Structure and function of traditional agroforestry systems in the western Himalaya. II. Nutrient cycling

Budget of nutrient in plant and soil, and their rates of recycling in three types of traditional agroforestry systems in mid elevations of the western Himalaya were studied. Concentration of nitrogen and potassium was significantly higher in fruits and leaves than in branches, boles, and litter. Grewia optiva, an important fodder tree, was found to be rich in potassium, vegetable crops in nitrogen, and weeds in phosphorus. Maximum store of nutrients was estimated in aboveground biomass of agrihortisilvicultural system: nitrogen 532, phosphorus 40, potassium 461, calcium 400, and magnesium 298 kg/ha; it was closely followed by that in agrihorticultural system, and both these had sbout 2-fold higher nutrient stock as compared to agrisilvicultural system. In the case of the former two systems, considerable quantity of nutrients up to nitrogen 169, phosphorus 14, potassium 165, calcium 97, and magnesium 65 kg/ha, for example in agrihortisilvicultural system, was recycled through debris of crops and weeds, and tree leaf litter, while the recycling rate was almost half in agrisilvicultural system. Through ‘harvest’ potassium was depleted in the largest quantity from the system, followed by nitrogen, magnesium, calcium, and phosphorus; much higher loss occurred through agricultural and/or horticultural edible parts as compared to fuel and fodder biomass. For example, the loss through agricultural crops and horticultural fruits for nitrogen in agrihortisilvicultural system was up to 75 percent and 38 percent respectively of the total annual uptake. This heavy loss makes the system unstable from a nutrient point of view particularly for nitrogen and potassium until external inputs are added through organic and inorganic fertilizers. Agrihortisilvicultural system is remarkably efficient in view of biomass productivity for fuel, fodder and fruits, and also from nutrient point of view provided losses through harvest are compensated externally.     

Shrub yield and fodder quality variations in a non-tropical dryland environment in West Asia

Integration of shrubs into the smallholder crop-rangeland-livestock farming systems in non-tropical dry areas could reduce feed gaps, rangeland degradation and desertification, but data on yield and fodder quality of most native and exotic shrubs are scanty. The study aimed at identifying shrubs for dryland agroforestry based on agronomic and fodder quality attributes. Fodder and wood yield, percent fodder (FBR) and fodder quality of 26 non-legume shrubs were determined from 6 to 8 months regrowth in north-west Syria. Seed yield was estimated from plants that were not cut. Fodder (31–87 Mg DM ha⁻¹), wood (16-2064 Mg DM ha⁻¹) and seed (0-132 Mg DM ha⁻¹) yield, FBR (24–87%), fodder concentrations of crude protein (69–195 g kg⁻¹.), acid detergent lignin (ADL) (24–109 g kg⁻¹), acid detergent fiber (ADF) (102–267 g kg⁻¹), neutral detergent fiber (NDF) (214–526 g kg⁻¹), in vitro organic matter digestibility (IVOMD) (391–526 g kg⁻¹), and in vitro gas production after 24 h of incubation (25–39 ml 200 mg⁻¹ DM) varied (P < 0.05) among the shrubs. Atriplex halimus-halimus and A. herba-alba from Spain, A. canescens, A. ploycarpa and A. lentiformis from USA, A. halimus and A. herba-alba from Syria and A. nummularia from Australia and South Africa had greater potential for development of dryland agroforestry technologies. The promising shrubs could be integrated into the rangeland-crop-livestock systems in non-tropical dryland environs to provide fodder, fuel-wood, shade, medicine, stabilize sand-dunes, and sequester carbon; thereby contributing to mitigation of rangeland degradation and global warming; if major constraints to adoption of fodder trees such as agronomic problems, low multipurpose value, land shortage and quality seed supply could be overcome.     

Carbon storage and nitrogen cycling in silvopastoral systems on a sodic in northwestern India

Tree-based land-use systems could sequester carbon in soil and vegetation and improve nutrient cycling within the systems. The present investigation was aimed at analyzing the role of tree and grass species on biomass productivity, carbon sequestration and nitrogen cycling in silvopastoral systems in a highly sodic soil. The silvopastoral systems (located at Saraswati Reserved Forest, Kurukshetra, 29°4prime; to 30°15prime; N and 75°15prime; to 77°16prime; E) consisted of about six-year-old-tree species of Acacia nilotica, Dalbergia sissoo and Prosopis juliflora in the mainplots of a split-plot experiment with two species of grasses, Desmostachya bipinnata and Sporobolus marginatus, in the subplots. The total carbon storage in the trees + grass systems was 1.18 to 18.55 Mg C ha⁻¹ and carbon input in net primary production varied between 0.98 to 6.50 Mg C ha⁻¹ yr⁻¹. Carbon flux in net primary productivity increased significantly due to integration of Prosopis and Dalbergia with grasses. Compared to 'grass-only' systems, soil organic matter, biological productivity and carbon storage were greater in the silvopastoral systems. Of the total nitrogen uptake by the plants, 4 to 21 per cent was retained in the perennial tree components. Nitrogen cycling in the soil-plant system was found to be efficient. Thus, It is suggested that the silvopastoral systems, integrating trees and grasses hold promise as a strategy for improving highly sodic soils. This revised version was published online in June 2006 with corrections to the Cover Date.     

Tree Performance and Root-Zone Salt Accumulation in Three Dryland Australian Plantations

Doubts exist about the effectiveness of establishing trees near saline discharge areas on farmland to manage dryland salinity. These centre on low rates of water uptake from saline water tables, salt accumulation in tree root zones and the consequent poor growth and survival of trees. Despite this, trees still survive in many plantations established adjacent to saline discharge areas and land-holders often favour such locations, as they do not compete for arable land such as that occurs with plantings in recharge areas. Tree performance and salt accumulation were assessed in three experimental plantations established adjacent to saline discharge areas 20–25 years ago. These were all in the 400–600 mm rainfall zone of south-western Australia. Mean soil salinity, within 1 m of the surface, ranged from 220 to 630 mS m⁻¹, while permanent ground-waters occurred within 2–5 m of the surface and had electrical conductivities ranging from 175 to 4150 mS m⁻¹. The study confirmed the low growth rates expected for trees established over shallow, saline water tables in a relatively low rainfall environment, with estimated wood volumes in Eucalyptus cladocalyx, E. spathulata, E. sargentii, E. occidentalis and E. wandoo of between 0.5 and 1.5 m³ ha⁻¹ yr⁻¹. Values of up to 3 m³ ha⁻¹ yr⁻¹ were obtained on soils with low salinity (<200 mS m⁻¹). The excellent survival (>70%) of several Eucalyptus species confirms that discharge plantations species can persist, despite increasing soil salinity. However, the long-term sustainability of such plantings (50–100 years) without broader landscape treatment of the present hydrological imbalance must be questioned.     

Production and phenology of the fruit shrub Eugenia stipitata in agroforestry systems in Costa Rica

Arazá (Eugenia stipitata), a fruit shrub originating from Western Amazonia, was evaluated growing in association with timber shade trees (Acacia mangium or Cordia alliodora) or with plantain (Musa sp.) as a potential commercial species for the tropical moist lowlands of Talamanca, Costa Rica. Height and crown width of the four-year-old shrubs varied between 2.7–2.8 m and 2.9–3.1 m, respectively. Flowering was positively correlated with initial fruit formation 1 month later and initial fruit formation with fully developed fruits a subsequent month after that. Three to four-year-old plants produced 20.0, 20.0, and 24.5 t ha⁻¹ yr⁻¹ (fresh fruits) in associations with A. mangium, C. alliodora and plantain, respectively, with higher production in rainy months. Fruit production in later years under C. alliodora (six to eight-year-old shrubs) was 26.5 t ha⁻¹ yr⁻¹. Where markets exist for E. stipitata fruit, the association can be recommended for tropical humid lowlands of Central America.     

Above- and belowground biomass,nutrient and carbon stocks contrasting an open-grown and a shaded coffee plantation

Coffee (Coffea canephora var robusta) is grown in Southwestern Togo under shade of native Albizia adianthifolia as a low input cropping system. However, there is no information on carbon and nutrient cycling in these shaded coffee systems. Hence, a study was conducted in a mature coffee plantation in Southwestern Togo to determine carbon and nutrient stocks in shaded versus open-grown coffee systems. Biomass of Albizia trees was predicted by allometry, whereas biomass of coffee bushes was estimated through destructive sampling. Above- and belowground biomass estimates were respectively, 140 Mg ha⁻¹ and 32 Mg ha⁻¹ in the coffee–Albizia association, and 29.7 Mg ha⁻¹ and 18.7 Mg ha⁻¹ in the open-grown system. Albizia trees contributed 87% of total aboveground biomass and 55% of total root biomass in the shaded coffee system. Individual coffee bushes consistently had higher biomass in the open-grown than in the shaded coffee system. Total C stock was 81 Mg ha⁻¹ in the shaded coffee system and only 22.9 Mg ha⁻¹ for coffee grown in the open. Apart from P and Mg, considerable amounts of major nutrients were stored in the shade tree biomass in non-easily recyclable fractions. Plant tissues in the shaded coffee system had higher N concentration, suggesting possible N fixation. Given the potential for competition between the shade trees and coffee for nutrients, particularly in low soil fertility conditions, it is suggested that the shade trees be periodically pruned in order to increase organic matter addition and nutrient return to the soil. An erratum to this article can be found at     

Dry matter production,morphology and nutritive value of <Emphasis Type="Italic">Dactylis glomerata</Emphasis> growing under different light regimes

Plant growth, morphology and nutritive value under shade can differ between temperate grasses. Therefore, the aim of this study was to quantify the dry matter (DM) production, sward morphology, crude protein (CP%), organic matter digestibility (OMD) and macro-nutrient concentrations (P, K, Mg, Ca and S) in a grazed cocksfoot (Dactylis glomerata L.) pasture under 10-year-old Pinus radiata D. Don forest. Four levels of light intensity were compared: full sunlight (100% photosynthetic photon flux density-PPFD), open + wooden slats (∼43% PPFD), trees (∼58% PPFD) and tree + slats (∼24% PPFD). The mean total DM production was 8.2 t DM ha⁻¹ yr⁻¹ in the open and 3.8 t DM ha⁻¹ yr⁻¹ in the trees + slats treatment. The changes in cocksfoot leaf area index (LAI) were related to variations in morphological aspects of the sward such as canopy height and tiller population. CP% increased as PPFD declined with mean values of 18.6% in open and 22.5% in the trees + slats treatment. In contrast, the intensity of fluctuating shade had little effect on OMD with a mean value of 79 ± 3.2%. The mean annual macro-nutrient concentrations in leaves increased as the PPFD level declined mainly between the open and the trees + slats treatments. It therefore appears that heavily shaded dominant temperate pastures in silvopastoral systems limit animal production per hectare through lower DM production rates and per animal through reduced pre-grazing pasture mass of lower bulk density from the etiolated pasture.     

Fine root dynamics in three central Himalayan high elevation forests ranging from closed canopied to open-canopied treeline vegetation

Fine root biomass, rates of dry matter production and nutrients dynamics were estimated for 1 year in three high elevation forests of the Indian central Himalaya. Fine root biomass and productivity were higher in closed canopied cappadocian maple forest (9.92 Mg ha⁻¹ and 6.34 Mg ha⁻¹ year⁻¹, respectively), followed by Himalayan birch forest (6.35 Mg ha⁻¹ and 4.44 Mg ha⁻¹ year⁻¹) and Bell rhododendron forest (6.23 Mg ha⁻¹ and 2.94 Mg ha⁻¹ year⁻¹). Both fine root biomass and productivity declined with an increase in elevation. Across the sites, fine root biomass was maximal in fall and minimal in summer. In all sites, maximum nutrient concentration in fine roots was in the rainy season and minimum in winter. Fine root biomass per unit basal area was positively related with elevation, Bell rhododendron forest having the largest fine root biomass per unit of basal area (0.53 Mg m⁻²) and cappadocian maple the least (0.18 Mg m⁻²). The production efficiency of fine roots per unit of leaf biomass also increased with elevation and ranged from 1.13 g g⁻¹ leaf mass year⁻¹ in cappadocian maple forest to 1.28 g g⁻¹ leaf mass year⁻¹ in Bell rhododendron forest. Present fine root turnover estimates showed a decline towards higher elevations (0.72 year⁻¹ in cappadocian maple and 0.58 year⁻¹ in Bell rhododendron forest) and are higher than global estimates (0.52).     

Interactions between Widely Spaced Young Poplars (Populus spp.) and Introduced Pasture Mixtures

Silvopastoral systems involving poplars are widespread in rural landscapes in a number of regions of New Zealand. The effect of widely spaced trees of Populus nigra × P. maximowiczii, aged 8–11 years, on the growth and botanical composition of understorey pasture mixes was determined over 3 years at a southern North Island hill country site. Pasture mixes comprised existing pasture, and two introduced test swards comprising new grass (Agrostis capillaris, Dactylis glomerata, Lolium perenne) and legume (Lotus uliginosus, Trifolium repens) cultivars. Pasture accumulation beneath trees (6.6 t DM ha⁻¹ year⁻¹) was 23% less than open (unshaded, no trees) pasture (8.6 t DM ha⁻¹ year⁻¹) and differences in accumulation occurred between tree aspects (North and South sides of trees) when trees were foliated. In early spring, North plots produced 11–14% more herbage than South plots whereas in late summer, South plots produced up to 44% more herbage than North plots. Yield of both test swards (6.4 and 8.0 t DM ha⁻¹ year⁻¹) was not significantly different from that of the existing sward (7.4 t DM ha⁻¹ year⁻¹) but productivity varied between swards in spring and summer. One test sward included Lotus uliginosus cv. Grasslands Maku, and the sward produced 30% more DM than the other swards in mid-summer. Dactylis glomerata cv. Grasslands Wana comprised 37% of the DM of swards in which it was sown and was the most successful cultivar, followed by Grasslands Maku. Both cultivars are recommended for silvopastoral systems where pasture is lightly grazed.     

<Emphasis Type="Italic">Pinus radiata</Emphasis> and sheep production in silvopastoral systems at Carngham,Victoria, Australia

This long term experiment provides production data for evaluation of combined forestry and livestock systems. Five systems were established in Pinus radiata planted in 1981 (620 mm average annual rainfall). Sheep were introduced in 1984 and trees were pruned in several lifts. Adjusted tree stockings were (1) 60 widely spaced trees ha⁻¹, (2) 200 widely spaced trees ha⁻¹, (3) 200 trees ha⁻¹ in five-row belts, (4) 1,090 trees ha⁻¹ (unpruned) and 815 trees ha⁻¹, and (5) no trees (open pasture). Tree growth, wool production, liveweight gain and pasture production were measured. At year 25, tree diameter under bark at 1.3 m (DBHUB) in Systems 1, 2, 3 and 4 (unpruned) was 46.0, 39.2, 33.5 and 24.1 cm, while volume of bark-free 6-m butt-logs was 49, 117, 86 and 233 m³ ha⁻¹. Inner rows of System 3 belts contained smaller trees than outer rows. Pruned System 4 trees had slightly greater diameter than unpruned trees. Pasture production declined with tree stocking and time, due to shading and competition. Wool production (WP ha⁻¹) and liveweight gain (LWG ha⁻¹) declined linearly from year 9 to 17 with increasing disparity among systems. In 1998 (year 17) WP ha⁻¹ in Systems 1, 2 and 3 was 64, 16 and 43% of that in open pasture. Further analysis is needed to evaluate the financial costs and returns of various systems under particular rotation lengths and market prices.     

Pasture production under different tree species and densities in an Atlantic silvopastoral system

We studied the effect of six tree species planted at six different densities on pasture production seven years after establishment. Annual and seasonal pasture production was studied every six months, over three years. Pasture production was lower under conifer trees (Pseudotsuga menziesii (Mirb.) Franco, Pinus pinaster Aiton, Pinus radiata D. Don) than under broadleaved trees (Betula alba L., Quercus rubra L. and Castanea sativa Mill.). Annual pasture production under Pseudotsuga menziesii and Pinus pinaster decreased progressively starting from 952 trees ha⁻¹, while decline in herbage production under Pinus radiata began to occur at 427 trees ha⁻¹. Tree density effect on pasture production was detected at 2,000 trees ha⁻¹ for all of the deciduous species studied. This effect on pasture production was more important in the first six months of the year (June sampling), while from June to December herbage production was less affected by tree density. The tree effect became more noticeable over time, with the last sampling showing the inverse relationship between tree density and herbage production most clearly. Seven years after tree establishment, pasture production was quite consistent under tree densities between 190 trees ha⁻¹ and 556 trees ha⁻¹ and declined remarkably from 556 trees ha⁻¹ to 2,500 trees ha⁻¹. The study also indicated that by the sixth growing season, annual pasture production under different tree species is inversely correlated with tree leaf area index.     

Biomass production and carbon stocks in poplar-crop intercropping systems: a case study in northwestern Jiangsu,China

The importance of agroforestry systems in CO₂ mitigation has become recognized worldwide in recent years. However, little is known about carbon (C) sequestered in poplar intercropping systems. The main objective of this study is to compare the effects of three poplar intercropping designs (configuration A: 250 trees ha⁻¹; configuration B: 167 trees ha⁻¹ and configuration C: 94 trees ha⁻¹) and two intercropping systems (wheat–corn cropping system and wheat–soybean cropping system) on biomass production and C stocks in poplar intercropping systems. The experiment was conducted at Suqian Ecological Demonstration Garden of fast-growing poplar plantations in northwestern Jiangsu. A significant difference in C concentration was observed among the poplar biomass components investigated (P ≤ 0.05), with the highest value in stemwood and the lowest in fine roots, ranging from 459.9 to 526.7 g kg⁻¹. There was also a significant difference in C concentration among the different crop components (P ≤ 0.05), and the highest concentration was observed in the corn ear. Over the 5-year period, the total poplar biomass increased with increasing tree density, ranging from 8.77 to 15.12 t ha⁻¹, while annual biomass production among the crops ranged from 4.69 to 16.58 t ha⁻¹ in the three configurations. Overall, total C stock in the poplar intercropping system was affected by configurations and cropping systems, and configuration A obtained the largest total C stock, reaching 16.7 t C ha⁻¹ for the wheat–soybean cropping system and 18.9 t C ha⁻¹ for the wheat–corn cropping system. Results from this case study suggest that configuration A was a relative optimum poplar intercropping system both for economic benefits and for C sequestration.     

Growth and yield of maize and timber trees in smallholder agroforestry systems in Claveria,northern Mindanao,Philippines

On-farm experiments were conducted in the Philippines to study over a 4-year period the growth of two timber trees, gmelina (Gmelina arborea R. Br.) and bagras (Eucalyptus deglupta Blume), and their impact on the grain yield of intercropped maize. The experiment consisted of maize monocropping plots (control) and maize intercropped between trees planted in block (2 × 2.5 m), and hedgerow arrangement (1 × 10 m). Three maize crops were planted in the block plots before canopy closure, and seven maize crops were planted in the hedgerow and monocropping plots. Maize grain yield in the hedgerow and in the block arrangement with gmelina were respectively 37% (16.58 tons ha⁻¹) and 68% (8.3 tons ha⁻¹) lower than in monocropping (26.21 tons ha⁻¹). In the plots with bagras, maize grain yield in hedgerow and in block arrangement were respectively 19% (24.8 tons ha⁻¹) and 66% (10.4 tons ha⁻¹) lower than in monocropping (30.6 tons ha⁻¹). For both tree species, the diameter at breast height (dbh) was greater in hedgerow than in block arrangement, with the difference being more pronounced with age. It was estimated that gmelina planted in hedgerows would produce 6–8 m³ ha⁻¹ of merchantable volume more than if planted in block. The study verifies the hypothesis that intercropping between widely-spaced trees rows (planted at 10 m or more) is more profitable and feasible to smallholders than either maize monocropping or woodlots, and concludes with recommendations on how to further improve the productivity of tree-intercropping systems.     

Intercropping field crops between rows of Leucaena leucocephala under rainfed conditions in northern India

A field study was conducted for six years (1981–1986) on sandy loam soil on intercropping hedgerows of Leucaena leucocephala (Lam) de Wit. with three field crops viz. maize (Zea mays L), black gram (Vigna mungo L) and cluster bean (Cyamopsis tetragonoloba L Taub.). In treatments 1 and 2 Leucaena hedges were planted as pure crops at close (25 cm × 75 cm) and wide (25 cm × 375 cm) spacings. In treatments 3, 4 and 5 the three field crops were intercropped between the hedgerows of Leucaena at the wide spacing, and in treatments 6, 7 and 8 the field crops were raised as pure crops. Leucaena was topped to 75 cm each time it attained a height of 175 cm. The pure crop of Leucaena at close spacing produced an average, over the six years, of 34 t ha⁻¹a⁻¹ of green fodder and 9.4 t ha⁻¹a⁻¹ of air dry fuelwood. The Leucaena at wide spacing produced 18.9 t ha⁻¹a⁻¹ of green fodder and 6.3 t ha⁻¹a⁻¹ of fuelwood. Intercropping with field crops decreased the yield of green fodder and fuelwood. The yield of all the field crops was less when raised as intercrops than as pure crops. Mean maximum net returns were obtained from intercrops of Leucaena and cluster bean (Rs 3540 ha⁻¹a⁻¹) which were significantly higher than the returns from pure crop of Leucaena at wide spacing but similar to the returns from pure crops of cluster bean. Leucaena with maize (Rs 3273 ha⁻¹a⁻¹) and black gram (Rs 3125 ha⁻¹a⁻¹) gave significantly higher net returns over pure crops of Leucaena at wide spacing, maize and black gram. ICRISAT = International Crops Research Institute for the Semi-Arid Tropics- Hyderabad, India. CIAT = Centro International de Agricultura Tropical - Cali - Columbia     

Structure and Function of <Emphasis Type="Italic">Populus deltoides</Emphasis> Agroforestry Systems in Eastern India: 1. <Emphasis Type="Italic">Dry matter dynamics</Emphasis>

Agroforestry systems based on poplar (Populus deltoides) are becoming popular in eastern and northern parts of India. Therefore studies on the structure and function of the systems are important. The investigations included allometric equations for above- and belowground tree components, crop and plantation floor biomass and litter fall estimation at Pusa, Bihar, India. Biomass, floor litter mass, litter fall and net primary productivity (NPP) of plantations increased with an increase in age of trees whereas, crop biomass for any specific crop interplanted with poplar decreased with the age of the plantation. The total plantation biomass increased from 12.08 to 90.59 Mg ha⁻¹ and NPP varied from 5.69 to 27.9 Mg ha⁻¹ year⁻¹. The biomass accumulation ratio ranged from 2.1 to 3.2. Total annual litter fall was in between 1.95 and 10.00 Mg ha⁻¹ year⁻¹, of which 92–94% was contributed by leaf litter. Compartmental models were developed for dry matter distribution in agroforestry systems involving young (3-year-old) and mature (9-year-old) poplar trees interplanted with various crops, the crops being grown in two rotations maize (Zea mays) – wheat (Triticum aestivum) – turmeric (Curcuma domestica) and pigeonpea (Cajanus cajan) – turmeric. This study substantiates the potential of Populus deltoides G₃ under agroforestry combinations.     

Growth and ecological impacts of traditional agroforestry tree species in Central Himalaya, India

A number of multipurpose tree species are conserved as scattered trees in settled farms on terraced slopes by the traditional farmers in Central Himalaya, India. Knowledge on growth rates and ecological impacts of these tree species is limited. Ten locally valued multipurpose tree species, viz., Albizzia lebbek, Alnus nepalensis, Boehmeria rugulosa, Celtis australis, Dalbergia sissoo, Ficus glomerata, Grewia optiva, Prunus cerasoides, Pyrus pashia and Sapium sebiferum, were established as mixed plantations at a degraded community forest land site and an abandoned agricultural land site in a village at 1200 m altitude in District Chamoli, India. At the abandoned agricultural land site, annual food crops were grown, along with planted trees, providing supplemental irrigation and organic manure following traditional farming practices. Survival, height, stem circumference, crown depth and width, number of branches, above-ground biomass and soil physico-chemical characteristics were monitored up to five years of plantation growth. Above-ground tree biomass accumulation at the abandoned agricultural land site was 3.9 t ha⁻¹ yr⁻¹ compared with 1.1 t ha⁻¹ yr⁻¹at the degraded forest land site. B. rugulosa, C. australis, F. glomerata, G. optiva, P. cerasoides and S. sebiferum showed more prominent differences in growth at the two sites compared with A. lebbek, A. nepalensis, D. sissoo and P. pashia. A. nepalensis and D. sissoo showed best growth performance at both the sites. A significant improvement in soil physico-chemical characteristics was observed after five years at both of the sites. Carbon sequestration in soil was higher than that in bole biomass. This revised version was published online in June 2006 with corrections to the Cover Date.     

Biomass production and C-sequestration of Gmelina arborea in plantation and agroforestry system in India

Tree based land use systems make a valuable contribution to sequester carbon and improve productivity and nutrient cycling within the systems. This study was conducted to determine biomass production, C-sequestration and nitrogen allocation in Gmelina arborea planted as sole and agrisilviculture system on abandoned agricultural land. At 5 years, total stand biomass in agrisilviculture system was 14.1 Mg ha⁻¹. Plantations had 35% higher biomass than agrisilviculture system. At 5 years, leaves, stem, branches and roots contributed 4.1, 65.2, 10.0 and 20.7%, respectively to total standing biomass (17.9 Mg ha⁻¹). Over the 5 years of study, trees had 3.5 Mg ha⁻¹ more C and 36 kg ha⁻¹ more N in plantation than agrisilviculture system. Biomass and C storage followed differential allocation. Relatively more C was allocated in above ground components in plantations compared to agrisilviculture system. C:N ratios for tree components were higher in stem wood (135–142) followed by roots (134–139), branches (123–128) and leaves (20–21). In agrisilviculture system crops recommended are: soybean and cowpea in rainy season; wheat and mustard in winter season. After 5 years, soil organic C increased by 51.2 and 15.1% and N by 38.4 and 9.3% in plantation and agrisilviculture system, respectively. Total C storage in abandoned agricultural land before planting was 26.3 Mg ha⁻¹, which increased to 33.7 and 45.8 Mg ha⁻¹ after 5 years in plantation and agrisilviculture system, respectively. Net C storage (soil + tree) was 7.4 Mg ha⁻¹ in agrisilviculture system compared to 19.5 Mg ha⁻¹ in G. arborea monoculture stands. The studies suggest that competitive interactions played a significant role in agrisilviculture system. Plantations were more efficient in accreting C than agrisilviculture system on abandoned agricultural land.     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica II. Cacao and wood production,litter production and decomposition

During 7 years (1979–1985) cacao harvests (beans and husks) have been recorded for the agroforestry systems ofTheobroma cacao underCordia alliodora andErythrina poeppigiana shade trees. The mean oven dry cacao yields were 626 and 712 kg.ha⁻¹.a⁻¹ cocoa beans underC. alliodora andE. poeppigiana respectively. Harvests have gradually increased over the years and the plantation has now reached maturity. Annual extraction of N, P, K, Ca and Mg in fruits, which is relatively small, was calculated on the basis of chemical analyses. The following average values were found (kg.ha⁻¹.a⁻¹): At the age of 8 years, theC. alliodora trees have reached 26.7 cm diameter (DBH) and 14.0 m in height. Mean annual growth (from age 5 to 7) is 14.6 m³.ha⁻¹.a⁻¹. Natural plant residue production has been measured for 4 years (Nov. 1981–Oct. 1985). UnderE. poeppigiana it has reached a value of 8.91 t.ha⁻¹.a⁻¹ and underC. alliodora 7.07 t.ha⁻¹.a⁻¹. The shade trees have contributed 57 and 47% respectively. Transference and decomposition rates are high and important in the nutrient cycles. The nutrient content of the litter was analysed and corresponding average yearly transfers were (kg.ha⁻¹.a⁻¹): For part I see Vol. 4, No. 3, 1986. Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesselschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Effect of wide spacing on increment and branch properties of young Norway spruce

The effects of stand density on increment and branch properties were studied in three spacing experiments of Norway spruce [Picea abies (L.) Karst.]. The stand densities ranged from 350 stems ha⁻¹, regarded as open-grown trees, up to 1,600 stems ha⁻¹, corresponding to the density recommended for forestry practice. Properties of all the branches were measured from the stem apex downwards. The study material included a total of 5,661 branches from 45 trees. Increasing stand density resulted in a decrease in radial increment as well as shorter and narrower crowns, but it had no effect on height increment. The average number of spike knots per tree was 0.87, 0.27, and 0.33 in densities of 350, 700 and 1,600 ha⁻¹, respectively. Additionally, in the widely spaced stands of 350 stems ha⁻¹, the fraction of trees having spike knots was high (over 50%). At a density of 1,600 ha⁻¹, the sample trees had somewhat less branches in a whorl compared with the more widely spaced plots. The most pronounced effect of stand density was the increase in branch diameter with decreasing stand density. At a density of 350 ha⁻¹, the maximum branch diameter of all the sample trees exceeded the diameter limit of quality class B in the European quality requirements for round wood. The results give some indication that trees subjected to severe competition would produce smaller branches per unit of crown projection area. However, the possibilities for reducing branch dimensions relative to stem and crown size through competition appear quite restricted.     

Nitrogen and fine root length dynamics in a tropical agroforestry system with periodically pruned Erythrina poeppigiana

The effect of pruning all branches (complete pruning) or retaining one branch (partial pruning) on the dynamics of nitrogen cycling in aboveground biomass, nitrogen supplying power of an amended Eutric Cambisol, and fine root length, was studied in an Erythrina poeppigiana (Walp.) O.F. Cook—tomato (Lycopersicon esculentum Mill.) alley cropping practice in Turrialba, Costa Rica during 1999–2000. Over the 1 year pruning cycle, in which trees were completely or partially pruned four times, respective aboveground biomass production was 4.4 Mg or 7 Mg ha⁻¹ (2-year-old trees) and 5.5 Mg or 9 Mg ha⁻¹ (8-year-old trees); N cycled in aboveground biomass was 123 kg or 187 kg ha⁻¹ (2-year-old trees) and 160 kg or 256 kg N ha⁻¹ (8-year-old trees); mean fine root length was 489 or 821 m (2-year-old-trees), 184 or 364 m per tree (8-year-old-trees). Pruning intensity did not significantly affect net N mineralisation and net nitrification rates during the tomato-cropping season. For the tomato crop, pre-plant mean net N mineralisation rate of 2.5 mg N kg⁻¹ soil day⁻¹ was significantly lower than 16.7 or 11.6 mg N kg⁻¹ soil day⁻¹ at the end of vegetative development and flowering, respectively. Mean net nitrification rates of 3.5, and 4.3 mg N kg⁻¹ soil day⁻¹, at pre-plant and end of vegetative development, respectively, were significantly higher than 0.3 mg N kg⁻¹ soil day⁻¹ at end of flowering. In humid tropical low-input agroforestry practices that depend on organic inputs from trees for crop nutrition, retention of a branch on the pruned tree stump appears to be a good alternative to removal of all branches for reducing N losses through higher N cycling in aboveground biomass, and for conserving fine root length for higher N uptake, although it might enhance competition for associated crops.