The effect of scalping on seedling establishment after seed tree cutting of Scots pine stands in drained peatlands in northern Finland

We studied the establishment of natural seedlings after seed tree cutting and scalping in two drained Scots pine peatland stands in northern Finland (Simo and Sievi). Approximately, 50?ha^?1 of Scots pine stems were retained on both sites. During the six subsequent years, five seedling surveys were conducted. The effect of site preparation was analysed using generalized linear mixed models (GLMM). The mean density of pine seedlings increased more quickly in the scalped plots and was ca. 1.1?m^?2 after six years compared to 0.87?m^?2 in the non-scalped plots, on average. Scalping increased the number of pine seedlings 2-fold compared to that in non-treated plots in Simo, however, in Sievi non-treated plots had 30% higher pine seedling density after six years. Downy birch seedling density was 9–10-fold compared to that of non-treated plots in both sites. In terms of seedling density, seed tree cutting led to satisfactory pine regeneration after just two growing seasons in drained pine peatlands, even without scalping. Scalping significantly increased the number of pine seedlings, but it also enhanced the germination of birch seedlings strongly.     

Ground vegetation has a major role in element dynamics in an ash-fertilized cut-away peatland

In Finland, recycling of wood-ash as a fertilizer has been widely studied in recent decades in peatlands drained for forestry. Wood-ash is reported to promote tree growth without any significant negative impacts on the environment and could, therefore, be a suitable option on cut-away peatlands, which are estimated to increase by 30,000 ha over the next ten years. However, the environmental effects of ash-fertilization on cut-away peatlands or on ground vegetation are not well known. We studied the effects of wood- and peat-ash application on the nutrient content in soil and plants and on the heavy metal content in the vegetation of a cut-away peatland. Six treatments of different quantities and mixtures of wood-ash, peat-ash, biotite, or Forest PK-fertilizer were replicated in three plots. Both wood- and peat-ash fertilization ensured an adequate level of nutrients for the early establishment and growth of ground vegetation and birch seedlings in a cut-away peatland. Dense pioneer moss cover was important in trapping the heavy metals, such as cadmium, which are present in the ash. Ground vegetation of mosses and herbaceous plants also had a major role in the retention of the nutrients that could otherwise have leached from the open cut-away peatland area during the early stages of afforestation. Both wood- and peat-ash proved to be suitable for the initial fertilization of afforested cut-away peatlands, but in order to guarantee a sufficient long-term potassium supply for tree growth, an application of biotite with the peat-ash is recommended.     

Comparison of ground disturbance of frozen peatland during stump harvesting using a stump drill and rake

ABSTRACT

Tree stumps could be a source of renewable energy, contributing to a reduced dependence on fossil fuels. In Finland, stumps are currently harvested when the ground is not frozen to avoid co-removal of large amounts of soil and stones. Hence, the machinery used for stump extraction is not operated year-round. On peatlands, stumps could potentially be harvested when the ground is frozen. However, peatlands are highly sensitive to ground disturbance. There is, therefore, a need to identify equipment that causes low ground disturbance. In this study, peatland ground disturbance at stump level caused by stump harvesting using either a stump drill or a conventional stump rake was evaluated and compared in winter conditions. Results show that the stump drill caused up to 90% less ground disturbance per harvested stump than the conventional stump rake, but harvested 32–53% of the stump wood. Additionally, the size and shape of the disturbed areas changed between the harvesting year and following year, indicating that frost heaving plays a role in filling holes caused by stump extraction. The stump drill also consumed similar time to the conventional stump rake when harvesting Scots pine stumps on mineral soils, but far more when harvesting Norway spruce stumps.     

Phosphorus adsorption by peat from drained mires in southern Finland

The phosphorus adsorption capacity of 20 peat soils and those soil properties likely to influence it (Al, Fe, Ca, Mg, pH, ash content) were studied. Phosphorus adsorption was correlated only weakly to the site type classification of peatlands. However, the P adsorption was strongly correlated with the concentration of Fe in the peat. Thus, soil chemical analysis should be used to support site type classification in order to assess the effectiveness of peatlands to adsorb P. When expressed on a volume basis, the P adsorption capacity varied from 7 kg ha^‐1 to 673 kg ha^‐1 in the 0–30 cm layer. Compared with the amounts of P used in practical peatland forest fertilization in Finland (40–45 kg ha^‐1), the P adsorption capacity of several peat soils studied was substantially lower. It was thus concluded that slowly soluble P fertilizers or applications <40–45 kg ha^‐1 should be used where the P adsorption capacity is low.     

Profitability of alternative management regimes in Scots pine stands on drained peatlands

In drained, forested peatlands, ditch network maintenance (DNM) is often considered necessary for tree growth, but it also constitutes additional management costs. Commercial thinnings, in turn, in addition to their silvicultural benefits, are generally applied to enhance the financial performance of stand management but results from peatland stands are scarce. In this study, our aim was to find financially feasible management practices for Scots pine-dominated stands on drained peatland sites in Finland. Using mainly inventory data sets, we compiled altogether 29 typical model-stands for four climatic areas, four site types, and two stand conditions according to need for silvicultural care. We used MOTTI stand simulator to predict the development of the model-stands according to different management regimes consisting of various combinations of 0–2 DNM and 0–2 thinnings with different timings and thinning intensities. We then calculated and compared the financial feasibility of the regimes using net present value (NPV; discount rate 3%) analysis. The separate effect of DNM on the profitability was marginal, but the positive effect of thinnings was clear. The harvesting removals varied within a wide range, depending on the timing and intensity of thinnings, but on average, the NPV doubled due to the thinnings. In the stands of initially good silvicultural condition, regimes including only one thinning and a DNM operation generally displayed a good financial result. In the stands of initially poor silvicultural condition due to neglected early care, regimes with two thinnings produced the best NPV regardless of the often low-yielding first thinning.     

Carbon pools and productivity in a 1-km heterogeneous forest and peatland mosaic in Minnesota,USA

Determining the magnitude of carbon (C) storage in forests and peatlands is an important step towards predicting how regional carbon balance will respond to climate change. However, spatial heterogeneity of dominant forest and peatland cover types can inhibit accurate C storage estimates. We evaluated ecosystem C pools and productivity in the Marcell Experimental Forest (MEF), in northern Minnesota, USA, using a network of plots that were evenly spaced across a heterogeneous 1-km² mosaic composed of a mix of upland forests and peatlands. Using a nested plot design, we estimated the standing C stock of vegetation, coarse detrital wood and soil pools. We also estimated aboveground net primary production (ANPP) as well as coarse root production. Additionally we evaluated how vegetation cover types within the study area differed in C storage. The total ecosystem C pool did not vary significantly among upland areas dominated by aspen (160 ± 13 Mg C ha⁻¹), mixed hardwoods (153 ± 19 Mg C ha⁻¹), and conifers (197 ± 23 Mg C ha⁻¹). Live vegetation accounted for approximately 50% of the total ecosystem C pool in these upland areas, and soil (including forest floor) accounted for another 35–40%, with remaining C stored as detrital wood. Compared to upland areas, total C stored in peatlands was much greater, 1286 ± 125 Mg C ha⁻¹, with 90–99% of that C found in peat soils that ranged from 1 to 5 m in depth. Forested areas ranged from 2.6 to 2.9 Mg C ha⁻¹ in ANPP, which was highest in conifer-dominated upland areas. In alder-dominated and black spruce-dominated peatland areas, ANPP averaged 2.8 Mg C ha⁻¹, and in open peatlands, ANPP averaged 1.5 Mg C ha⁻¹. In treed areas of forest and peatlands, our estimates of coarse root production ranged from 0.1 to 0.2 Mg C ha⁻¹. Despite the lower production in open peatlands, all peatlands have acted as long-term C sinks over hundreds to thousands of years and store significantly more C per unit area than is stored in uplands. Despite occupying only 13% of our study area, peatlands store almost 50% of the C contained within it. Because C storage in peatlands depends largely on climatic drivers, the impact of climate changes on peatlands may have important ramifications for C budgets of the western Great Lakes region.     

欧洲和北美地区退化泥炭地修复技术与借鉴

泥炭地是重要的自然湿地类型,具有固碳、水文调节、生物多样性维护等多种生态系统服务功能。受侵蚀、火灾等自然灾害以及排水疏干、过度放牧和基础建设等强烈的人为干扰,全球泥炭地面积不断缩减,成为全球气候变化背景下的碳源。文中从水文修复、植被修复和生态监测与评估等方面,综述了欧洲和北美国家退化泥炭地修复技术研究进展,结合若尔盖湿地修复技术现状进行了研究展望,旨在为我国高原退化湿地的生态修复提供参考。     

Carbon stock changes of forest land in Finland under different levels of wood use and climate change

Context

Prediction of the effect of harvests and climate change (CC) on the changes in carbon stock of forests is necessary both for CC mitigation and adaptation purposes.

Aims

We assessed the impact of roundwood and fuelwood removals and climate change (CC) on the changes in carbon stock of Finnish forests during 2007–2042. We considered three harvest scenarios: two based on the recent projections of roundwood and fuelwood demand, and the third reflecting the maximum sustainable cutting level. We applied two climate scenarios: the climate was in the state that prevailed around year 2006, or it changed according to the IPCC SRES A1B scenario.

Methods

We combined the large-scale forestry model MELA with the soil carbon model Yasso07 for mineral soils. For soils of drained, forested peatlands, we used a method based on emission factors.

Results

The stock change of trees accounted for approximately 80 % of the total stock change. Trees and mineral soils acted as carbon sinks and the drained peatland soils as a carbon source. The forest carbon sink increased clearly in both of the demand-based scenarios, reaching the level of 13–20 Tg C/year (without CC). The planned increase in the use of bioenergy reduced the forest sink by 2.6 Tg C/year. CC increased the forest carbon sink in 2042 by 38 %–58 % depending on the scenario. CC decreased the sink of mineral soils in the initial years of the simulations; after 2030, the effect was slightly positive. CC increased the emissions from the drained peatland soils.

Conclusions

It is likely that forest land in Finland acts as a carbon sink in the future. The changes in carbon stocks of trees, mineral soils, and peatland soils respond differently to CC and fuelwood and roundwood harvests.     

Growth and herbage productivity of Gliricidia in Panicum maximum pasture as influenced by seedbed preparation,planting and weed control techniques

Interplanting Gliricidia (Gliricidia sepium [Jacq.] Walp.) into poor quality native grass can improve the quality of pasture. But information on methods to establish the tree legume into existing pasture is scarce. This study was designed to compare the effects of disc-ploughing, hoeing, or rotary tillage operations, and direct seeding or planting out potted seedlings on growth and dry matter yields of Gliricidia in grass pasture. The Guinea grass (Panicum maximum) pasture was first mowed to a height of 10 cm. Then the seedbed was prepared by either hoeing 0.5 m strip or rototilling 0.5 m strip with a 2-wheel tractor or disc-ploughing 1.0 m strip with a 4-wheel tractor. Eight weeks old potted seedlings were planted or seeds were sown direct. At 6 months after planting (MAP), trees in disc-ploughed strips averaged 81 cm in height, about 35% taller than trees in hoed- or rotary-tilled strips. Potted seedlings (average height 77 cm) grew 37% taller than direct-seeded trees. At 12 MAP trees in disc-ploughed strips yielded 95% more leaf DM (1170 vs. 600 kg/ha) than trees in hoed- or rotary-tilled strips. In order to intercrop Gliricidia into existing Guinea grass pasture, it is necessary to plant potted seedlings previously raised in nursery into 1.0 m wide strips disc-ploughed with 4-wheel tractor. Weed control is necessary.     

Ash-fertilization improves germination and early establishment of birch (Betula pubescens Ehrh.) seedlings on a cut-away peatland

Common objectives of afforestation of cut-away peatlands are the diminution of nutrition loads and the production of energy wood in an efficient and economic way. We studied the effect of fertilization and afforestation methods on the early establishment of downy birch (Betula pubescens Ehrh.) seedlings on a cut-away peatland. Six treatments of peat-ash, wood-ash, biotite, or PK-fertilizer were replicated in three blocks in different mixtures corresponding to 50 kg/ha of phosphorous. Birch seeds were sown in randomized halves of split plots, while other halves were naturally afforested. Natural afforestation was equal to sowing in terms of seedling amounts due to closeness of surrounding birch stands with abundant seed crops. After the first growing season the greatest quantities and dominant heights of birch seedlings appeared on the ash-fertilized plots, whereas on the unfertilized areas both the establishment and growth of seedlings was very poor. On the other hand, the quantity of Scots pine (Pinus sylvestris L.) seedlings was more than three times higher on the unfertilized areas compared to the ash-fertilized areas. The differences between the fertilized and the unfertilized areas accentuated during the four observation years. Our results support regular use of ash-fertilizers for the afforestation of cut-away peatlands instead of the ash being deposited as a waste in landfills.     

竹柏不同播种育苗方式的生长效果分析

采用容器(10 cm×10 cm营养钵)和田间苗床2种播种育苗方式,进行了竹柏不同播种密度、不同遮荫处理的比较试验,结果表明:2种育苗方式对竹柏种子的发芽率没有显著影响,田间苗床播种苗的生长量大于容器苗;田间苗床的播种密度以株行距10 cm×20 cm最为适宜,生长量最大;生长期遮荫对竹柏播种苗有极显著影响,遮荫处理的苗木保存率达92.3%,未遮荫的苗木保存率只有37.5%。     

Framework for designing and applying peak runoff control structures for peatland forestry conditions

Drainage-induced diffuse pollution and erosion are key water quality problems in peatland forestry. A major part of the pollutant load is transported during peak runoff periods after snowmelt or intense rainfall. This study investigated possibilities to increase retention time of runoff waters in drained peatland catchments on purpose to diminish peak runoff and improve settling conditions of suspended solids (SS). To create retention, a peak runoff control (PRC) structure was developed and its functioning, dimensioning and practical applications were studied in five partly or completely ditch-drained catchments in Central Finland. The method reduced runoff peaks by 10–73% or 5.07–57.63 l⁻¹ s⁻¹ km⁻², and functioned especially well during largest runoff peaks. The effectiveness of the PRC method depended on (i) catchment topography (slope) and available detention volume, (ii) dimensioning and location of the PRC structure, and (iii) runoff rates. The PRC structure is cheap and can easily be created with forest drainage machinery during the ditching and ditch network maintenance operations. Different issues relating to the structural design, water quality benefits, and impacts on forestry are discussed.     

Windthrow after Shelterwood cutting in Picea abies Peatland forests

The aim of the study was to address the question of whether shelterwood cutting should be discouraged as a forest regeneration method in Norway spruce (Picea abies (L.) Karst.) forests on highly productive peatlands due to a higher risk of windthrow in such forests. The total extent of windthrow was observed during six years after shelterwood cutting in nine spruce forests on fertile peatlands in north, central and south Sweden. Shelterwoods at densities of 140 and 200 stems ha^‐1 were studied. At the end of the six year study period, 43% of all trees in the sparser shelterwoods and 38% of the trees in the denser shelterwoods had blown down. With respect to results from recent studies of the development of naturally regenerated and planted seedlings under the shelter trees, the extent of windthrow was regarded acceptable. Analysis of wind data from national weather stations close to the experiments showed that the acceptable extent of windthrow could not be explained by low frequencies of high winds during the shelterwood period. On the contrary, according to the wind observations made at the weather stations selected in central and south Sweden, the annual number of days with high winds (>21 ms^‐1) during the six years after the shelter‐wood cutting was larger than the average high wind frequency for the last 40 years. Observations of maximum wind speeds show that the mean annual gale force during the shelterwood period was higher than the long term average. Thus, the results from this study do not justify avoidance of shelterwood cutting in spruce peatland forests because of the risk of windthrow. Each of the shelterwoods in the experiment was located close to a large clearcut, and the shelterwood cuttings were carried out as heavy thinnings. For further reduction of windthrow, denser shelterwoods and no clearcutting in adjacent stands are suggested when using shelterwood systems in practical forestry.     

Rhizoctonia solani AG 2-1 as a causative agent of cotyledon rot on European beech (Fagus sylvatica)

Rhizoctonia solani was frequently isolated in the Italian Alps from nursery-grown European beech (Fagus sylvatica) seedlings displaying symptoms of cotyledon rot. Koch's postulates were verified and mode of infection of the associated isolates was investigated with light and scanning electron microscopy. Population structure of the pathogen was investigated by scoring the anastomosis reaction type in pairings between different isolates from the same seedbed. One pathogen genotype showed a large distribution area within the seedbed, this implying that the inoculum had been building up in the seedbed over a longer time period. Hyphal anastomosis tests and sequence analysis of the internal transcribed spacer (ITS) region of ribosomal DNA indicated that the pathogen belongs to AG 2-1 of R. solani. ITS sequence analysis indicates that the isolates from beech are closely related to R. solani isolates causing a disease on tulip. The striking similarities between the two diseases are discussed.     

Effects of Seedbed Substrate on Moisture Conditions and Germination of Pinus sylvestris Seeds in a Clearcut

The objective of the investigation was to describe moisture conditions and germination of Scots pine (Pinus sylvestris L.) seeds in different seedbed substrates (feather mosses, mineral soil, humus and ground organic material) with and without irrigation during 2 weeks, in a clearcut in northern Sweden. Without irrigation no seeds germinated, which could be explained as an effect of the low moisture conditions of the seedbed substrates. This was shown in the seeds' low moisture content, with highest values in mineral soil, 10-20%, and for the other seedbed substrates 5-15%. The moisture conditions were highest during the night and lowest in the afternoon. With irrigation, moisture conditions improved (seed moisture content 20-50%) and diurnal changes were moderated. Seeds germinated in all substrates with most seedlings in mineral soil (60% of sown germinable seeds). The moisture conditions of the seedbed substrates proved to be in poor agreement with the seeds' requirement for germination, especially during warm weather with no precipitation.     

Effects of forest floor planting and stock type on growth and root emergence of Pinus contorta seedlings in a cold northern cutblock

A two-year field trial was conducted to determine the growth response, and root emergence pattern of interior lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) seedlings in response to container type and forest floor removal. Seedlings were grown in Styroblocks^TM, Copperblocks^TM, or AirBlocks^TM, and were planted directly into the undisturbed forest floor or into manually prepared planting spots where the forest floor had been scraped away to expose the mineral soil. Seedlings planted into scalped planting spots exhibited marginally but significantly (7%) greater above-ground growth rates (seedling stem volume); whereas seedlings planted into the forest floor produced significantly more (11%) new roots. There were no differences in above- or below-ground biomass. Seedlings grown in Copperblock^TM containers produced a higher proportion of roots near the top of the plug when tested at lifting, however this pattern was not observed in the field. Given that scalping is more costly than forest floor planting, and that the increased shoot growth was relatively small, we recommend that forest floor planting be considered as an alternative to manual spot scalping for sites, such as the site tested here: those with cold, but well-drained soils and where competition from other plants is not a serious problem.     

Airborne small-footprint discrete-return LiDAR data in the assessment of boreal mire surface patterns, vegetation, and habitats

Boreal mires encompass high diversity in species and habitats, many of which are endangered. In Finland, extensive use of peatlands has resulted in habitat fragmentation. A need for accurate and cost-efficient vegetation mapping and monitoring of habitat changes exists in mire conservation, restoration and peatland forestry. LiDAR is an emerging and excellent tool for probing the geometry of vegetation and terrain. Modern systems measure the backscattered signal accurately and also provide radiometric information. Experiments were carried out in a complex minerotrophic–ombrotrophic eccentric raised bog in southern Finland (61°47′N, 24.18′E). First, we tested discrete-return LiDAR for the modeling of mire surface patterns and the detection of hummocks and hollows, as well as the effect of mire plants on the Z accuracy of the surface echoes. Secondly, the response of different mire vegetation samples in LiDAR intensity was examined. Thirdly, we tested area-based geometric and radiometric features in supervised classification of mire habitats to discover the meaningful variables. The vertical accuracy of LiDAR in mire surface modeling was high: 0.05–0.10 m. A binary hummock-hollow model that was estimated from a LiDAR-based elevation model matched flawlessly in aerial images and had moderate explanatory power in habitat classification trials. The intensity of LiDAR in open-mire vegetation was mainly influenced by the surface moisture, and separation of vegetation classes spanning from ombrotrophic to mesotrophic vegetation proved to be difficult. Area-based features that characterize the height distribution of LiDAR points in the canopy were the strongest explanatory variables in the classification of 21 diverse mire site types. Actual qualifying differences in the ground flora were unattainable in the LiDAR data, which resulted in inferior accuracy in the characterization of ecohydrological conditions and nutrient level of open mires and sparsely forested wet sites. Mire habitat classification accuracy with LiDAR surpassed earlier results with optical data. The results suggested that LiDAR constitutes an efficient aid for monitoring applications. We propose the co-use of images and LiDAR for enhanced mapping of open mires and tree species. In situ calibration and validation procedures are required until invariant geometric and radiometric features are discovered.     

Scenario analyses on the effects of fertilization,improved regeneration material,and ditch network maintenance on timber production of Finnish forests

We used national scenario analyses to examine the effects of fertilization, use of improved regeneration material and ditch network maintenance (DNM), both separately and simultaneously, on timber production of Finnish forests under the current climate. We also analyzed how the area of artificial regeneration, forest fertilization, and DNM developed in different management and harvesting intensity scenarios. The initial data were obtained from the 11th National Forest Inventory of Finland, excluding protected forests. Four sets of even-flow harvesting scenarios with annual timber harvest targets of 60, 70, 80, and 90 million m³ were developed for 90-year simulation period. Use of improved material in artificial forest regeneration was assumed to result in 10% higher diameter and height increment compared to naturally regenerated seedlings. Sub-xeric pine-dominated and mesic spruce-dominated sites were fertilized, and 40% of drained peatlands were maintenance-ditched when they fulfilled a set of predetermined criteria for temperature sum, stand basal area, and mean tree diameter. As a result, when fertilization, improved regeneration material, and DNM were all used, the mean annual volume increment over the 90-year simulation period increased by 3.4–5.4 million m³ depending on harvesting intensity. The maximum sustainable harvest of timber would be almost 80 million m³ yr^?1. The simulated fertilization area was about four times larger than the presently fertilized area, and the simulated DNM area was about the same as the current. Fertilization gave the largest additional 90-year volume increment and the DNM the smallest when they were used separately. The use of improved regeneration material gave the largest additional volume increment in southern Finland and fertilization in central and northern Finland.     

Probability models for pine twisting rust (Melampsora pinitorqua) damage in Scots pine (Pinus sylvestris) stands in Finland

Factors affecting the probability that pine twisting rust (Melampsora pinitorqua) damage occur in a Scots pine (Pinus sylvestris) stand were analysed using the 7th Finnish National Forest Inventory data (NFI7) from southern Finland in 1977–1983. The inventory was based on systematic sampling. The NFI7 data was measured in clusters, each of which consisted of 21 sample plots. In addition to the stand and site characteristics measured for forest management planning purposes, the data included records of damage by pine twisting rust and occurrence of aspens (Populus tremula, the other host plant of the pathogen) in the stands. Two multilevel logit models were developed for predicting the overall probability of pine twisting rust damage and the probability of severe pine twisting rust damage. Site and stand characteristics were used as explanatory variables in the models. Residual variance in the models was studied on the inventory crew, cluster and year levels. The occurrence of aspens and site fertility were the most important factors increasing the probability that pine twisting rust damage will occur in a stand. The damage probability also decreased with increasing effective temperature sum calculated for the location. The overall damage probability was equally high on peatlands and on mineral soil if there were aspens in the stand. If, however, there were no aspens in the stand, the probability of damage was higher on mineral soils than on peatlands. In addition, the overall probability was lower in naturally regenerated stands than in planted or sown stands, and it decreased with increasing mean age of pines. In both models, the residual variance was significant on the both the inventory crew and the cluster levels.     

秦岭冷杉幼苗适应性的研究

通过对秦岭冷杉不同海拔、光照、苗床类型、覆土深度等条件下的野外播种试验,分析了不同条件下种子萌发、幼苗存活和幼苗生长率的差异,揭示了秦岭冷杉种子及幼苗在天然环境下的适应性.天然生境下秦岭冷杉的平均出苗率只有17.7％,不及实验室发芽率(36％)的一半.海拔、光照、苗床类型、埋藏深度对秦岭冷杉的种子萌发、幼苗存活和幼苗生长率都有不同程度的影响.生产上秦岭冷杉的复壮一定要遵循适地适树的原则.