Study of structure and dynamics of Korean pine forest in the Xiaoxing’an Mountains

The thesis for the Doctorate consist of two parts. The first is to study on structure, dynamics and tree growth of natural Korean pine forest. The second is to research for tree architecture, growth and stand structure of artificial Korean pine (Pinus koraiensis) forests.The thesis compares the stand structure, process of regeneration, and various tree growth patterns between natural forests and artificial forests. Its purpose is to give a basis for forest management and silviculture. The study was carried out in Liangshui Experimental Forest Farm, which is located in the southern Xiaoxing’an Mountains.     

Stone Pine Forests in Siberia

hitt.oductionStOneglnefOrestsinRussiaareco111POsedofthreestonePinespeCesfSiberianstonepine(PinIissibiIjcaDuTour),Koreanshnepine(PinlisKoIxiiensisSieb.EtZucc.),andJapanesestonepine(PinIispumila[Pall.]Ropl).AllthesespeciesoccuPyabout4Omillionhm',SiberianstO…     

Tree recruitment in gaps of various size,clearcuts and undisturbed mixed forest of interior British Columbia,Canada

Tree seedling recruitment was monitored after various types of logging in mixed conifer and deciduous forests of northern British Columbia, Canada. Predicting tree seedling recruitment after disturbance is fundamental to understanding forest dynamics and succession and is vital for forest management purposes. Seedling recruitment success in multi-species northern latitude forests varied as a function of mature tree canopy cover, gap size and position in a gap. Recruitment was abundant within canopy gaps across a wide range of gap sizes (20–5000 m²), but recruit numbers dropped off rapidly under the closed forest canopy and in the open conditions of clearcuts. Inside canopy gaps, recruitment was similar by gap position in small gaps (<300 m²) but, in these northern latitude forests, exhibited a trend of increasing density from the sunny north to shady south end of larger gaps. This was true for all tree species regardless of their shade tolerance ranking. There was no evidence of gap partitioning by any of the tree species during the regeneration phase suggesting that adaptation to the subtleties of gap size during early recruitment are not well developed in these tree species. Favorable locations for emergence and early establishment of germinants were less favorable for growth and survival of established seedlings, i.e. the regeneration niches in these forests were discordant. Tree abundance and species diversity appears to be controlled more by differentiation among growth and survival niches than by the regeneration niches. From the perspective of forest management, abundant natural regeneration of all the dominant tree species of these mixed-species forests can be obtained after partial cutting.     

Structure and regeneration dynamics of dominant tree species along altitudinal gradient in a dry valley slopes of the Bhutan Himalaya

The aim of our study was to characterize the structure and regeneration dynamics of dominant tree species along altitudinal gradient in dry valley slopes of the Bhutan Himalaya. In the typical dry valley slopes of west-central Bhutan, we analyzed forest stratification, tree crown projection, and population structure from lower dry valley bottom (1520 m a.s.l.) to upper humid ridge top (3370 m a.s.l.) across five major forest types: i.e. (1) dry pine forest with Pinus roxburghii as a dominant species; (2) mixed broad-leaved forest with Quercus lanata (evergreen), Quercus griffithii, (deciduous), and Rhododendron arboreum (evergreen) as major canopy species; (3) evergreen broad-leaved forest dominated by Quercus semecarpifolia, Quercus glauca and Quercus oxyodon; (4) cool conifer forest dominated by Abies densa and Tsuga dumosa; (5) cold conifer forest with A. densa and Juniperus recurva as dominants. In general, regeneration pattern of major dominant species shifted from inverse-J (lower altitudes), to sporadic (mid-altitudes), and to uni-modal type (upper altitudes) corresponding to three regeneration trends: (1) invasive at the lower, warm, dry forest under relatively strong human disturbances; (2) stable/balanced at the mid-altitude, in a relatively stable, mature moist evergreen broad-leaved forest with gap regeneration; and (3) poor/low regeneration at the upper, cool, humid conifer forest with a continuously cattle-grazed understory. Overall, regeneration patterns were balanced in all forest types, however care should be taken to prevent excessive exploitation of dry pine forest of lower valley bottom, and to control cattle grazing at higher altitudes. Our findings can be incorporated into management plans for sustainable management and conservation of mountain forests.     

Variation in post-wildfire regeneration of boreal mixedwood forests: underlying factors and implications for natural disturbance-based management

To test the direct regeneration hypothesis and support natural disturbance-based forest management we characterized the structure and composition of boreal mixedwood forests regenerating after large wildfires and examined the influence of pre-fire stand composition and post-fire competing vegetation. In stands which had been deciduous (Populus sp.)-dominated, conifer (white spruce)-dominated, or mixed pre-fire we measured regeneration stocking (presence in 10 m² plots), density and height 10–20 years post-burn in five wildfires in Alberta, Canada. Most plots regenerated to the deciduous or mixed stocking types; plots in the older fire and in stands that were pure conifer pre-fire had higher amounts of conifer regeneration. Surprisingly, regeneration in pre-fire ‘pure’ white spruce stands was most often to pine, although these had not been recorded in the pre-fire inventory. Pre-fire deciduous stands were the most resilient in that poplar species dominated their post-fire regeneration in terms of stocking, density and height. These stands also had the highest diversity of regenerating tree species and the most unstocked plots. High grass cover negatively affected regeneration density of both deciduous and conifer trees. Our results demonstrate the natural occurrence of regeneration gaps, pre- to post-fire changes in forest composition, and high variation in post-fire regeneration composition. These should be taken into consideration when developing goals for post-harvest regeneration mimicking natural disturbance.     

Proactive intervention to sustain high-elevation pine ecosystems threatened by white pine blister rust

Only recently have efforts begun to address how management might prepare currently healthy forests to affect the outcome of invasion by established non-native pests. Cronartium ribicola, the fungus that causes the disease white pine blister rust (WPBR), is among the introductions into North America where containment and eradication have failed; the disease continues to spread. Ecosystem function is impaired by high rust-caused mortality in mature five-needle white pine forests. This paper evaluates five proactive management options to mitigate the development of impacts caused by white pine blister rust in threatened remote high-elevation five-needle pine ecosystems of western North America. They are: reducing pest populations; managing forest composition; improving host vigor; introducing resistant stock with artificial regeneration; and diversifying age class structure to affect the natural selection process for resistance. Proactive intervention to manage and facilitate evolutionary change in the host species may sustain host populations and ecosystem function during pathogen naturalization.     

STUDY OF STRUCTURE AND DYNAMICS OF KOREAN PINE FOREST IN THE XIAOXING′AN MOUNTAINS

THEMAINCoNTENToNTHENATURALFORESTAgeStTucture,SpatialPattern,andDynamicsofKoreanPineForestSpa-halpatternoftrees,basalstemincrementandheightwereexaminedina5.ohm2mixeddeciduous/Pinuskoraiensisfor-est.ConcIusionsareasfollowingtAgestructureAgedeterminationsonthestudyplotshowthatKoreanpineisalonglivedtreespeciesandunevenagedpopuIation.ThegeneraIecoIogicalIongevi-tyisabout3OO-4oOyear-o1d.TheoverstoryKoreanpinetreesincludeawiderangeofage,themaximumageofKoreanpinetreeonthestudyp1ot…     

STUDY ON THE CUTOVER LAND OF THE HARDWOODKOREAN PINE FOREST FOR REPRODUCTION IN THE XIAOXINGAN MOUNTAINS OF NORTHEAST CHINA

lllehardt`ood-Korcanpincforcstsintl1cXiaoxinganMts.havebecncxtCnsivclycx-ploitcdsinccl948forn1orethan4oycars.Uptono`t.thcrcarcIiuIcKorcanpincresourccstobecxploitedinmostbureaus(rCgions),andthercare2/3ofbureaus(regions)havingPoOrKoreanpincrcsourccs.Inl95Os,Korcanpincforestsoccupicd53.8%ofthcforestcdarcaand74.3%ofthevolumcinthewholercgion.NaturaIKoreanpineforesthasgrcatlydc-creasedto5.4%and9.8%initsareaandvol-umeresPectively,andthcarcaandvoIuncofhardx`oodforcstincrcascdcorrcsPOndingIy.ln…     

Effects of some ecological variables on carabid communities in native and non native forests in the Ibaizabal basin (Basque Country: Spain)

? Carabid beetles were investigated at five different forest types in the Ibaizabal basin (northern Spain). The landscape is characterized by the presence of remnants of native forest surrounded by conifer plantations.

? Carabids were trapped in 52 stands of mixed forest, beech forest, holm oak forest, mixed pine and Monterey pine plantations in 2005 and 2006. The main objectives of the study were: compare carabid diversity, recognise the characteristic species, and study the effects of ecological variables on carabid assemblages in the different forest types.

? No significative differences in species abundance, richness and diversity were found among the studied forests. Most of the trapped beetles were identified as forest generalists, nevertheless some native and non-native forest specialist species were also found. Distribution of carabid communities overlapped and, except for beech forest, no specific communities were distinguished. Altitude, percentage of grass coverage and temperature were the main variables influencing species distribution.

? The results suggest high habitat homogeneity, caused by regeneration in pine plantations of the indigenous understorey, and by poor habitat quality in native forest (patchy remnants enclosed in conifer plantations). This situation could explain the similar carabid diversity.

     

Effects of Forest Management on Field Layer Vegetation: A Comparison Between Finnish and Russian Karelia

The effects of forest management on the field layer vegetation were compared between Finnish and Russian Karelia in a total of 46 mature stands representing pine, spruce and deciduous forests. In Finland, the number of species was slightly higher in each forest type. Many grassland species, such as Galium boreale , Lathyrus pratensis , and Veronica officinalis , were more abundant in Finnish deciduous stands (14.4 species, mean cover 15.8%) than in Russian stands (9.6 species, mean cover 11.5%). Dwarf shrubs, such as Calluna vulgaris (mean cover: Fin 3.3%, Rus 7.7%) and Vaccinium vitis-idaea (Fin 25.0%, Rus 26.9%) in pine stands, and Vaccinium myrtillus (Fin 22.9%, Rus 31.5%) in spruce stands, were more abundant in Russia. Ferns and graminoids exhibited no differences between the two countries. The observed differences were likely to result from the lower tree stand density, more extensive fragmentation and fertilisation of forests in Finland, the higher frequency of forest fires and more intensive grazing in Russian forests, and from differences in natural conditions between Finnish and Russian Karelia.     

Effects of gaps on regeneration of woody plants: a meta-analysis

Forest gaps, openings in the canopy caused by death of one or more trees, have a profound effect on forest regeneration and drive the forest growth cycle. It is therefore necessary to understand the effects of forest gaps on regeneration for modern forest management. In order to provide a quantitative assessment of the effects of forest gaps on regen-eration of woody plants, we conducted this review of gap effects on woody plant regeneration on the basis of 527 observations from 42 indi-vidual papers, and reported the results of these data in a meta-analysis. Overall, densities of regenerated woody plants were significantly greater （359%） in forest gaps than on the closed-canopy forest floor. The regen-eration density in gaps of plantation forests was significantly greater （P＆lt;0.05） than that of natural forest because the regeneration in gaps of plan-tation forests was improved by both gap effects and experimental meas-ures. Similarly, in comparison to natural gaps, regeneration was better enhanced in artificial gaps. Regeneration density exhibited a significantly positive correlation with gap size, but a negative correlation with gap age because the gap size decreased with increasing gap age. Shade tolerance of woody plants affected regeneration density in gaps and understory. Average regeneration density of shade-tolerant species exhibited a sig-nificantly positive response to gaps but densities remained lower in total than those of intermediate and shade-intolerant species. Gap effects on regeneration decreased in response to increasing temperature and pre-cipitation because of the limiting effects of lower temperature and moisture on woody plant regeneration. In summary, forest gaps enhance woody plant regeneration, and the effects of gaps varied by forest type, gap characteristics, environmental factors and plant traits. The results of this meta-analysis are useful for better understanding the effects and roles of gaps on forest regeneration and forest management.     

Restoration of mixed-species forest by gap technique in Sanjiang Plain

IntroductionKoreanpine(PinuskoraiensiSSieb.etZucc.)forestmixedwithbroad-leavedtreesisacIimaxvegetationtypeinNoFtheastChina(Wang1994;Nie1995iZhOu1994).BecausetheforestdominatedbyPinuskoraiensiSthathasveryhighecologicaIandeconomicvalue(Wang1994,Nie1995lZhou1994),itisoneofthemostcompIicatedplantcommunitytypesinNoFtheastAsia.ThisforestisveryimpoFtantforthemaintenanceofbiodiversityandtheconserva-tionofwaterandsoiIintheregionalarea(Liu1957,Zhang1957,Zhang1986,JiLin199O,Wang1994,Nie1995)…     

近自然森林经营在德国的应用成效分析

[目的]分析德国推行近自然森林经营20年来的经营效果,总结德国推行近自然森林经营的经验。[方法]在明晰近自然森林经营原则的基础上,基于两期(间隔期10年)资源清查数据对比,对德国大面积推行近自然森林经营20年的经营效果进行分析。[结果]两期资源清查数据表明,德国采用近自然经营20年后,针叶林如挪威云杉和欧洲赤松的面积显著减少,阔叶林和针阔混交林面积持续增加;阔叶树种蓄积平均增长量约为15 m~3·hm~(-2),针叶树种蓄积平均增长4 m~3·hm~(-2),挪威云杉是唯一蓄积下降的树种,下降了5 m~3·hm~(-2);近自然度等级变化表明,人工林面积在减少,而近自然森林的面积在增加,甚至符合原始林等级的森林面积也在增加。[结论]实现近自然森林经营的目标是一个长期的过程,联邦政府促进和保护森林的政策以及林场主所采用的先进的森林经营技术也是德国森林每公顷蓄积达到336 m~3的重要原因;在近自然森林经营的原则下,德国森林中针阔混交林比例显著增加;将近自然程度不高的人工林经营转化为近自然的森林生态系统是德国森林经营所面临的主要问题。     

Effects of using certain tree species in forest regeneration on volume growth,timber yield,and carbon stock of boreal forests in Finland under different CMIP5 projections

We studied how the use of certain tree species in forest regeneration affected the volume growth, timber yield, and carbon stock of boreal forests in Finland under the current climate (1981–2010) and recent-generation global climate model (GCM) predictions (i.e., multi-model means and individual GCMs of CMIP5), using the representative concentration pathways RCP4.5 and RCP8.5 over the period 2010–2099. Forest ecosystem model simulations were conducted on upland national forest inventory plots throughout Finland. In a baseline management regime, forest regeneration was performed by planting the same tree species that was dominant before the final cut. In alternative management regimes, either Scots pine, Norway spruce, or silver birch were planted on medium-fertility sites. Other management actions over rotation were done as in a baseline management. Compared to baseline management, an increased planting of birch resulted in relative sense highest increase in the volume growth, timber yield, and carbon stock in forests in the south, especially under severe climate projections (e.g., multi-model mean RCP8.5, and GCMs such as HadGEM2-ES RCP8.5 and GFDL-CM3 RCP8.5). This situation was opposite for Norway spruce. In the north, the volume growth, timber yield, and carbon stock of forests increased the most under severe climate projections (e.g., multi-model mean RCP8.5 and CNRM-CM5 RCP8.5), regardless of tree species preference. The magnitude of the climate change impacts depended largely on the geographical region and the severity of the climate projection. Increasing the cultivation of birch and Scots pine, as opposed to Norway spruce, could be recommended for the south. In the north, all three species could be cultivated, regardless of the severity of climate change.     

Influence of different forest conversion strategies on ground vegetation and tree regeneration in pine (<Emphasis Type="Italic">Pinus sylvestris</Emphasis> L.) stands: a case study in NE Germany

The conversion of anthropogenic into more natural, self-regenerating forests is one of the major objectives of forestry throughout Europe. In this study, we present investigations on permanent plots with different silvicultural treatment in NE German pine stands. Management of old-growth pine stands on acidic and nutrient-poor sandy sites differs in fencing, thinning, and planting of certain tree species. The investigations were carried out on the community, population, and individual level of the pine forest ecosystems. Thus, vegetation changes, size and height of tree populations, and height increment of tree individuals were observed over a time span of 6 years. Special attention was paid to short-lived tree species such as, e.g., Frangula alnus and Sorbus aucuparia, as well as to Fagus sylvatica as one of the most typical forest tree species of Central Europe. Vegetation changes are interpreted as a consequence of natural regeneration of formerly degraded forest sites, involving an increase in nutrient availability. High browsing pressure can be considered as a key factor for the inhibition of tree seedlings and growth of saplings. Some Sorbus aucuparia individuals, however, succeeded in growing out of the browsing height also in unfenced stands. Few found specimens of Fagus sylvatica proved that this species is able to establish spontaneously on these relatively dry, acidic sites under continental climate influence. Such natural regeneration processes, also including spontaneous rejuvenation of trees, can be integrated into silviculture as passive forest conversion management. An active management like thinning of stands, planting of trees, and fencing can accelerate forest conversion with regard to height growth and species number of trees.     

Regeneration and structure of mixed conifer forests under single-tree harvest management in the western Bhutan Himalayas

We examined the regeneration and structure of mixed conifer forests under single-tree harvest management in western Bhutan. Sixteen 900 m² (30 m × 30 m) plots were sampled at four Forest Management Units (FMUs; Chamgang, Gidakom, Paro-Zonglela, and Haa-East) representing the forest type, including half the plots in single-tree harvest stands and half in unlogged stands. In addition, we solicited information on traditional forest management practices from informants using survey questionnaires and collected tree species data from felling records from respective local forest offices. Rural timber demand is concentrated on the removal of straight and well-formed bluepine trees for beams, planks, and scaffolding. Single-tree harvest, however, has not significantly altered stand structures from unlogged stands. Similarly, tree regeneration is not different when comparing single-tree harvest and unlogged stands, except at Chamgang FMU, where seedling densities were generally higher in harvested stands than in unlogged stands. These results indicate that single-tree harvest is not detrimental to regeneration and utilization of mixed conifer forests in western Bhutan.     

Comparison of tree growth and undergrowth development in aerially seeded and planted Pinus tabulaeformis forests

Direct seeding is a less expensive practice than planting and has the potential to become a viable alternative to transplanting for afforestation and regeneration purposes. As an effective and a less costly regeneration method, aerial seeding has been applied with several tree species. As early as 1956, Chinese people engaged in aerial seeding and stands with a total of 2.97×10⁷ hm² have been developed up to 2004. Our study tested whether the growth of planted Chinese pine (Pinus tabulaeformis Carr.) seedlings and its undergrowth development in northwest aspects differ from that of aerially sown seedlings on the northern and northwestern aspects of slopes. In 2007, we collected data such as height, diameter at breast height (DBH), clear bole height and canopy widths of trees, abundance, coverage, and frequency of shrubs and herbs from 21-year-old planted Chinese pine stands on a northwestern aspect (PNW), aerially sown stands in a northwest aspect (ANW) and aerially sown stands in a northern aspect (AN). Results showed that the relation of crown area and mean DBH was best fitted by a double inverse model for the ANW and AN forests and by a quadratic model for the PNW forest. There was no difference in the growth between ANW and AN forests, while growth was significantly higher in the PNW forest than in the ANW and AN forests. That was consistent with the Sorenson diversity indices in the shrub and herb layers, indicating that there was a large number of the same species in both aerially seeded stands, although their locations were different. Both the number of species in the undergrowth and the Shannon-Wiener index in the shrub layer were higher in the PNW stands than in the ANW and AN stands. Dominant families for all three stands were Rosaceae and Compositae in the shrub and herb layer, respectively. The dominant species for all three stands was Spiraea pubescens in the shrub layer, while the dominant species was different from each other in the three stands. The discrepancy in diversity and composition of species in the herb layer show that herbs are sensitive to shrubs in the three forests. High mortality and skewed diameter distributions reflect severe competition and too high a density in the aerially seeded forests. Thus, aerial seeding is a viable and effective regeneration technique, but management practices, such as thinning, should be applied to these forests.     

An alternative management regime of selection cutting for sustaining stand structure of mixed forests of northern Japan: a simulation study

In uneven-aged conifer–broadleaved mixed forests in Hokkaido, northern Japan, single-tree selection cutting has been a common management practice since the early twentieth century. This practice is expected to produce timber without major changes in stand structure or tree species composition. The demographic response of forests to this practice has often been unexpected, and degradation of stand properties has been widely observed. We propose here a sustainable management regime of selection cutting, based on an individual-based forest dynamics simulation model, SORTIE-ND. Our simulations, based on demographic data from 15 long-term monitoring stands, suggest that selection cutting using a lower cutting intensity together with a longer rotation period and reduced removal of small trees and conifer species is more appropriate than traditional systems in terms of maintaining stand structure and tree species composition, as well as being profitable financially. Supplemental regeneration practices, which can counter accidental mortality incurred during harvesting operations, would also be necessary to ensure tree recruitment.     

Comparison of stand structure and growth between artificial and natural forests of Pinus sylvestiris var, mongolica on sandy land

Mongolian pine (Pinus sylvestiris Linnaeus var. mongolica Litvinov) as a valuable conifer tree species has been broadly introduced to the sandy land areas in “Three North” regions (North, northwest and northeast of China), but many prob-lems occurred in the earliest Mongolian pine plantations in 7hanggutai, 7hangwu County, Liaoning Province (ZZL). In order to clarify the reason, comprehensive investigations were carried out on differences in structure characteristics, growth processes and ecological factors between artificial stands (the first plantation established in ZZL in 1950s) and natural stands (the origin forests of the tree species in Honghuaerji, Inner Mongolia) on sandy land. The results showed that variation of diameter-class distributions in artificial stands and natural stands could be described by Weibull and Normal distribution models, respectively.Chapman-Richards growth model was employed to reconstruct the growth process of Mongolian pine based on the data from field investigation and stem analysis. The ages of maximum of relative growth rate and average growth rate of DBH, height, and volume of planted trees were 11,22 years, 8, 15 years and 35, 59 years earlier than those of natural stand trees, respectively. In respect of the incremental acceleration of volume, the artificial and natural stands reached their maximum values at 14 years and 33 years respectively. The quantitative maturity ages of artificial stands and natural stands were 43 years and 102 years respectively. It was concluded that the life span of the Mongolian pine trees in natural stands was about 60 years longer than those in artificial stands. The differences mentioned above between artificial and natural Mongolian pine forests on sandy land were partially attributed to the drastic variations of ecological conditions such as latitude, temperature, precipitation, evaporation and height above sea level. Human beings‘‘ disturbances and higher density in plantation forest may be ascribed as additional reasons. Those results may be potentially useful for the management and afforestation of Mongolian pine plantations on sandy land in arid and semi-arid areas.