The effect of summer management of perennial ryegrass-dominant swards on plant and animal responses in the autumn when grazed by sheep. 1. Tissue turnover and sward structure

An experiment was conducted lo compare the effects of the grazing by ewes and weaned lambs on aftermath and previously continuously grazed perennial ryegrass-dominant swards, at two sward heights (4 and 8 cm) in (he autumn, on changes in structure and growth of the swards. The experiment had a factorial design, was replicated twice and was conducted from mid-August to early November with measurements being made on three occasions in the autumn. Aftermath swards had lower tiller population densities and lower herbage masses than those that had been previously continuously grazed, the differences being greatest in August. On an area basis net growth rates of herbage on aftermaths were lower than those on previously continuously grazed swards in August and September but not in October. Growth rates of herbage were higher on the taller sward, but the senescence rates were similar at the two sward heights. It was concluded that autumn swards may be managed at taller sward heights than summer swards without increasing senescence of the sward and a consequent reduction in efficiency of utilization. The effects of previous sward management on tissue turnover in the autumn were not long lasting.     

Effects of grazing by sheep or goats on sward composition in ryegrass/white clover pasture and on subsequent performance of weaned lambs

An experiment was conducted to assess the effects of grazing a perennial ryegrass (Lolium perenne) / white clover (Trifolium repens) sward by sheep or goats on sward composition and structure and on subsequent diet selection, herbage intake and liveweight gain by weaned lambs. From mid-May to late July (phase 1), ewes with twin lambs or yearling Scottish Cashmere goats grazed continuously swards maintained at 4- or 8-cm sward surface height. From mid-August to the end of September (phase 2), weaned lambs continuously grazed the same swards maintained at 4 cm (treatment 4–4) or at 8 cm (treatment 8–8) or which had been allowed to increase from 4 cm to 8 cm (treatment 4–8). By the end of phase 1, swards grazed by goats had higher proportions of white clover in the whole sward (0.377 vs. 0.181; s.e.d 0.0382; P < 0.001) than those grazed by sheep, irrespective of sward height treatment. This resulted in phase 2 in a higher proportion of white clover selected ( P <0.001), higher herbage intakes ( P < 0.001) and higher liveweight gains ( P < 0.001) by weaned lambs grazing swards previously grazed by goats compared with those previously grazed by sheep. There were higher proportions of clover present in the swards from treatment 4–8 at the beginning of phase 2 compared with the other sward height treatments and consequently weaned lambs had, on this treatment, a higher proportion of clover in their diet ( P <0.001), higher herbage intakes ( P <0001) and higher liveweight gains ( P <0.001). It is concluded that goats can be integrated into sequential grazing systems with sheep on grass/clover swards and this can result in an increase in the proportion of clover in swards and increased sheep performance.     

Effect of sward surface height on the performance of ewes and Iambs continuously grazed on grass/clover and nitrogen-fertilized grass swards

In three successive years, sward height was maintained at 3, 5, 7 or 9 cm on grass swards receiving a total of 300 kg N ha^?1 in six equal monthly dressings from April, and on grass/clover swards receiving 50 kg N ha^?1 as a single dressing in early spring. From turnout in April until weaning in July, 64 ewes and their lambs (mean litter size 1·5) were continuously grazed at the four sward surface heights on the two sward types. White clover content of grass/clover swards remained low throughout the experiment ranging from 0·2 to 7·4% of the herbage mass. During the first two years, lamb gains averaged over sward types were 204, 260, 285 and 308 g d^?1 up to weaning, while in the third year gains were 238, 296, 296 and 260 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Ewes lost live weight on 3 cm swards but apart from this sward height had little effect on performance. During the autumn, weaned lambs gained — 27, 87, 147 and 167 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Sward type had only a small effect on the performance of lambs up to weaning but in the autumn, mean gains of weaned lambs were lower on grass/N swards (73 g d^?1) than on grass/clover swards (115 g d^?1). Relative to 3 cm swards, carrying capacities of 5, 7 and 9 cm swards were 0·76, 0·57 and 0·52 respectively from turnout to weaning and 0·66, 0·52 and 0·44 respectively during autumn. Grass/clover swards carried 0·67 of the ewes carried by grass/N swards from turnout to weaning and 0·51 of the live weight carried by grass/N swards during autumn. The reaction of the two sward types to sward height did not appear to differ but in the third year there was evidence of a reduction in white clover content when swards were grazed at 9 cm. The data suggest that lamb growth rates will increase as sward height increases up to 9 cm and the evidence for this was stronger with weaned lambs in autumn than with suckling lambs in spring.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

Intake and behaviour responses by sheep to changes in sward characteristics under rotational grazing

A 10-week grazing experiment was conducted on a perennial ryegrass sward with lactating ewes and their twin lambs. Three paddocks were rotationally grazed with rest periods of from 4 to 5 weeks. Sward surface heights at the start of each grazing were 145, 259 and 250mni for treatments RG1, RG2 and RG3. A further four paddocks were maintained by continuous variable stocking (CS) at sward surface heights (SSHs) of about 30, 60, 90 and 120mm. Sward and animal measurements were made on the two different grazing managements as the RG swards were grazed down, giving measurements at similar sward heights for treatments RG and CS.
There was less green leaf and the total herbage mass present under RG was less than on CS swards at the same sward heights, demonstrating the differences in structure between rotationally and continuously grazed swards.
Regression analysis of animal factors on sward factors showed that grazing behaviour was more highly correlated with green leaf mass than SSH or any of the other sward measurements. On the RG swards, maximum intake per animal was reached at about 1500 kg green leaf mass ha⁻¹. A SSH of 60mm allowed the CS ewes to achieve the highest intake rate, but at this height the ewes on treatments RG2 and RG3 were restricted to approximately half this rate. The results suggest that green leaf mass or leaf area index, rather than sward surface height, could be used as a rational basis to relate intake of herbage to sward state for swards changing rapidly in leaf to stem ratio.     

Intake and behaviour responses by sheep to changes in sward characteristics under continuous stocking

Perennial ryegrass pastures were maintained at sward surface heights (SSH) of 30, 60, 90 and 120 mm by continuous variable stocking using lactating ewes and their twin lambs in spring and non-lactating (dry) ewes in autumn.
The effects of SSH on ingestive behaviour, herbage intake, animal performance and sward structure and production were measured in spring, and again in autumn, when an additional SSH of 20 mm was also established.
In spring, differences in sward structure were quickly established with mean tiller numbers of 41 000, 30 000, 21 000 and 19 000 m⁻² and leaf area indices (LAI) of 1·5, 2·2, 3·3 and 4·1 at SSH of 30 to 120, respectively. By autumn the mean tiller numbers had fallen to 26 000, 26 000, 23 000, 18 000 and 13 000 m⁻² and LAI to 1·0, 1·1, 1·5. 2·0 and 1·5 for treatments 20 to 120.
The mean stocking rates for the ewes in spring were 27, 20, 22, and 19 ewes ha⁻¹ and growth rates of their lambs were 208, 275, 250 and 263 g d⁻¹ for treatments 30 to 120. In autumn the stocking rates maintained for dry ewes were 22, 15, 9, 4 and 0 for treatments 20 to 120.
In spring, both grazing times (GT) and prehension biting rate (BR) were negatively correlated with SSH, while bite mass (BM) was positively correlated with SSH. However, mean daily intakes of organic matter (OM), were reduced only for animals grazing the 30 mm sward.
In autumn, intake rate was lower only for the animals grazing the 20mm sward and GT and BR were not significantly affected by SSH.
Relationships between SSH and, components of ingestive behaviour and intake, are presented and the control mechanisms involved are discussed. The results showed that the optimum sward surface height for continuously stocked swards, grazed by sheep, Was between 30 mm and 60 mm.     

The effect of herbage allowance on intake and performance of ewes suckling twin lambs

The effect of herbage allowance on the herbage intake and performance of ewes and their twin lambs at pasture was investigated. Daily herbage allowances of 40,80.120 and 160 g organic matter (OM) kg^-1 ewe live weight, based on herbage mass measured to ground level, were offered during the first 12 weeks of lactation.
The sheep were grazed rotationally around four paddocks of a perennial ryegrass pasture for 7-day periods and herbage mass, extended tiller length and digestibility of the herbage consumed by the animals were estimated. Herbage intake by the ewes was estimated during weeks two to twelve and live weights were recorded weekly: during the last two weeks of the experiment grazing behaviour of one ewe on each treatment was recorded continuously.
Herbage intakes by the ewes (164, 1–81, 2–42 and 268 ± 0153 kg d^-1) and live weight gains of the lambs (202, 245, 274 and 300 ± 7–3 g d^-1) increased with increasing herbage allowances. Herbage intakes by the ewes and growth rates of their lambs increased up to a herbage allowance which was over five times the amount of herbage eaten by the ewes.
As the animals reduced herbage mass and sward height, biting rates by the ewes during grazing increased by 4 (± 0·08) bites min^-1 cm^-1 and masticating rates decreased. Mastications reached a maximum of 90 (± 3·5) min^-1 at a sward surface height of 9 cm.     

Effect of breed and sward height on sheep performance and production per hectare during the spring and autumn in Northern Spain

Abstract Effects of breed and sward surface height on ewe liveweight and body condition changes and on lamb liveweight gains during grazing in the spring and autumn were studied. The output per hectare in each season was also calculated. A total of 112 Gallega ewes (35·6 kg live weight) and 204 Latxa ewes (47·5 kg live weight) with their lambs were used during the spring (March–June) to study the effect of five target sward heights (3·0, 4·5, 5·5, 6·5 and 8·0 cm) and the interaction with breed. In autumn (October–December) 155 Gallega and 126 Latxa ewes were used to study the effect of three target sward heights (4·0, 5·5 and 7·0 cm) and the interaction with breed. Each study was replicated twice. A quadratic relationship was found between sward height and ewe liveweight and body condition changes and lamb liveweight gain during the spring grazing season, with the maximum individual performance being achieved at around a sward height of 6·0 cm. There were significant breed × sward height treatment interactions for ewe liveweight and body condition changes, and lamb liveweight gains. Higher liveweight gains were achieved by Latxa ewes and lambs in swards taller than 6·5 cm but they also sustained higher liveweight losses in swards shorter than 4·5 cm. Nevertheless, a higher output (liveweight gains of ewes + lambs) per hectare for a given weaning date or age was achieved by the Gallega ewes. In autumn, the relationship between liveweight change and sward height was linear, with a breed–sward height interaction similar to that observed in the spring. Sward surface heights of 4·0–4·5 cm and 4·5–5·0 cm were required to maintain live weight and body condition in spring and autumn respectively. Lamb and ewe liveweight gains per hectare decreased considerably in swards taller than 6·5 cm.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 5. The effect of stocking rate and reduced levels of nitrogen fertilizer

The implications for UK upland sheep systems of reducing nitrogen fertilizer application to perennial ryegrass/white clover swards were studied over 3 years. Sward height (3·5–5·5 cm) was controlled for ewes with lambs until weaning using surplus pasture areas for silage; thereafter, ewes and weaned lambs were grazed on separate areas, and sward height was controlled by adjusting the size of the areas grazed and using surplus pasture areas for silage if necessary. Combinations from three stocking rates [10, 6 and 4 ewes ha⁻¹ on the total area (grazed and ensiled)] and four nitrogen fertilizer levels (150, 100, 50 and 0 kg ha⁻¹) provided six treatments that were replicated three times. Average white clover content was negatively correlated with level of nitrogen fertilizer. The proportion of white clover in the swards increased over the duration of the experiment. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain on all treatments. All treatments provided adequate winter fodder as silage. It is concluded that the application of nitrogen fertilizer can be reduced or removed from upland sheep pastures without compromising individual animal performance provided that white clover content and sward height are maintained. Resting pastures from grazing by changing ensiled and grazed areas from year to year sustained white clover content over a 3-year period.     

The effect of daily herbage allowance and sward characteristics upon the ingestive behaviour and herbage intake of calves under strip-grazing management

Measurements of herbage intake and ingestive behaviour were made on British Friesian calves 4–9 months old, which were strip-grazed on swards of S24 perennial ryegrass at three (Experiment A) or four (Experiment B) levels of daily herbage allowance in three (A) or four (B) 12-d periods. Herbage intake declined by approx. 18% as daily herbage allowance was reduced from 90 to 30 g DM per kg LW in both experiments, but the relationship between intake and allowance was not consistent between periods. Bite size and rate of biting were greater in calves on low than on high allowances after entry to a new strip of herbage but the positions were reversed towards the end of grazing. Reduced herbage intakes at low allowance resulted from reductions of approx. equal magnitude in bite size, rate of biting and grazing time, though not all of these effects were significant. It is suggested that these responses result from the increasing difficulty of prehending and ingesting herbage as swards are grazed down. The calves at low herbage allowances did not increase grazing time, probably because their behaviour was conditioned by anticipation of a new allowance of herbage. Bite size and daily herbage intake were greater on a spring sward than on an autumn sward, but within experiments herbage intake was not consistently related to any of the sward characteristics measured.     

Animal production evaluation of herbage varieties. 1. Comparison of Aurora with Frances, Talbot and Melle perennial ryegrasses when grown alone and with clover

Comparative herbage characteristics and sheep production (ewes and lambs until weaning and lambs thereafter) from swards of Aurora (very early flowering), Frances (early flowering), Talbot (intermediate flowering) and Melle (late flowering) varieties of perennial ryegrass ( Lolium perenne ) were studied under a continuous variable stocking management based on sward surface height guidelines. The varieties were assessed as grass-only (215 kg N ha⁻¹ fertilizer) and grass/clover (83 kg N ha⁻¹ fertilizer) pastures.
Over 2 years (1985-86) total annual lamb production per hectare from grass-only swards of Aurora was 19% more than that from Frances despite similar herbage productivity. Differences between the varieties in lamb output were more pronounced as grass/clover swards, with Aurora producing 29 and 18% more lamb than Melle and Talbot respectively and with Frances also giving 16% more than Melle. Overall lamb production from grass/clover swards was 10% more than that from the grass-only pastures, mainly due to 69% advantage in individual lamb growth rates after weaning. Herbage organic matter digestibility, during the post-weaning period, was higher on grass/clover than on grass-only swards, and similarly with Aurora versus the other varieties. Under a frequent cut simulated grazing regime the relative herbage productivity of the four varieties differed, with Aurora 13% less productive than Frances.
The results are discussed in relation to the limitations of assessing performance under cutting, and the significance of the interaction in animal productivity between the two sward types.     

Effect of early season sward management on sward quality and lamb liveweight gain during autumn

In two experiments (1990 and 1991) perennial ryegrass (cv. Talbot) pasture was subjected to six different management treatments by adjusting stock numbers to achieve set sward heights for set periods between turn-out in early April and weaning in early July. The main objective of the experiments was to measure the effect of differences in sward management up to July on subsequent sward type, and the effect of sward type on the performance of weaned lambs between mid-July and early October when all swards were grazed at a sward height of 7 cm. In Experiment 1 swards were grazed at 7 cm initially and then reduced to 3 cm at various dates and maintained at 3 cm until early July. In Experiment 2 swards were grazed initially at 7 cm, and this was reduced to 3 cm and then allowed to return to 7 cm at various dates up to early July. Grazing at a sward height of 7 cm up to weaning, in early July, gave mean twin lamb growth rates of 310 g d^?1, while at a tower sward surface height of 3 cm lamb growth rate was reduced to 206 g d^?1 The density of seed heads produced from July was strongly influenced by earlier management. Grazing at 7 cm up to July gave a high density of seed heads (213m^?2), and this reduced (126 m^?2) by grazing to 3 cm. Seed heads were almost eliminated (17 m^?2) by grazing at 7 cm until late June followed by grazing down to 3 cm in early July. Reducing sward surface height from 7 cm to 3 cm at an earlier date tended to increase the development of seed heads from early July. Where swards were grazed from 7 cm to 3 cm and then allowed to return to 7 cm, the later in the pre-weaning period this took place the greater the reduction in seed head development. Greater seed head density was associated with a significant decrease in the nitrogen content of the herbage on offer in Experiment 2, but an increase in modified acid detergent (MAD) fibre content was not significant. Despite the differences in the sward type on offer, the effect on weaned lamb growth rate was small. Lamb liveweight gain during autumn was significantly reduced in Experiment 2 following earlier grazing at 7 cm compared with 3 cm, but liveweight gain differences in Experiment 1 were not significant.     

Variations in the surface characteristics of the sward and the short-term rate of herbage intake by calves and lambs

The results are reported of an examination of detailed measurements on the ingestive behaviour of calves and lambs in response to variations in the surface characteristics of swards under strip-grazing and continuous stocking managements. It is shown that intake per bite and the short-term rate of herbage intake were both sensitive to the height of the surface horizon above ground level but, at least under strip-grazing management, not to variations in herbage density in the grazed horizon. The rate of biting was less sensitive to variations in sward conditions, particularly under strip-grazing.
Both intake per bite and rate of intake were more sensitive to variations in grazing height under strip-grazing, where sward changes were rapid, than under continuous stocking, where they were slow. Under continuous stocking, ingestive behaviour was more sensitive to changes in sward conditions in lambs than in calves.     

The effect of a high-magnesium perennial ryegrass variety on the magnesium status of sheep

An early-heading variety of perennial ryegrass (Ramore) bred for high magnesium (Mg) content was compared with an early control variety (Frances) for its ability to increase Mg intake, Mg utilization and blood Mg concentration in sheep, and so reduce the incidence of hypomagnesaemic tetany. Swards of the two varieties were established and each was grazed by ewes suckling twin lambs from turn-out to weaning and thereafter by dry ewes. In early and late season herbage from a proportion of each sward was cut daily and fed fresh to ewes housed indoors in metabolism crates. Under grazing conditions herbage from Ramore swards had, over the 1992 and 1993 grazing seasons, a higher Mg concentration than Frances (2.35 g kg^?1 vs. 2.18 g kg^?1; s.e.m. 0.019, P< 0.001). During the spring period there was a higher (P 0.001, s.e.m. 0.0074) Mg concentration in the blood of ewes grazing Ramore (0.854 m mol 1^?l) swards compared with those grazing Frances (0.793 m mol 1 ^?1) whereas during the autumn period blood Mg concentrations were higher in ewes grazing Frances. Ewe and lamb liveweight gains, recorded in 1992 only, were higher on Ramore swards than on Frances swards, although the difference was not significant. Indoor feeding studies showed that Mg intake was significantly higher (P< 0.001, s.e.m. 0.026) in ewes offered Ramore (1.98 g d^?1) herbage compared with those offered Frances (1.74 g d^?1) herbage with no significant difference in dry-matter intake between the two varieties. Apparent availability and retention of Mg were higher for animals fed Ramore during the spring feeding periods than those fed Frances, but during the autumn feeding periods the reverse was true. Under indoor feeding conditions, variety had no significant effect on blood Mg concentrations. It was concluded that under grazing conditions a high Mg variety could provide an effective means of reducing the incidence of hypomagnesaemic tetany.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 2. Animal production

Performance of continuously stocked Mule ewes nursing Suffolk-cross twin lambs over three grazing seasons, between April and August, was compared on swards of N-fertilized diploid perennial ryegrass (D), tetraploid perennial ryegrass (T) and tetraploid perennial ryegrass with white clover (TC), the latter receiving no fertilizer N. Sward height was maintained by variable stocking rate close to a target of 4–6 cm (constant treatment) from turnout and compared in July and August with a rising sward height treatment (target 6–8 cm). Lambs on TC swards had significantly higher (P <0·001) liveweight gains compared with lambs on T swards by 41 gd^-1 in April–June and by 68gd^-1 in July-August. Live weight and body condition score of ewes in August were significantly higher (P<0·001) on TC compared with T swards, by 11·3 kg and 0·75 respectively. Rising sward heights in July–August increased live-weight gain of lambs compared with constant sward heights by 102, 39 and 54gd^-1 in consecutive years, associated with sward height increases of 0·9, 0·5 and 0·6cm respectively. Rising sward height increased ewe live weight and body condition score by 5·1 kg and 0·3 respectively, compared with results from constant sward heights. Effects of sward height and sward type were additive. T swards had a significantly (P<0·01) 16% greater overall lamb output than the D swards due mainly to a 10% higher achieved stocking rate. Stocking rates of ewes on TC vs T swards were 40, 13 and 12% lower in April-August in successive years. The higher liveweight gain of lambs on the TC swards resulted in lamb outputs of 76, 105 and 101% of the T swards in successive years, showing that grass/clover swards containing over 20% clover could produce similar lamb output ha^-1 to grass swards given 150–180 kg N ha^-1.     

Variations in herbage mass and digestibility, and the grazing behaviour and herbage intake of adult cattle and weaned calves

Two experiments are described in which groups of lactating cows, non-lactating cattle and weaned calves grazed a sequence of swards varying in maturity and herbage mass under strip grazing management at a daily herbage allowance of 60 g dry matter per kg live weight. Lactating cows ate 43% and 76% more herbage than non-lactating cattle of similar weight in the two experiments but herbage OM intakes per unit live weight by the calves and lactating cows were similar.
Variations in diet digestibility and herbage intake in the lactating and non-lactating cattle with changes in sward conditions were similar. In the first experiment the calves were experienced grazers; the variation in the digestibility of the herbage selected was less in the calves than the adult cattle, but the variation in herbage intake was greater. The calves in the second experiment were younger, and they were inexperienced grazers; their ability to increase herbage intake in response to changing sward conditions was poorer than that of adult cattle, though variations in diet digestibility were similar in all classes.
The practical implications of these results are discussed briefly.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK: 6. The effect of reducing stocking rate and application of fertilizer nitrogen in Wales

Abstract The implications for the agricultural productivity of the UK upland sheep systems of reducing nitrogen fertilizer application and lowering stocking rates on perennial ryegrass/white clover swards were studied over 4 years at a site in Wales. The system involved grazing ewes and lambs from birth to weaning on swards maintained at a constant height with surplus herbage made into silage, thereafter ewes and weaned lambs grazed on separate areas until the onset of winter with adjustments to the size of the areas grazed and utilizing surplus pasture areas for silage. Four stocking rates [SR 18, 15, 12 and 9 ewes ha^?1 on the total area (grazed and ensiled)] and two levels of annual nitrogen fertilizer application (N 200 and 50 kg ha^?1) were studied in five treatments (N200/SR18, N200/SR15, N50/SR15, N50/SR12 and N50/SR9). Average white clover content was negatively correlated with the level of annual nitrogen fertilizer application. White clover content of the swards was maintained over the duration of the experiment with an increasing proportion of clover in the swards receiving 50 kg N ha^?1. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain from birth to weaning in all treatments but fewer lambs reached the slaughter live weight by September at the higher stocking rates and with the lower level of fertilizer application. Three of the five treatments provided adequate winter fodder as silage (N200/SR15, N50/SR12 and N50/SR9). Because of the failure to make adequate winter fodder and the failure of white clover to fully compensate for reduction in nitrogen fertilizer application, it is concluded that nitrogen fertilizer can only be reduced on upland sheep pastures if accompanied by reduced stocking rates.     

Herbage intake and liveweight gain of bulls and steers continuously stocked at fixed sward heights during autumn and spring

Two experiments were conducted to examine the relationship between sward surface height, herbage intake and liveweight gain in beef cattle grazed on pasture. In Experiment 1, two 'animal types' (18 Charolais × Angus steers and 18 Friesian bulls) were continuously grazed for 22 days during the late autumn on replicated swards maintained at sward surface heights of 6, 10 and 15 cm. Herbage intakes, assessed from the faecal concentration of chromium delivered from an intraruminal controlled release capsule and the in vitro digestibility of hand-plucked herbage samples, were curvilinearly related to sward height (r = 0·76, p <0·0·01). Average liveweight gains were 0·02, 0·61 and 1·31 kg d^-1 ( P <0·05) and increased linearly ( r = 0·84, P <0·001) with sward height. The maintenance organic matter intakes of the steers and bulls, with initial mean (± s.e.) live weights of 225 ± 15 kg and 172 ± 15 kg respectively, were estimated to be 3·6 and 3·5 kg d^-1 respectively. In Experiment 2 (spring) 36 cattle, including 35 of those used in Experiment 1, were reallocated to sward heights of 5, 10 and 15 cm using the same design as for Experiment 1. Average liveweight gains were 0·94, 1·57 and 1·68kg d^-1 ( p 0·05) and were curvilinearly related to sward height ( r = 0·093, p <0·05). Maintenance intakes could not be reliably extimated for the cattle in Experiment 2 because few animals had liveweight gains close to zero. These trials confirm that liveweight gain in continuously grazed finishing steers and bulls increases with sward surface height to maximum of 8–10cm with spring ryegrass/white clover pastures while, in autumn, swards of 12–15cm height are required to achieve maximum performance.     

Influence of structure and composition of ryegrass and prairie grass-white clover swards on the grazed horizon and diet harvested by sheep

Diet selection from ryegass-and prairie grass-white clover swards, vertically stratified into three horizons (A > 6 cm, B 3–6 cm, C > 3 cm), was studied using oesophageally fistulated sheep during summer and autumn. Animals grazed for 3-day periods. Apparent herbage intake was calculated from total herbage disappearance. The composition of each horizon and of the diet selected was measured daily.
Herbage mass (DM ha^-1) and sward height (cm) prior to grazing were not significantly different between swards in each season, and were 2·0 and 20 in summer and 1·6 and 10 in autumn. In summer, 36% and 5% of the green grass leaf (GGL) for prairie grass and ryegrass, respectively, was distributed in horizons A and B. In autumn 39% and 29% of GGL occurred above 3 cm for prairie grass and ryegrass, respectively. GGL distribution determined which sward horizons were grazed. Sheep grazed horizon C (0–3 cm) of summer ryegrass pasture, and the surface canopy (>3 cm) of all other swards.
In summer, apparent intake achieved by sheep grazing prairie grass swards was 87% higher than that achieved on ryegrass swards. In autumn a greater GGL distribution above 3 cm with prairie     

The effects of sward characteristics and supplement type on the herbage intake and milk production of summer-calving cows

High (H; 27 350 m^?1:) or low (L. 13 300 m-¹) tiller density perennial ryegrass swards were created in the mid- and late grazing season by imposing different sward heights in the spring. Summer-calving cows then grazed these swards from 6 June to 2 September 1992 and were offered 5 kg fresh weight hd^?1 d^?1 of either a barley (S) or a molassed sugar beet pulp (F) based supplement. The factorial combination of sward and supplement types resulted in four experimental swards being grazed by thirteen Holstein/Friesian cows each. Supplement F contained more crude fibre (110 vs. 58 g kg^?1) and less metabolizable energy [12–5 vs. 13–2 MJ kg^?1 dry matter (DM)] than supplement S. Herbage on the H sward contained more metabolizable energy (11–9 vs. 104 MJ kg^?1 DM) and crude protein (232 vs. 205 g kg^?1 DM), had fewer rejected areas f 16–5 vs. 26–9%) and a higher live-dead tiller ratio (4–6 vs. 2–1) than that on the L sward. Sward, but not supplement type, significantly affected the intake of grazed herbage (P<0–001). On average, the herbage intakes of cows grazing II swards were higher than for L swards (14–5 vs. 11 6kg DM d^?1) and those of cows on the S and F supplements were 12–6 and 13–5 kg DM d^?1 respectively. Averaged over the grazing period, sward and supplement had no significant effects on milk yield, milk composition or yield of constituents. When expressed on an average weekly basis, cows grazing an L sward and offered the F supplement on occasions had significantly lower milk yields and higher milk fat contents (P<005) than those grazing an H sward and offered the S supplement. There were no significant effects on cow live weight or condition score change. The results suggested that grazing swards with a high density of live tillers increased herbage intakes and on occasions milk yield, relative to low density swards. However, small increases in energy intake from sward and supplement effects were used primarily to ameliorate liveweight loss.