An evaluation of silvopastoral systems in pine plantations in the Central Highlands of Ecuador

One of the most serious problems facing Andean nations today is population growth and the resultant pressures placed on limited natural resources. Currently,thousands of hectares of pine plantations in the central highlands of Ecuador are left unmanaged and unutilized, while at the same time local agriculturalists lack grazing land,firewood and other basic necessities which could be provided by appropriate forest management. This paper evaluates ten pine plantations in central Ecuador for their potential as silvopastoral systems, and discusses the possibilities of integrating local farmers into the appropriate management and use of them.

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Resúmen Uno de los problemas mas graves afrontando a los paises Andinos es el crecimiento de la población y las presiones que resultan sobre los recursos naturales. Al momento, miles de hectáreas de plantaciones de pino en la Sierra central del Ecuador se quedan sin que nadie les maneje ni les utilize; mientras tanto los agricultores cercanos sufren por falta de pastoreo, leña y otras necesidades que se podría conseguir a través del manejo forestal. Este documento examine diez rodales de pino en el Ecuador por su potencial como sistemas silvopastoriles, y discute las posibilidades de integrar agricultores pequeños dentro del manejo y uso apropriado de ellos.

     

Contribution of Erythrina protein banks and rejected bananas for improving cattle production in the humid tropics

In view of the low pasture productivity in the Central American humid tropics where cattle rearing is a major land-use activity, it is important to examine the potential of alternate feed sources for ruminant feeding. Erythrina berteroana Uban, locally known as poro, and green banana (Musa AAA) fruits have been identified as two such potential sources. The effects of feeding poro as a grazing supplement and diet supplementation with green banana fruits on cattle productivity were evaluated in a long-term trial in Costa Rica. Daily liveweight (LW) gains were measured and samples of all feed material were analyzed for crude protein and in vitro dry matter (DM) digestibility. Pasture availability was high, averaging 0.17 t DM ha day^–1 over the grazing period. The main grass species were Paspalum fasciculatium Poiret, Axonopus compressus Swartz and to a lesser extent African star grass (Cynodon nlemfluensis Vanderyst). DM yields of poro declined significantly with time (> 50%) when it was managed with a two-month resting period but remained higher when subsequently managed with a three-month resting period. Average daily LW gain of animals was 21% to 26% higher with two hours daily browsing of poro than for animals only grazing pasture. Highest liveweight gain was achieved when diets were supplemented with banana and there was no additional benefit when poro was fed in addition to banana. This suggests that fodder banks of poro or supplements with green banana can improve cattle nutrition in the humid tropics.This revised version was published online in November 2005 with corrections to the Cover Date.     

Decomposition of woody legume nodules in two tree/grass associations under contrasting environmental conditions

Turnover of nitrogen-rich root nodules follows the pruning of legume trees, forming a potentially important yet little studied way of N release to the soil. The effects of soil moisture, herbivory by soil mesofauna and microbial decomposition on the disappearance rate of woody legume nodules was studied in two tree/grass forage production associations. Litter bags containing nodules of Erythrina variegata L. (Papilionoideae: Phaseoleae) were incubated for four weeks in grass-covered alleys between Gliricidia sepium (Jacq.) Walp. (Papilionoideae: Robinieae) hedgerows, established on a deep alluvial Oxisol under a humid tropical climate and on a shallow Vertisol under a subhumid tropical climate in Guadeloupe, French Antilles. Soil moisture was regulated by irrigating or covering small plots from natural rainfall. Fine nylon mesh bags were used to study the rate of microbial decomposition, and open-ended perforated cylinders were used to estimate nodule herbivory. The chemical traits, especially the lignin: nitrogen ratio, of E. variegata and G. sepium nodules were similar (lignin: N 1.70 and 1.55, respectively), and suggest that the results are probably also applicable to the G. sepium nodules in the associations. Both soil moisture and decomposing agent (microbes or mesofauna) had a significant effect on the nodule disappearance rate, but soil type did not have any apparent effect. The nodule half-life varied from three to seven days under different treatments. The N release rate from the nodules was high, with N half-life varying from three to five days. Herbivory accounted for ca. 10% of total mass and N loss from nodules during the four-week field incubation period, but its importance increased towards the end of the incubation, especially in Vertisol, after the most easily decomposable part of the nodules had decayed. After pruning, the nodule N is released to soil more rapidly than from mulch. This revised version was published online in June 2006 with corrections to the Cover Date.     

The espinal: agroforestry systems of the mediterranean — type climate region of Chile

The Central Valley of Chile is largely occupied by a pseudo-savanna called espinal consisting of a single dominant tree species, Acacia caven (Mol.) Mol. (Leguminosae), and some 215 annuals, most of which were inadvertently introduced from the Mediterranean Basin. The probable origin and current distribution of the espino (Acacia caven) and the espinales in Chile are described. Predominant production systems of the espinal in both the semiarid and subhumid regions of Chile are outlined, with emphasis on the later region. Causes of the current low levels of productivity are discussed, and various strategies for long-term improvement of the espinal are presented.     

On-farm research in a silvopastoral project: a case study

The Silvopastoral Project is being implemented by CATIE in the Atlantic humid lowland of Costa Rica, in order to develop alternatives to the current destructive mode of livestock production. The overall approach is briefly described and two superimposed on-farm experiments are discussed in more detail. The main constraints for on-farm research within the project are analysed: the limited previous knowledge of the area and of the local species, the difficulty of achieving a common understanding about research with collaborating farmers, the need for additional staff supervision and the complexity of the statistical analyses. An attempt is made to draw lessons from the ongoing project regarding how to resolve the apparent contradiction between rigorous scientific requirements and questions of immediate value to extension, the compromises to be achieved between different types of evaluation (statistical, risk, etc.) and the potentials and limitations of farmers' participation in research.     

Productivity of two Douglas fir/subclover/sheep agroforests compared to pasture and forest monocultures

Resource sharing between tree and forage plant components in silvopastoral systems includes a complex set of facilitative and competitive interactions. To the extent that facilitation exceeds competition, agroforests are expected to outyield monocultures of their components. Pasture and Douglas fir (Pseudotsuga menziesii) tree production of young agroforests was compared to pasture and forest monocultures under both grid and cluster patterns of tree planting near Corvallis, Oregon, USA, during 1983–1987. The height and diameter growth of forest and agroforest trees was similar, regardless of tree planting pattern. Five-year average annual forage production was 6500, 5800, and 2800 kg ha^–1 on pasture, agroforest, and forest plots, respectively. The total cumulative 1982–1987 above-ground phytomass yield of forage plus trees was similar for pasture and conventional grid forest monocultures. The total productivity of agroforests, however, was over 30% greater than either pasture or forest components grown in monoculture. Approximately 1.6 ha (0.96 ha forest + 0.64 ha pasture) of monocultures would be needed to equal the productivity of 1 ha of agroforest.Oregon Agricultural Experiment Station Technical Paper No. 10, 825.     

Management of secondary tree species in agroforestry systems to improve production sustainability in Amazonian Ecuador

Three production systems (robusta coffee, cattle pastures and swidden agriculture), all associated with trees, have provided a strategy for regional development in the Ecuadorian Amazon. Feedback from more than 250 on-farm demonstrations, project evaluations and visitors have all contributed to strengthening the ability of the Ecuadorean Ministry of Agriculture's Forestry Directorate to provide colonists and indigenous groups with management guidelines for low-input sustainable agroforestry production. The existing market for 20 fastgrowing native tree species presents an opportunity for the diversification of farm income (currently dependent on robusta coffee) and for the development of a forest industrial base.The collective area of land owned and operated by these small holders constitutes a national resource and a Collective Forest Estate that can be individually managed. Commercial timber production in these extensive secondary forests can help alleviate conversion of primary forest without taking agricultural land out of production.     

Cattle grazing under Pinus caribaea

An analysis is made of the animal stocking rate used in commercial Pinus caribaea plantations 2.5–8.5 yr old. Pines were planted at 2.5 × 2.5 m and programmed for clear-cut harvest at 10 yr rotations (pulpwood). Cattle grazing was introduced to reduce weeding costs.Stocking rate in 1983–1985 was 0.67 AU/ha/yr (AU = animal units 350 kg live weight). A quadratic model calculated using 1984 data (S = 0.0108 + 0.385A–0.0418A²) describes the relationship between animal stocking rate (S) and stand age (A 9.5 in yr).The model predicts a maximum of 0.89 AU/ha/yr at plantation age 4.5 yr. Grazing started at plantation ages 2.5 yr; no plantations older than 9.5 yr are grazed.     

Woody legumes as live support systems in yam cultivation

This paper reports on the use of three leguminous perennials (Leucaena leucocephala, Flemingia macrophylla and Gliricidia sepium) as live support systems in yam cultivation. In a planting arrangement in which yam rows alternated with rows of the woody species the tuber yields per ha were 3.4 (leucaena), 5.3 (flemingia) and 10.1 (gliricidia) ton fresh weight. TUKEY's L.S.D. value equalled 2.9 ton. Leucaena leucocephala is unsuitable as live support since the species shows a strong competitive power expressed in terms of leaf productivity and relative density of the root mass in the upper soil stratum, the zone explored by the yam crop. Flemingia macrophylla is unsuitable as support species mainly because of its structural weakness: Its branches do not sufficiently lignify to carry the yam leaf mass. The significantly higher tuber yield of the yam crop grown with Gliricidia sepium is a function of specific properties of the tree species: Low leaf productivity, a relatively weakly developed root system and an open architecture, which leaves sufficient space for a yam crop grown in association.     

Comparative performance of four multi-purpose trees associated with four grass species in the humid regions of Southern India

The results of a field trial conducted at the Livestock Research Station, Thiruvazhamkunnu, Kerala to study the compatibility of different components in a silvo-pastoral system revealed that growth and yield of fodder species were significantly influenced by the tree components only after tree canopy formation. The fodder species such as Pennisetum purpureum Schum., Panicum maximum Jacq., Brachiaria ruziziensis Griseb. and Euchlaena mexicana Schrad. grown in association with Casuarina equisetifolia J. R. & G. Forst. and Ailanthus malabarica DC recorded comparatively higher forage yield even after canopy formation. However, forage crops grown in combination with Acacia auriculiformis A. Cunn. ex Benth. and Leucaena leucocephala (Lam.) de Wit. registered relatively lower values for growth and yield. Increased light infiltration into the understorey due to the cladophyllous canopy can be attributed as reason for the higher fodder productivity under Casuarina. Due to the combination of crown size and shape, tree height and spacing the amount of light intercepted by Ailanthus also was very low. Among the four multi-purpose trees used, Acacia recorded the maximum growth rate followed by Casuarina, Ailanthus and Leucaena. Forage productivity of the four species was in the order: Pennisetum purpureum > Panicum maximum > Brachiaria ruziziensis > Euchlaena mexicana. Casuarina with Pennisetum/ Panicm were found to be optimal tree — forage combinations for silvo-pastoral systems.     

Manual recording of minirhizotron data and its application to study the effect of herbicide and nitrogen fertiliser on tree and pasture root growth in a silvopastoral system

The use of a simple counting method using a borescope and minirhizotron system to study the interaction of roots of wild cherry trees (Prunus avium L) and pasture in an upland silvopastoral system is described. Operator variation was greatest when distinguishing different tree root categories (e.g. white, brown, woody) but was not significant in relation to total tree or grass root counts or species differences. Analysis of covariance was used to take these differences into account in a field trial using several operators. The seasonal patterns of tree and pasture root growth were different. Tree roots peaked in May before maximal shoot growth. The peak in pasture roots occurred two weeks earlier. Tree height growth was increased by the use of herbicides to control grass growth but this effect was only apparent during August when the soil was dry and when pasture root counts were reduced.     

Effects of understorey vegetation on tree establishment and growth in a silvopastoral system in northern Greece

The interaction between understorey plants and trees in a young silvopastoral system was studied in a sub-humid Mediterranean environment with cold winters in northern Greece. The experimental design was a split-split plot with three replications and included: three understorey treatments (grass, legume and control), two tree species (Acer pseudoplatanus and Pinus sylvestris) and two spacings (2.5×2.5 m and 3.5×3.5 m). The competition between herbaceous plants and trees resulted in significant differentiation in tree growth while their spacing did not produce any significant differences. The influence of trees on herbage yield either by plant species or spacing was not significant. During the early stages of establishment, a significant positive correlation was observed between sycamore growth and soil moisture, leaf weight, leaf area and leaf number as well as nitrogen and potassium concentration in leaves. Six years after planting the height increase of the sycamore trees was largest in the control treatment (415%), lowest in the grass treatment (134%) and intermediate (192%) in the legume treatment. Much higher was the increase obtained for the diameter, 161%, 207% and 536% respectively for the grass, legume and control treatments. The Scots pine trees grew faster than sycamore over the course of the experiment. Height increased by 397%, 351% and 400% and diameter by 518%, 443% and 683% respectively for the grass, legume and control treatments. This revised version was published online in June 2006 with corrections to the Cover Date.     

Survival and development of immature Harmonia axyridis (Pallas) feeding on Chaitophorus populeti (Panzer) propagated on transgenic Populus alba × P. glandulosa

Laboratory feeding experiments with the poplar aphid, Chaitophorus populeti (Panzer), feeding on transgenic poplar (P. alba × P. glandulosa) varieties C13-5 and C013-5, were carried out to study the effect of transgenic poplar on the ladybird Harmonia axyridis (Pallas). The mortality and development time of the immature stages, the eclosion rate and body mass of H. axyridis were measured. The results indicated that C. populeti feeding on different varieties of transgenic plants had no statistically significant effect on the mortality of H. axyridis larvae. The development time of larval and pupal stages were not significantly different between the two transgenic poplars and a non-transgenic poplar. Furthermore, the body mass and eclosion rate did not show any difference between the H. axyridis feeding on aphids reared on transgenic plants and those from non-transgenic plants. It is suggested that transgenic plants have no deleterious effect on the predatory ladybird.     

寄生小蜂对小青×黑杨树皮及杨圆蚧挥发油的行为反应

Four-armed airflow olfactometer was used to determining the foraging behavior ofPteroptrix longgiclava (Girault) (Hymenoptera: Aphelinidae) andEncarsia gigas (Tshumakova) (Hymenoptera: Aphelinidae) to the essential oils which emitted from the healthy bark ofPopulus pseudo-simonii×P. nigra, the infested bark injured byQuadraspidiotus gigas (Thiem & Gerneck), the body and scale of fixed 1st-instar-nymph ofQ. gigas. The results from these experiments showed that the volatile oils produced from the injured bark and from the scale of fixed 1st-instar-nymph had a higher attractive ability to female adults of the two species of wasps. The essential oil produced from the scale of the pest at dosages of 3–7 μL and the essential oil emitted by injured bark at dosages of 5–9 μL had a stronger alluring effect on the host searching behavior ofPteroptrix longgiclava. The essential oil from the body of fixed 1st-instar-nymph ofQ. gigas also had certain effect on the host locating effort ofPteroptrix longgiclava andEncarsia gigas. Those two wasps did not shown any reaction to the essential oil produced by the healthy bark of poplar. Foundation item: This paper was supported by National Natural Science Foundation of China (39970620) and the “TRAPOYT” Biography: CHI De-fu (1962-), male, Ph.D., Professor in Northeast Forestry University, Harbin 150040, P.R. China Responsible editor: Chai Ruihai     

Agroforestry in central,northern, and eastern Europe

Integrated management of natural resources and the multiple use of trees and forests have prevailed in most European societies since prehistoric times. In the Middle Ages, expanding and intensified agriculture resulted in the separation of trees from agricultural fields. During the last century, with the introduction of sustainable and highly productive forest management, the goal of increased wood production has been achieved in most parts of central, northern, and eastern Europe. Today, agroforestry is not considered to be an important land-use option within the region; however, there are many practices that could rightfully be classified as agroforestry. These include tree/crop systems in which trees provide products and/or environmental benefits, and tree/animal systems in which animals are grazed in forests or open woodlands.The future seems to offer some prospects for agroforestry. Large areas, hitherto used for food production, are either marginally suited to agriculture, or will probably be taken out of production due to agricultural policy considerations. Agroforestry may, at least in part, offer alternatives for the use of such lands. The availability of (surplus) fertile soils, capital, and labor may provide incentives for site-adapted forms of agroforestry, including improved fallow management. The focus of such systems would be on maintenance of biodiversity in the landscape, environmental protection, recreation, and product diversification.There are numerous expectations as to what agroforestry might provide for the land holder and for society as a whole. These expectations should be carefully analyzed and evaluated prior to political decisions on future land use. The promotion of agroforestry requires overall investment; agroforestry does not happen by itself. A set of integrated actions — not isolated efforts — must be implemented if agroforestry is to become a successful land-use option.