林前水平,第3-5年逐年增加。 相似文献
15.
为了解不同施肥处理对普陀樟(Cinnamomum japonicum var.Chenii)、全缘冬青(Ilex integra Thunb.)、红楠(Machilus thunbergii Sieb.et Zucc)3种海岛树种苗木生长及造林效果的影响,设置了7个不同的施肥处理,即(1)N(对照),(2)N+P,(3)N+2P、(4)N+K、(5)N+2K、(6)N+1P+1K、(7)N+2P+2K,于2011年5月至2012年1月观测苗高和地径,并测定植物各部分的生物量。结果表明:3种苗木不同施肥处理下,通过测算苗木质量指数QI可知,在N+1K施肥组合下,全缘冬青的苗木质量最好,而红楠在N+2P+2K的施肥组合下苗木质量最高,普陀樟在N+1P+1K的施肥组合下,苗木质量要好于其他处理。 相似文献
16.
Niinemets U 《Tree physiology》2012,32(5):505-509
Plant canopies are characterized by huge within-canopy gradients in environmental and physiological characteristics, but distributions of foliage photosynthetic capacity in plant canopies do not follow simple optimality criteria. Various hypotheses have been put forward to explain the lack of "optimality", including missing physiological and structural constraints and/or missing fundamental relationships. In this issue, Dewar et al. (2012) highlight important optimality constraints due to finite minimum leaf dry mass per unit area the plants can achieve, while Peltoniemi et al. (2012) proposes that limited acclimation of hydraulic conductance to within-canopy light conditions is a key missing factor in canopy optimality. This commentary advocates that future large-scale model exercises need to introduce more realism in model predictions both by including hydraulic controls as well as consider additional constraints shaping foliage structure, chemistry and physiological activity. 相似文献
17.
The survival and growth of Quercus germana Schltdl. & Cham., Q. xalapensis Humb. & Bonpl. and Magnolia dealbata Zucc. was evaluated in three treatments: (1) under the canopy of Pinus maximinoi H.E. Moore; (2) under the canopy of Liquidambar macrophylla Oersted, and (3) in open areas. The following hypothesis was presented: P. maximinoi and L. macrophylla canopy facilitate the establishment of Quercus germana, Q. xalapensis, and Magnolia dealbata. After one year, the survival of seedlings and the growth rate in height and basal diameter were significantly different between treatments and species (p < 0.05). The results suggest a facilitation establishment of Magnolia dealbata under the canopy of Pinus and Liquidambar, whereas the tolerance was observed for Quercus germana and Q. xalapensis. The results will allow to identify restoration strategies of the mountain cloud forest in Mexico.Resumen. Se evaluó la supervivencia y crecimiento de Quercus germana, Q. xalapensis y Magnolia dealbata, en tres tratamientos; (1) bajo el dosel de Pinus maximinoi, (2) bajo el dosel de Liquidambar macrophylla y (3) en áreas abiertas. La hipótesis planteada fue; la cobertura de P. maximinoi y L. macrophylla facilitan el establecimiento de Quercus germana, Q. xalapensis y Magnolia dealbata. Después de un año, la sobrevivencia de plántulas, y las tasas de crecimiento en altura y diámetro de la base, fueron significativamente diferentes entre tratamientos y especies (p < 0.05). Los resultados sugieren que el establecimiento de Magnolia dealbata es facilitado bajo el dosel de Pinus y Liquidambar, mientras que para Quercus germana y Q. xalapensis se observó tolerancia. Los resultados permitirán identificar estrategias de rehabilitación del bosque mesófilo de montaña en México. 相似文献
18.
?Context
Tree species differ largely in their capability to produce characteristic shade leaves with effective morphological and physiological acclimation to low light.?Aims
By examining the sun/shade leaf differentiation in leaf morphology, foliar nitrogen and photosynthetic capacity in five temperate tree species of different successional status, we aimed at identifying those leaf traits that determine the development of a typical shade crown with low light-acclimated leaves.?Methods
Leaf morphology, foliar N content, photosynthetic capacity (V cmax, J max and A max) and leaf dark respiration (R d) were measured in the canopies of 26 adult trees of Fraxinus, Acer, Carpinus, Tilia and Fagus species.?Results
Six traits (the sun/shade leaf differentiation in specific leaf area, leaf size, A max per leaf area or per mass, photosynthetic N use efficiency and R d) were found to characterise best the degree of low light acclimation in shade leaves. All five species exhibited certain modifications in leaf morphology and/or physiology in response to low light; Fagus sylvatica showed the highest and Fraxinus excelsior the lowest shade leaf acclimation.?Conclusions
Our results indicate that the five early/mid- to late-successional species have developed species-specific low light acclimation strategies in their shade crowns which differ in terms of the relative importance of leaf morphological and physiological acclimation. 相似文献19.
To determine the effects of shade on biomass, carbon allocation patterns and photosynthetic response, seedlings of loblolly pine (Pinus taeda L.), white pine (Pinus strobus L.), red maple (Acer rubrum L.), and yellow-poplar (Liriodendron tulipifera L.) were grown without shade or in shade treatments providing a 79 or 89% reduction of full sunlight for two growing seasons. The shade treatments resulted in less total biomass for all species, with loblolly pine showing the greatest shade-induced growth reduction. Yellow-poplar was the only species to show increased stem height growth in the 89% shade treatment. The shade treatments increased specific leaf area of all species. Quantum efficiency, dark respiration and light compensation point were generally not affected by the shade treatments. Quantum efficiency, dark respiration, maximum photosynthesis and light compensation point did not change consistently between the first and second growing seasons. We conclude that differences in shade tolerance among these species are not the result of changes in the photosynthetic mechanism in response to shade. 相似文献
20.
以辽宁省朝阳地区8个枣树品种为试验对象,对净光合速率、蒸腾速率、水分利用效率及其影响因子、气体交换特性进行研究,结果表明:不同枣树品种叶片净光合速率、蒸腾速率和水分利用效率日变化均呈"双峰型"曲线。光合速率最大值依次为金丝王枣金丝王蜜枣金丝蜜枣冬枣三星枣大平顶枣山西枣树大铃铛枣;蒸腾速率最大值依次为金丝蜜枣金丝王枣金丝王蜜枣大铃铛枣冬枣=山西枣树三星枣大平顶枣;水分利用效率最大值依次为金丝王枣金丝王蜜枣金丝蜜枣三星枣山西枣树冬枣大平顶枣大铃铛枣。金丝王枣、金丝王蜜枣和金丝蜜枣3个品种水分利用效率较高,更适合朝阳等干旱地区的气候特点。胞间CO2浓度和气孔导度是枣树水分利用效率最直接的影响因子。 相似文献