Tithonia diversifolia as a green manure for soil fertility improvement in western Kenya: A review

Tithonia diversifolia, a shrub in the family Asteraceae, is widely distributed along farm boundaries in the humid and subhumid tropics of Africa. Green biomass of tithonia has been recognized as an effective source of nutrients for lowland rice (Oryza sativa) in Asia and more recently for maize (Zea mays) and vegetables in eastern and southern Africa. This paper reviews the potential of tithonia green biomass for soil fertility improvement based on recent research in western Kenya. Green leaf biomass of tithonia is high in nutrients, averaging about 3.5% N, 0.37% P and 4.1% K on a dry matter basis. Boundary hedges of sole tithonia can produce about 1 kg biomass (tender stems + leaves) m^–1 yr^–1 on a dry weight basis. Tithonia biomass decomposes rapidly after application to soil, and incorporated biomass can be an effective source of N, P and K for crops. In some cases, maize yields were even higher with incorporation of tithonia biomass than with commercial mineral fertilizer at equivalent rates of N, P and K. In addition to providing nutrients, tithonia incorporated at 5 t dry matter ha^–1 can reduce P sorption and increase soil microbial biomass. Because of high labor requirements for cutting and carrying the biomass to fields, the use of tithonia biomass as a nutrient source is more profitable with high-value crops such as vegetables than with relatively low-valued maize. The transfer of tithonia biomass to fields constitutes the redistribution of nutrients within the landscape rather than a net input of nutrients. External inputs of nutrients would eventually be required to sustain production of tithonia when biomass is continually cut and transferred to agricultural land.This revised version was published online in November 2005 with corrections to the Cover Date.     

Combining Tithonia diversifolia and fertilizers for maize production in a phosphorus deficient soil in Kenya

The ability of Tithonia diversifolia, fertilizers and their combination to improve maize production in a P-deficient Ferralsol was studied in western Kenya. Tithonia and fertilizers were applied separately or combined in different proportions to give equal rates of165 kg N ha⁻¹, 15.5 kg P ha⁻¹ and 155 kg K ha⁻¹ in two consecutive maize growing seasons, followed by two residual maize crops. Maize grain yields and P recovered in the above-ground biomass were higher in the pure Tithonia than the pure fertilizer treatments. Maize yields increased with increasing proportions ofTithonia in the mixed treatments. When less than 36% of theP applied in the mixture was supplied by Tithonia, there was no additional yield benefit in the combined treatments compared to the pure fertilizer treatments. However an added value ranging from 18 to 24% in yield was observed at greater Tithonia proportions. Economic returns were greater when Tithonia was applied alone than when fertilizers were used, with a larger profit whenTithonia was collected from existing niches than when produced on site. Collecting Tithonia from current niches resulted also in larger net returns for all the combined treatments compared to the fertilizer treatments. The results of this study indicate that a high quality organic residue such as Tithonia can increase maize production to a greater extent than fertilizers. In low input systems, the combination of Tithonia and fertilizers is a valuable alternative when resources are scarce and an added benefit can be obtained by maximizing the proportion of Tithonia in the mixture. This revised version was published online in June 2006 with corrections to the Cover Date.     

Coppicing improved fallows are profitable for maize production in striga infested soils of western Kenya

Striga hermonthica (striga) weed is a major threat to crop production in sub-Saharan Africa, and short duration improved fallow species have recently been found to reduce the effects of this weed because of their ability to replenish soil nitrogen. The objective of this study was to compare the efficacy and profitability of coppicing improved fallow species (Gliricidia sepium [gliricidia], Leucaena trichandra [leucaena] and Calliandra calothyrsus [calliandra]) and non-coppicing species (Sesbania sesban [sesbania], Mucuna pruriens [mucuna], and Tephrosia vogelii [tephrosia]), in controlling striga. Natural fallow and a sole maize crop were included as control treatments. The fallow treatments were split into two and either fertilized with N or unfertilized. The results showed that coppicing fallows produced higher biomass than non-coppicing fallows. For example, Callindra (coppicing fallow species) produced 19.5 and 41.4 Mg ha⁻¹ of leafy and woody biomass, respectively after four cumulative harvests as compared with Sesbania (non-coppicing species), which produced only 2.3 and 5.9 Mg ha⁻¹ leaf and woody biomass, respectively. Improved fallows reduced striga population in proportion to the amount of leafy biomass incorporated into the soil (r = 0.87). N application increased cumulative maize yield by between 15–28% in improved fallow systems and by as much as 51–83% in the control treatments. Added total costs of the coppicing fallows did not differ significantly from those of the non-coppicing fallows and control treatments. However, the added net benefits of the coppicing fallows were significantly higher (US$ 527 for +N and 428 for −N subplots; P < 0.01) than those of the non-coppicing fallows (US$ 374 for +N and 278 for −N), and the least for the control treatments. The most profitable fallow system was Tephrosia with net added benefits of US$ 453.5 ha⁻¹ season⁻¹ without N, and US$ 586.7 ha⁻¹ season⁻¹ with added N.     

Improved fallows come of age in the tropics

Improved fallows are the deliberate planting of fast-growing species — usually legumes — for rapid replenishment of soil fertility. Improved fallows are rapidly spreading in several regions of the tropics as a sensible way for in situ accumulation of large quantities of N in vegetation and soil, as well as for providing sustainability enhancing services. Research on improved fallows increased after the mid 1980s with the development of what is known as the second soil fertility paradigm, which is based on sustainability considerations. Many lessons have emerged from short-term improved fallows (<5 years duration). These include the diversity of farm sizes where improved fallows are used, the advantage of sequential versus simultaneous systems, the utilization of dry seasons unfavorable for crop production, the comparative advantages of woody versus herbaceous leguminous fallows, the magnitude of N accumulation, the strategic use of N fertilizers, and the importance of P. Other key services provided by fallows include fuelwood production, recycling of nutrients besides N, provision of a C supply to soil microorganisms, weed suppression, Striga control, and improved soil water storage. Natural fallows of non-legume shrubs belonging to the Asteraceae family, collectively called daisy fallows, may provide lessons for the development of improved fallows. The maintenance of genetic diversity in fallows is paramount. The main limiting factor in Africa is clearly the supply of germplasm of improved fallow species. This must be overcome though large-scale seed orchards and nursery development before impact at the scale of millions of farmers can take place.This revised version was published online in November 2005 with corrections to the Cover Date.     

The root-knot nematode problem in sesbania fallows and scope for managing it in western Kenya

Sesbania (Sesbania sesban), which is promoted as a short-duration-fallow species in eastern and southern Africa, is an alternative host to root-knot nematode (Meloidogyne spp.). Therefore, it is important to know the extent of sesbania infection by the nematode on farms and its effects to crops grown in rotation with sesbania. A survey of 14 farms in western Kenya indicated that sesbania was infected by root-knot nematode in all the farms. The potential to build up the nematode population was much greater on farms where maize (Zea mays) was previously intercropped with a susceptible bean (Phaseolus vulgaris). Screening of 30 single plant accessions of four sesbania provenances (Kakamega, Siaya, Kisumu, and Kisii) from western Kenya in pots indicated limited scope for selecting material resistant to the nematode. All four provenances were infested by the nematode, and they increased its population in soil and root. A rangewide collection of sesbania germplasm needs to be screened to determine the scope of finding nematode-resistant material. Transplanted seedlings produced in heat-sterilized, nematode-free soil showed less nematode infection than direct-seeded plants. However, both direct seeding and transplanting seemed to increase nematode population similarly. A number of Crotalaria spp. that were found resistant to nematodes can be considered as alternatives to sesbania-planted fallows, but further field-scale testing is necessary to select appropriate species based on biomass production. Among other species suitable for short-duration fallows, only pigeonpea (Cajanus cajan) and senna (Senna siamea) showed promise. Species such as neem (Azadirachta indica), casuarina (Casuarina junghuhniana), and grevillea (Grevillea robusta) that are suitable for boundary plantings were free from the root-knot nematode.This revised version was published online in November 2005 with corrections to the Cover Date.     

Potential of improved fallows to increase household and regional fuelwood supply: evidence from western Kenya

Fuelwood is the main energy source for households in rural Africa, but its supply is rapidly declining especially in the densely populated areas. Short duration planted tree fallows, an agroforestry technology widely promoted in sub-Sahara Africa for soil fertility improvement may offer some remedy. Our objective was to determine the fuelwood production potential of 6, 12 and 18 months (the common fallow rotation periods) old Crotalaria grahamiana, Crotalaria paulina, Tephrosia vogelli and Tephrosia candida fallows under farmer-managed conditions in western Kenya. Based on plot-level yields, we estimated the extent to which these tree fallows would meet household and sub-national fuelwood needs if farmers planted at least 0.25 hectares, the proportion of land that is typically left under natural fallows by farmers in the region. Fuelwood yield was affected significantly (P < 0.05) by the interaction between species and fallow duration. Among the 6-month-old fallows, T. candida produced the highest fuelwood (8.9 t ha⁻¹), compared with the rest that produced between 5.6 and 6.2 t ha⁻¹. Twelve months old T. candida and C. paulina also produced significantly higher fuelwood yield (average, 9.6 t ha⁻¹) than T. vogelli and C. grahamiana of the same age. Between the fallow durations, the 18-month fallows produced the most fuelwood among the species evaluated, averaging 14.7 t ha⁻¹. This was 2–3 times higher than the average yields of 6 and 12-month-old fallows whose yields were not significantly different. The actual fuelwood harvested from the plots that were planted to improved fallows (which ranged from 0.01 to 0.08 ha) would last a typical household between 11.8 and 124.8 days depending on the species and fallow duration. This would increase to 268.5 (0.7 years) and 1173.7 days (0.7–3.2 years) if farmers were to increase area planted to 0.25 ha. Farmers typically planted the fallows at high stand densities (over 100,000 plants ha⁻¹ on average) in order to maximize their benefits of improving soil fertility and providing fuelwood at the same time. This potential could be increased if more land (which fortunately exists) was planted to the fallows within the farms in the region. The research and development needs for this to happen at the desired scale are highlighted in the paper.     

Carbon pools in tree biomass and the soil in improved fallows in eastern Zambia

This study quantified tree and soil C stocks and their response to different tree species and clay contents in improved fallows in eastern Zambia. From 2002 to 2003, soil, and destructively harvested two-year old tree, samples were analysed for C. There were significant differences (P < 0.05) in aboveground tree C stocks, and in net organic C (NOC) intake rates across coppicing tree species at Msekera and Kalunga. Aboveground C stocks ranged from 2.9 to 9.8 t ha^-1, equivalent to NOC intakes of 0.8–4.9 t ha^-1 year^-1. SOC stocks in non-coppiced fallows at Kalichero and Msekera significantly differed (P < 0.05) across treatments. SOC stocks to 200 cm depth ranged from 64.7 t C ha^-1 under non-coppicing fallows at Kalunga to 184.0 t ha^-1 in 10-year-old coppicing fallows at Msekera. Therefore, tree and soil C stocks in improved fallows can be increased by planting selected tree species on soils with high clay content.     

Modelling of planted legume fallows in Western Kenya using WaNuLCAS. (I) Model calibration and validation

Poor soil fertility is the biggest obstacle to agricultural productivity in Sub-Saharan Africa. Improved fallows can help to raise agricultural productivity in these systems of low financial capital, however, experimental testing of their potential application domain and design is costly and time consuming. Models can evaluate alternative systems relatively quickly and at relatively low cost, but must first be validated to assess satisfactory simulation of the target systems. Specific climatic, edaphic, crop and fallow growth data was used from five sites in Western Kenya to calibrate and validate simulations of maize and improved fallow growth using the Water, Nutrient and Light Capture in Agroforestry Systems (WaNuLCAS) model. The model predicted continuous maize yields across the sites with an R ² of 0.72, an EF (model efficiency) of 0.66 and a CD (coefficient of determination) of 2.73, although the default pedotransfer functions (PTF) for volumetric soil water content used in the model had to be substituted for a tropical soils specific PTF before this was achieved. Predicted maize yield was consistently related to fallow biomass (i.e. higher fallow biomass correlated with higher subsequent maize yields) at two sites and the model predicted maize yields following fallow growth from this subset of the data with an R ² of 0.42. This relationship of fallow biomass to subsequent maize yield was not observed across the whole data set due to incomplete fallow litterfall data, factors not included in the model and associated poor model prediction of recycled tree biomass. After site and tree calibration, the model can thus be applied to assess fallow management strategies for sites limited by water and nitrogen.     

Modelling of planted legume fallows in Western Kenya. (II) Productivity and sustainability of simulated management strategies

Improved fallow is a technology that can help to raise agricultural productivity in systems of poor soil fertility and low financial capital. Models, once calibrated, can be used to investigate a range of improved fallow systems relatively quickly and at relatively low cost, helping to direct experimental research towards promising areas of interest. Six fallow crop rotations were simulated using the WaNuLCAS model in a bimodal rainfall setting in Kenya over a 10 year period: (A) alternating fallow and crop seasons, (B) one season fallow followed by three seasons crop, (C) one season fallow followed by four seasons crop, (D–F) 1–3 seasons fallow periods followed by 3–5 seasons crop. The strategies were tested using a number of fallow growth rates, soil clay contents, and rainfall amounts to determine the interaction of fallow rotation and biophysical variables on maize (Zea mays (L.)) yield and sustainability (organic matter, N₂ fixation, leaching). The best simulated fallow strategies doubled maize yield compared to continuous maize over a 10 year period. Across all biophysical treatments strategy A and B of no more than three consecutive cropping seasons and of one consecutive fallow season yielded the most maize. This was because fallow benefits were largely due to the immediate fallow soil fertility benefit (IFB) rather than the cumulative benefit (CFB). The difference in yield between the two strategies was through a balance between (1) their interaction with the biophysical variables affecting accumulation of organic matter, hence increasing soil fertility and (2) the extra intrinsic soil fertility used for maize productivity by the inclusion of more cropping seasons within the rotation. We propose the following conceptual framework to manage fallows for maximum maize yield: when environmental factors are strongly limiting to fallow and crop growth then fallow strategy A would be the best strategy to employ (less risk but more labour) and when factors are less limiting then strategy B would be the best to employ.     

Conversion of deciduous forest to silvopasture produces soil properties indicative of rapid transition to improved pasture

Differences in soil properties between forests and pastures have been well documented in the literature, especially under coniferous forests. However, since nearly all of these reports have been time-point comparisons, utilizing long-term paired-sites, properties of transitional states and time of their appearance can only be inferred at present. In this study, a deciduous forest ecosystem was converted to a silvopasture ecosystem by tree thinning, fertilization, and sheep incorporation of seed and forest litter. After 2 years, topsoil (0–15 cm) physico-chemical properties, particularly P fractions, and phosphatases were monitored over the growing season in these ecosystems, and a nearby pasture ecosystem. Initially, before spring vegetative growth, differences were found for pH, exchangeable cations and soil moisture, most of which could be explained by management history. Compared to forest, organic-C (C_o) and organic-N (N_o) concentrations were reduced in silvopasture by 17 and 9%, respectively, indicative of substantial litter decomposition. Most values for all these physico-chemical properties for silvopasture were intermediate between forest and pasture, and generally remained so throughout the growing season. Initial total P (TP_t), organic-P (TP_o) and inorganic-P (TP_i) concentrations were generally as anticipated for the forest and pasture. Silvopasture, however, had 36 and 23% greater TP_o than forest and pasture, respectively, presumably due to fertilizer-P immobilization induced by incorporation of forest litter. Total P components remained essentially constant over the growing season in all ecosystems, with the exception of pasture, likely due to high forage TP_i uptake. Bray I-extractable-organic-P (BrP_o) and bicarbonate-extractable-organic-P (BiP_o) concentrations, although consistently highest in the forest and silvopasture soils, were not reflective of the increase in TP_o under silvopasture. Acid phosphatase (PMEac) activities were highest in spring in all the ecosystems, then gradually declined to typically 25–50% initial activities. Alkaline phosphatase (PMEal) activities showed a broadly-similar pattern, with exception of forest and silvopasture soils, which exhibited low activities throughout the season. For the entire data set, PMEac and PMEal activities were poorly correlated with TP_i, BrP_i, and BiP_i. These results demonstrate that conversion of forest to silvopasture results in soil changes indicative of its rapid transition to pasture and that an increased P_o reservoir results that should be taken into account in fertilizer-P recommendations for temperate silvopastures.     

肯尼亚西部印度田菁改良休耕地对玉米生产力和黄独脚金寄生杂草(Sesbania sesban)侵扰的作用

在撒哈拉以南的非洲地区,黄独脚金寄生杂草（Strigahermonthica）侵扰是限制小农产自给性农业生产的主要因素之一。土壤肥力低加之总体环境退化是寄生杂草侵扰产生的重要原因。引入改良的耕作制度来解决寄生杂草侵扰和土壤肥力下降的问题势在必行。本文对肯尼亚西部双峰高原地区内,用豆科植物--印度田菁改良的休耕地对玉米产量和农田寄生杂草侵扰的作用进行了研究。实验处理分阶段进行,处理包括田菁改良6和18个月的休耕地、未经耕作自然植物再生6和18个月的休耕地、连续种植玉米未施肥的耕地和连续种植玉米同时施加氮和磷肥的耕地。结果表明,与未施肥玉米地相比,田菁改良休耕地明显（p〈0.5）增加玉米产量.除草管理降低了第一季度（428000&#177;63000株&#183;hm-2）、第二季度（51000&#177;1500株&#183;hm-2）玉米地寄生杂草植株种群。实验周期内,除草管理降低玉米地土壤中寄生杂草种子种群数。短期田菁改良休耕地对玉米产量的促进作用明显好于未施肥的玉米地,但是短期杂草休耕地对玉米产量无显著影响。种植玉米和除草控制寄生杂草效果要好于休耕。     

Effect of vegetation and land use on soil fertility in Wutai Mountains of Shanxi Province

IntroductionSoilpropertiesdependonclimate,vegetationtypes,parentmaterials,landformandsoilderivedage(Bei-jingForestryCoIlege1982).VegetationpIaysasig-nificantroIeintheformationofsoiIparticuIarIyforthepropertiespfSurfBcesoil.PlantsabsorbselectivelynutrientfromsoilandbuiIdtheirbodies.ThenutrientpartofIitterdecomposedgradualIybymicrobeswouldraturntoground-TheroOtsystemOfplantaIsoplaysasignificantroleinsoiIproperties.EffectofpIantsonsoildependonthevegdstiontype,speciescomposi-tion,age,dens…     

Simulated biomass and soil carbon of loblolly pine and cottonwood plantations across a thermal gradient in southeastern United States

Changes in biomass and soil carbon with nitrogen fertilization were simulated for a 25-year loblolly pine (Pinus taeda) plantation and for three consecutive 7-year short-rotation cottonwood (Populus deltoides) stands. Simulations were conducted for 17 locations in the southeastern United States with mean annual temperatures ranging from 13.1 to 19.4 °C. The LINKAGES stand growth model, modified to include the “RothC” soil C and soil N model, simulated tree growth and soil C status. Nitrogen fertilization significantly increased cumulative cottonwood aboveground biomass in the three rotations from a site average of 106 to 272 Mg/ha in 21 years. The equivalent site averages for loblolly pine showed a significant increase from 176 and 184 Mg/ha in 25 years with fertilization. Location results, compared on the annual sum of daily mean air temperatures above 5.5 °C (growing-degree-days), showed contrasts. Loblolly pine biomass increased whereas cottonwood decreased with increasing growing-degree-days, particularly in cottonwood stands receiving N fertilization. The increment of biomass due to N addition per unit of control biomass (relative response) declined in both plantations with increase in growing-degree-days. Average soil C in loblolly pine stands increased from 24.3 to 40.4 Mg/ha in 25 years and in cottonwood soil C decreased from 14.7 to 13.7 Mg/ha after three 7-year rotations. Soil C did not decrease with increasing growing-degree-days in either plantation type suggesting that global warming may not initially affect soil C. Nitrogen fertilizer increased soil C slightly in cottonwood plantations and had no significant effect on the soil C of loblolly stands.     

Ecosystem fertility and fallow function in the humid and subhumid tropics

The regeneration of natural vegetation (fallowing) is a traditional practice for restoring fertility of agricultural land in many parts in the tropics. As a result of increasing human population and insufficient fertilizer inputs, the ecosystem fertility functions of traditional fallows must now be improved upon via the use of managed fallows. Interactions between vegetation and soil determine nutrient losses and gains in crop—fallow systems and are influenced by fallow species, patterns and rates of biomass allocation, and crop and fallow management. Nutrient losses occur through offtake in crop harvests during the cropping phase and through leaching, runoff, and erosion in the cropping phase and the initial stage of fallows $#x2014; when nutrient availability exceeds nutrient demand by vegetation. Gains in nutrient stocks in later stages of fallow are generally more rapid on soils with high than low base status due to greater quantities of weatherable minerals and lack of constraints to N₂ fixation, deep rooting, and retrieval of subsoil nutrients by fallow vegetation. On low base status soils (exchangeable Ca < 1 cmol_c kg^–1), N₂ fixation and atmospheric inputs are likely to be the main sources of nutrient additions. On high base status soils limited by N, gains in N stocks by inputs from N₂ fixation and retrieval of subsoil nitrate can occur relatively rapidly; hence short-term fallows can often improve crop performance. Large losses of Ca associated with soil organic matter (SOM) mineralization and soil acidification during cropping and fallow establishment, combined with chemical barriers to root penetration, suggest that long-duration fallows (> 5 yr) are needed for recovery of cation stocks and crop performance on low base status soils. On both soils, however, residual benefits of fallows on crop yields usually last less than three crops.This revised version was published online in November 2005 with corrections to the Cover Date.     

Modelling soil organic carbon turnover in improved fallows in eastern Zambia using the RothC-26.3 model

Scarcity of simple and reliable methods of estimating soil organic carbon (SOC) turnover and lack of data from long-term experiments make it difficult to estimate attainable soil C sequestration in tropical improved fallows. Testing and validating existing and widely used SOC models would help to determine attainable C storage in fallows. The Rothamsted C (RothC) model, therefore, was tested using empirical data from improved fallows at Msekera in eastern Zambia. This study (i) determined the effects of nitrogen fixing tree (NFT) species on aboveground organic C inputs to the soil and SOC stocks, (ii) estimated annual net organic C inputs to the soil using the RothC, and (iii) tested the performance of RothC model using empirical data from improved fallows. Soil samples (0–20 cm) were collected from coppicing and non-coppicing fallow experiments in October 2002 for determination of SOC by LECO CHN-1000 analyser. Data on surface litter, maize and weed biomasses, and on weather, were supplied by the Zambia/ICRAF Agroforestry Project. Measured SOC stocks to 20 cm depth ranged from 32.2 to 37.8 t ha⁻¹ in coppicing fallows and 29.5 to 30.1 t ha⁻¹ in non-coppicing fallows compared to 22.2–26.2 t ha⁻¹ in maize monoculture systems. Coppicing fallows accumulated more SOC (680–1150 g m⁻² year⁻¹) than non-coppicing fallows (410–789 g m⁻² year⁻¹). While treatments with NFTs accumulated more SOC than NFT-free systems, SOC stocks increased with increasing tree biomass production and tree rotation. For food security and C sequestration, coppicing fallows are a potentially viable option.     

Species screening for short-term planted fallows in the highlands of western Kenya

Short-term improved fallow technology, which is characterised by deliberate planting of fast growing N₂ fixing legumes species in rotation with crops is currently being promoted for soil fertility replenishment in the small holder farms in the tropics. Recent research and extension efforts on this technology have mainly focused on a narrow range of species. There is a need to evaluate more alternative species in order to diversify the options available to farmers and hence reduce the risks of over dependence on fewer species. We evaluated twenty-two shrubby and herbaceous species for their site adaptability, biomass and nutrient accumulation, biomass quality and maize yield response to soil incorporated plant biomass after the fallow (six and twelve months) in three different field experiments on a Kandiudalfic Eutrudox in western Kenya. Species which yielded large amounts ofthe most biomass N adequate for two to three maize crops were Sesbania sesban, Tephrosia vogelii, Tephrosia candida, Crotalaria grahamiana, Dodonea viscosa, Colopogonium mucunoides, Desmondium uncinatum, Glycine wightii and Macroptilium atropurpureum. Most fallow species tested recycled <22 kg P ha^–1 in plant biomass. Significant amounts of K were recycled through plant biomass of Sesbania sesban, Tithonia diversifolia, Tephrosia candida, Crotalaria grahamiana, Dodonea viscosa, Colopogonium mucunoides, Desmondium uncinatum, Glycine wightii, Macroptilium atropurpureum and natural weed fallows. Recyclable K in plant biomass ranged between 4 and 188 kg ha^–1Two methods of establishing S. sesban and T. vogelii fallows did not result in significant differences in biomass and nutrient yields at the end of the fallow period. Shrubby species gave Hhigh lignin (>10%) and polyphenol (>2%) concentrations. were found only in the shrubby species, and the (Ppolyphenol + lignin ): N ratio varied widely (0.3–5) amongst the species. evaluated. Maize yield increased by two-fold in the first season following the fallow phase compared with continuous maize for most species. Results suggest that there are a wide variety of legumes that could be used for use in improved fallow technologies aimed at ameliorating nutrient degraded soils and subsequently enhancing crop yields.This revised version was published online in November 2005 with corrections to the Cover Date.     

Dynamics of soil carbon in a beechwood chronosequence forest

Accurate estimates of forest soil organic matter (OM) are now crucial to predictions of global C cycling. This work addresses soil C stocks and dynamics throughout a managed beechwood chronosequence (28–197 years old, Normandy, France). Throughout this rotation, we investigated the variation patterns of (i) C stocks in soil and humic epipedon, (ii) macro-morphological characteristics of humic epipedon, and (iii) mass, C content and C-to-N ratio in physical fractions of humic epipedon. The fractions isolated were large debris (>2000 μm), coarse particular OM (cPOM, 200–2000 μm), fine particular OM (fPOM, 50–200 μm) and the mineral associated OM (MaOM, <50 μm).     

Replacement of wildfire by whole-tree harvesting in jack pine forests: Effects on soil fertility and tree nutrition

Large areas of northern coniferous forests once naturally maintained by stand-replacing wildfires have shifted to an anthropogenic disturbance regime of clearcut harvesting followed by natural or artificial regeneration, with unknown consequences for soil biogeochemical processes. We used a comparative approach to investigate the effects of whole-tree harvesting (WTH) vs. stand-replacing wildfire (WF) on soil C and nutrient availability, and nutrition and growth of the succeeding stand, in jack pine (Pinus banksiana) forests of northern Lower Michigan. We compared total carbon (C), total nitrogen (N), potential N mineralization, and extractable phosphorus (P), potassium (K), calcium (Ca) and magnesium (Mg) among stands regenerated via WTH or WF in two age classes (4–7 years and 12–18 years). We also measured jack pine foliar nutrition and height growth in these same stands, as well as estimating the contribution of legacy dead wood to ecosystem nutrient capital in young stands. We found some evidence in support of our hypothesis that WTH would leave behind greater pools of soil C and N, but lower pools of P and base cations. However, the differences we observed were confined entirely to surface organic horizons, with the two disturbance regimes indistinguishable when viewed cumulatively to our maximum sampling depth of 30 cm. Estimates of nutrient pools in legacy wood inherited by young jack pine stands were also small in comparison to total soil pools (ranging from 1 to 9% depending on the element), suggesting that decomposition and nutrient release from this material is not likely to result in noticeable differences in soil fertility later in stand development. Similar levels of soil nutrients between WTH- and WF-origin stands were reflected in our measures of jack pine foliar nutrition and height growth, which were both unaffected by mode of stand origin. Results from this study suggest that soil nutrient levels following WTH fall within the natural range of variation produced by WF in these jack pine forests; however, comparison with a similar study on boreal jack pine suggests that latitudinal effects on O-horizon nutrient capital may influence the degree to which WTH matches the effects of WF on soil nutrient availability.     

A review on the potential of improved fallows and green manure in Rwanda

Agricultural production in the densely populated highlands of Rwanda is subject to serious soil fertility constraints. As the use of imported mineral fertilizers is beyond the economic means of resource-poor farmers, research and extension efforts of several projects, institutes and organizations concentrated during the last 15 years on the development and promotion of improved fallows with woody and herbaceous legumes, like Tephrosia sp., Cajanus sp., Crotalaria sp., Sesbania sp., Mucuna sp., and Mimosa sp., planted over one or more seasons as pure green manure, in hedgerows (alley cropping), or on fields as seasonal inter- or relay-crop. Green manuring proved to be a risky enterprise, due to highly variable biomass production and residual effects. Yield increments on-farm of up to 74% in the first season and 46% in the second season did not compensate loss of yields and labour investments during green manuring. Even where biomass production was sufficient, residual effects were in most cases unsatisfactory, due to rapid nutrient leaching (N, K) or inappropriate foliage incorporation on-farm. In researcher-managed trials, residual effects were in general somewhat higher, but more than a mere compensation of lost yields was not possible and farmers' adoption of these labour-intensive technologies was rather low. Due to acute land shortage, farmers were reluctant in allocating land to fallows or hedgerows also, with the exception of fields already out of production. Consequently, the concept of improving soil fertility and crop yields with the help of planted fallows or green manure in rotation failed. Woody legumes might have a future on abandoned fields and in wide spaced contour hedges, mainly for the production of firewood and bean stakes. For soil fertility management, the production and availability of farmyard manure and country-own mineral fertilizers, such as travertin and volcanic ashes should be supported. The question is raised as to whether sustainable agricultural development is possible without a credit system for small farmers, reallocating land and creating off-farm employment.     

Effects of the second-generation larch plantations onsoil fertility and tree growth

Larch is one of the most important plantation species in the northeast region of China. After clear cutting of larch plantations it is the urgent problem to besolved that whether the second generation is continuously plant6d, whether the soil fertility is decreasing and the plantstion is how to be managed onthe forestry produCtion. In odder tO realize the secondgenerahon Of larch plantstions and to provide thetheoretical base and the reasonable managementmeasures, we inventoried the groWth o…