Improving the Traditional Acacia Senegal-Crop System in Sudan: The Effect of Tree Density on Water Use, Gum Production and Crop Yields

The traditional Acacia senegal bush-fallow in North Kordofan, Sudan, was disrupted and the traditional rotational fallow cultivation cycle has been shortened or completely abandoned, causing decline in soil fertility and crop and gum yields. An agroforestry system may give reasonable crop and gum yields, and be more appealing to farmers. We studied the effect of tree density (266 or 433 trees ha⁻¹) on two traditional crops; sorghum (Sorghum bicolor) early maturing variety and karkadeh (Hibiscus sabdariffa), with regard to physiological interactions, yields and soil water depletion. There was little evidence of complementarity of resource sharing between trees and crops, since both trees and field crops competed for soil water from the same depth. Intercropping significantly affected the soil water status, photosynthesis and stomatal conductance in trees and crops. Gum production per unit area increased when sorghum was intercropped with trees in low or high density. However, karkadeh reduced the gum yield significantly at high tree density. Yields of sorghum and karkadeh planted within trees of high density diminished by 44 and 55% compared to sole crops, respectively. Intercropping increased the rain use efficiency significantly compared to trees and field crops grown solely. Karkadeh appears to be more appropriate for intercropping with A. senegal than sorghum and particularly recommendable in combination with low tree density. Modification of tree density can be used as a management tool to mitigate competitive interaction in the intercropping system.     

Wind protection in a hedged agroforestry system in semiarid Kenya

In semiarid Laikipia (Kenya) severe crop damage and loss of mulch material may be caused by south to south-easterly winds from June to September. Demonstration agroforestry systems which surround farms with live fences had some success in protecting crops, mulch and soil, but great care must be taken, because air may be channelled through or over them. For demonstration purposes, a deliberate gap was made in a two meter high Coleus barbatus live fence to study its effect on wind speed and damage to crops. The effectiveness of protection given by this hedge together with intercropped Grevillea robusta trees was quantified using electrical cup anemometers. The combination of hedges and trees gave protection to a maize/bean intercrop but the biomass distribution was not the most suitable one. This picture was complicated by variable wind direction and interactions between wind and the biomass of hedges and trees. The lowest efficiency of wind reduction occurred closest to the deliberate gap, where also the lowest protection by the Grevillea trees was found. South to south-easterly winds increased the gap effect and caused gradients in tree protection perpendicular to the southern hedge. This gave at times wind speeds even higher than outside the system. Only visually wind effects could be detected. Trees and hedges strongly competed with the crops if not root pruned.This revised version was published online in November 2005 with corrections to the Cover Date.     

Productivity and nutrient cycling in young Acacia senegal farming systems on Vertisol in the Blue Nile region, Sudan

Acacia senegal, the gum arabic-producing tree, is the most important component of traditional dryland agroforestry systems in the␣Sudan. The spatial arrangement of trees and the type of agricultural crop used influence the interaction between trees and crops. Tree and crop growth, gum and crop yields and nutrient cycling were investigated over a period of 4 years. Trees were grown at 5 × 5 m and 10 × 10 m spacing alone or in mixtures with sorghum or sesame. No statistically significant differences in sorghum or sesame yields between the intercropping and control treatments were observed (mean values were 1.54 and 1.54 t ha⁻¹ for sorghum grain and 0.36 and 0.42 t ha⁻¹ for sesame seed in the mixed and mono-crop plots, respectively). At an early stage of agroforestry system management, A. senegal had no detrimental effect on crop yield; however, the pattern of resource capture by trees and crops may change as the system matures. A significant positive relationship existed between the second gum picking and the total gum yield. The second gum picking seems to be a decisive factor in gum production and could be used as an indicator for the prediction of the total gum yield. Soil organic carbon, N, P and K contents were not increased by agroforestry as compared to the initial levels. Soil OC was not increased by agroforestry as compared to sole cropping. There was no evidence that P increased in the topsoil as the agroforestry plantations aged. At a stocking density of 400 trees ha⁻¹ (5 × 5 m spacing), A. senegal accumulated in its biomass a total of 18.0, 1.21, 7.8 and 972 kg ha⁻¹ of N, P, K and OC, respectively. Agroforestry contributed ca. 217 and 1500 kg ha⁻¹ of K and OC, respectively, to the top 25-cm of soil during the first four years of intercropping.     

Effect of increasing distance fromAcacia nilotica trees on wheat yield

Planting trees in the cultivated fields of Pakistan is a controversial topic between foresters and agriculturists. Some farmers believe that trees significantly reduce their crop yields; therefore they avoid planting trees in their fields. On the other hand, many foresters claim trees are beneficial for crops. This study was conducted to determine the influence of boundary trees (Acacia nilotica) on the growth and yield of associated wheat crops under irrigated conditions in Punjab, Pakistan. Results indicate that close proximity to trees adversely affected tillers/m², weight/1000 grain and the grain yield of wheat planted up to a distance of 8.5 m from the trees. Plant height and grains/spike were least affected. In general, the growth and yield of wheat improved as distance from the tree increased. Tree size did not affect wheat height, tillers/m², grains/spike or weight/1000 grain. The grain yields were slightly lower near the largest trees (50–54.9 cm DBH trees).     

The effect of young Leucaena leucocephala (Lam) De Wit hedges on the growth and yield of maize,sweet potato and cowpea in an agroforestry system in Sierra Leone

An investigation was undertaken at Senehun — Kamajei, a high rainfall region in Sierra Leone, to assess the effects of Leucaena leucocephala, on the growth and productivity of maize, cowpea and sweet potato. The experiment was laid out in a completely randomised design with four replicates of each treatment. The treatments and controls were: (i) pure crops of maize, cowpea, sweet potato, Leucaena clean weeded and unweeded and (ii) intercrops of Leucaena with the food crops, both (iii) with and without applied fertilizers after the first year.The growth of Leucaena was slow but nevertheless tended to reduce grain yields of maize, and tuber and vine yields of sweet potato in the rows in the immediate vicinity of the trees, especially in the nitrogen-treated plots. The maize, in particular, caused an improvement in the early height growth of the trees.     

Agroforestry systems with Ilex paraguariensis (American holly or yerba mate) and native timber trees on small farms in Misiones, Argentina

In Misiones, in the northeast of Argentina, agroforestry systems of timber trees and perennial cash crops are becoming increasingly common. We evaluated the productivity of Ilex paraguariensis St Hill (South American holly or yerba mate, Aquifoliaceae) in association with indigenous trees: Enterolobium contortisiliquum (Vellozo) Morong. (timbó, Leguminosae, a N₂-fixing tree), and two timber species, Balfourodendron riedelianum (Engler) Engler (guatambú, Rutaceae) and Tabebuia heptaphylla (Vellozo) Toledo (lapacho negro, Bignoniaceae). Five years after planting, the tree species were 2.7 to 5.0 m high and 3 to 7 cm in diameter at breast height, and the yerba mate produced its first harvest. Additionally, the production of associated crops of subsistence covered the annual needs of the farmer. These systems are promising for sustainable use of deforested lands in the region.This revised version was published online in November 2005 with corrections to the Cover Date.     

Benefits and costs of intercropping reforestation in the Atlantic lowlands of Costa Rica.

Agroforestry options as a means of promoting reforestation were testedby establishing with 10 farmers simple comparisons between pure plantations andtimber trees with annual or perennial intercrops. Two year-old intercroppedtrees on-farm had 22–48% greater height and 24–38% greater diameterthan trees in pure plantations. The exception was Cordiaalliodora that did not respond significantly to intercropping withcassava (Manihot esculenta). Only intercrops with the mostvaluable crops on agricultural soils produced a return to labour above thedailywage rate. On non-agricultural soils, farmers only intercrop a small area oftheir plantations with subsistence crops to meet home consumption needs. In thecontext of Costa Rica intercropping appeared to have a limited potential tofinance the reforestation costs of farmers.     

Eucalyptus plantations in Israel: an assessment of economic and environmental viability

This paper discusses a plantation management approach involving a combination of “short” and “long” rotations designed to allow farmers to receive income from trees as soon as possible after establishment. We present results from two plots that represent extreme conditions: (a) a seasonally waterlogged, non-saline site (Nahalal), and (b) a saline site (Ginnegar) located in the Yizre’el Valley, Israel. Six improved seed sources, four of Eucalyptus camaldulensis and two of E. occidentalis, were examined. The local Israeli seed source of E. camaldulensis (HA) performed best at both sites. In Nahalal, the short rotation thinning of the slower growing (50%) plantation trees could provide economic returns approximately five years after establishment. The calculated mean annual increment (MAI) of these trees reached 12.2 t ha⁻¹ year⁻¹. The long rotation, or better performing half of the plantation trees, could be used as a source of sawn timber, providing higher-value products. By nine years after establishment, the average DBH of the various seed sources reached 25.8 ± 1.9 cm. The calculated MAI of the combined cutting rotations reached 48.3 t ha⁻¹ year⁻¹. Eucalyptus grown under the combined (short- and long-term) management approach at Nahalal was more profitable than many other non-irrigated local crops. Eucalyptus production in Ginnegar would be less profitable than in Nahalal. However, an additional ecological benefit was provided by the crop’s ability to lower the water table. When this contribution to regional drainage is taken into account, trees become economically competitive with other non-irrigated field crops under saline conditions. Jim Morris–Deceased.     

Soil water dynamics in cropping systems containing <Emphasis Type="Italic">Gliricidia sepium</Emphasis>, pigeonpea and maize in southern Malawi

The water dynamics of cropping systems containing mixtures of Gliricidia sepium (Jacq.) Walp trees with maize (Zea mays L.) and/or pigeonpea (Cajanus cajan L.) were examined during three consecutive cropping seasons. The trees were pruned before and during each cropping season, but were left unpruned after harvesting the maize; prunings were returned to the cropping area in all agroforestry systems to provide green leaf manure. The hypothesis was that regular severe pruning of the trees would minimise competition with crops for soil moisture and enhance their growth by providing additional nutrients. Neutron probe measurements were used to determine spatial and temporal changes in soil moisture content during the 1997/98, 1998/99 and 1999/00 cropping seasons for various cropping systems. These included gliricidia intercropped with maize, with and without pigeonpea, a maize + pigeonpea intercrop, sole maize, sole pigeonpea and sole gliricidia. Soil water content was measured to a depth of 150 cm in all treatments at 4–6 week intervals during the main cropping season and less frequently at other times. Competition for water was apparently not a critical factor in determining crop performance as rainfall exceeded potential evaporation during the cropping season in all years. The distribution of water in the soil profile was generally comparable in all cropping systems, implying there was no spatial complementarity in water abstraction by tree and crop roots. However, available soil water content at the beginning of the cropping season was generally lower in the tree-based systems, suggesting that the trees continued to deplete available soil water during the dry season. The results show that, under rainfall conditions typical of southern Malawi, the soil profile contains sufficient stored water during the dry season (ca. 75–125 mm) to support the growth of gliricidia and pigeonpea, and that gliricidia trees pruned before and during the cropping season did not deleteriously compete for water with associated crops. Water use efficiency also appeared to be higher in the tree-based systems than in the sole maize and maize + pigeonpea treatments, subject to the proviso that the calculations were based on changes in soil water content rather than absolute measurements of water uptake by the trees and crops.     

Karité (Vitellaria paradoxa) and néré (Parkia biglobosa) associated with crops in South Mali

The effect of two tree species, karité (Vitellaria paradoxa) and néré (Parkia biglobosa) on soil condition, development and production of cotton, sorghum and pearl millet, main crops in South Mali, has been investigated.The soil under the trees is slightly richer (organic matter content and several cations) compared to adjacent tree-less sites. Also tree-specific effects exist.Soil enrichment, however, is mainly a matter of redistribution of locally available nutrient resources.Of the six associations studied, only karité-cotton shows no tree-induced reduction in crop production. All other associations suffer greatly from reduced crop outputs caused by the trees, usually in the order of 60%.There are at least two reasons that explain tree-induced yield reduction. Because of increased humidity in the immediate surroundings of trees, in both soil and air, lower numbers of crop plants survive up to maturity, presumably because plants are attacked by fungi. Secondly, crop plants that mature show reduced output because of inter-specific competition for light and nutrients.In case of néré, in order to minimize the trees' negative influence lopping is advised. Such practice, however, is less suitable in case of karité, because of its assumed lower potential to regenerate, as compared to néré.     

Acacia albida and other multipurpose trees on the fur farmlands in the Jebel Marra highlands,Western Darfur,Sudan

This paper describes the traditional agroforestry systems based on Acacia albida and other multipurpose trees as practised by the sedentary Fur people on the lower slopes and highlands of the Jebel Marra massif, Sudan. The basic agrosilvopastoral system consists of terraced village fields, where semipermanent rainfed cropping of staple millet and other subsistence crops takes place under stands of multipurpose trees dominated by Acacia albida, Cordia abyssinica and Ziziphus spina-christi. Trees have been retained primarily for food, wood and fodder. Thorn from cut and browsed branches makes a good fencing materal.This system has been able to sustain self-sufficiency of a densely settled population over centuries. However, recent out-migration of people from the montane dry-farming areas has caused gradual return to shifting cultivation. As a consequence, the present-day subsistence farming in the region is characterized by a general level of carelessness and exploitative management and this is reflected in a successive decrease of the tree cover.The evaluation of the AF practices described includes a discussion on their regional importance and extrapolability within the framework of similar situations, especially in Africa, emphasis being given to mountain and highland conditions. The outstanding potential for Acacia albida-based AF systems to be sustained and spread almost all over semiarid to semihumid Africa is highlighted by illustrating its ecologic and economic variability. Extrapolation of such examples, however, is not feasible, without thorough feasibility studies concerning the ecologic, ethnologic and socio- economic conditions in the respective project areas. Basic research needs for improving and extending the system are also indicated.     

Beneficial Effects of Intercropping on the Growth and Nitrogen Status of Young Wild Cherry and Hybrid Walnut Trees

Growing concern for economic and environmental issues emphasizes the potential value of intercropping systems in temperate regions. However, the selection of relevant tree species to be associated with crops has been little documented. The growth and the nitrogen nutrition of two economically valuable species, wild cherry (Prunus avium L.) and hybrid walnut (Juglans nigra L.×Juglans regia L.), were compared over six years after plantation. These two species were associated with non-irrigated cereal crops in the agroforestry treatment or grown separately (weeded control and fallow). Intercropping increased diameter growth as soon as year 2 in the two species. Leaf biomass assessment using allometric models showed an earlier and greater leaf biomass increase in hybrid walnut than in wild cherry tree. After six years, the relative growth increase of the agroforestry trees with respect to the control trees varied with the parameter considered (diameter at breast height from +26 to +65%, leaf biomass from +54 to +142%) and with the tree species (higher relative growth for hybrid walnut trees). The beneficial effect on tree growth can be accounted for in terms of enhanced nitrogen nutrition. The tree–crop association in intercropping systems, which improves tree growth, might thus allow the planting of more demanding trees on soils of lower fertility.     

An evaluation of the Acacia albida-based agroforestry practices in the Hararghe highlands of Eastern Ethiopia

Growing Acacia albida as a permanent tree crop, on farmlands with cereals, vegetables and coffee underneath or in between, is an indigenous agroforestry system in the Hararghe highlands of Eastern Ethiopia. However, there is practically no systematic record or data on the merits and benefits of this practice.The paper presents the results of an investigation into the effects of the presence of A. albida on farmlands on the yield of maize (Zea mays L.) and sorghum (Sorghum bicolor L. Moench). Twenty seven plot pairs each consisting of one plot underneath the A. albida foliage cover and the other in the open, away from the tree-on farmers' fields, in a 40 km radius around the Alemaya College of Agriculture, were sampled and the yield components analyzed. A statistically significant increase in crops yields by 56% on average was found for the crops under the tree canopies compared to those away from the trees. This increase was caused by the improvement in 1000 grain weight and number of grains of plants under the tree, indicating that the trees enhanced the fertility status of the soil and improved its physical conditions in terms of crop growth.Additional benefits from the A. albida trees include supply of fuelwood and fodder. Quantitative estimates of these outputs as well as their monetary values are presented in the paper. However, in order to realize these benefits to a discernible extent, higher stand densities of the tree than at present are required.Based on an enquiry about the farmers attitude towards A. albida, the prospects for an extension of this promising agroforestry technique are discussed against the background of the state and trends of development of agriculture in the area. It is surmised that despite some shortcomings like the relatively slow and highly variable growth of A. albida and a conflict with the spreading cultivation of Ch'at (Catha edulis Forsk.), the prospects of extension of this technique are good. It is recommended that its propagation should be incorporated into the programmes of the extension agencies of the various governmental agencies concerned with land use.     

Water use efficiency and uptake patterns in a runoff agroforestry system in an arid environment

Water is the most limiting factor for plant production in arid to semiarid regions. In order to overcome this limitation surface runoff water can be used to supplement seasonal rainfall. During 1996 we conducted a runoff irrigated agroforestry field trial in the Turkana district of Northern Kenya. The effects of two different Acacia saligna (Labill.) H. Wendl. tree planting densities (2500 and 833 trees per ha), tree pruning (no pruning vs. pruning) and annual intercrops (no intercrop vs. intercrop: Sorghum bicolor (L.) Moench during the first season and Vigna unguiculata (L.) Walp. during the second season) on water use were investigated. The annual crops were also grown as monocrops. Water consumption ranged from 585 to 840 mm during the first season (only treatments including trees). During the second season, which was shorter and the plants relied solely on stored water in the soil profile, water consumption was less than half of that during the first season. Highest water consumptions were found for non-pruned trees at high density and the lowest were found for the annual crops grown as monocrops. Tree pruning decreased water uptake compared to non-pruned trees but soil moisture depletion pattern showed complementarity in water uptake between pruned trees and annual intercrops. The highest values of water use efficiency for an individual treatment were achieved when the pruned trees at high density were intercropped with sorghum (1.59 kg m^–3) and cowpea (1.21 kg m^–3). Intercropping and high tree density increased water use efficiency in our runoff agroforestry trial. We ascribe the observed improvement in water use efficiency to the reduction of unproductive water loss from the bare soil.This revised version was published online in November 2005 with corrections to the Cover Date.     

Spice crops agroforestry systems in the East Usambara Mountains,Tanzania: growth analysis

A scarcity of cultivation land calls for more intensive and productive land use in the East Usambara Mountains in NE Tanzania. Spice crops could generate cash in higher parts of the mountains, but the present cultivation methods are depleting the valuable forest resources. The trial was established at the end of 2000 to find out how the two popular cash crops, cardamom (Elettaria cardamomum (L.) Maton.) and black pepper (Piper nigrum L.), normally grown under the natural forest, will produce in intensive agroforestry system with two multipurpose farm trees, Grevillea robusta A.Cunn. and nitrogen fixing Gliricidia sepium Jacq. Results from 6 years showed that cardamom produced better with grevillea than in natural forest; 5.5 times more in the fourth year than the average in the area. The Land Equivalent Ratios for black pepper and cardamom showed that pepper intercropped with grevillea produced 3.9 times more than in monoculture whereas cardamom intercropped with grevillea and pepper produced 2.3 times more than in monoculture. Gliricidia improved the nitrogen and organic matter content of the soil over the levels found in natural forest. Soil acidity was, however, preventing the plants from using the available mineral nutrients more effectively.     

Tree/crop complementarity in an arid zone runoff agroforestry system in northern Kenya

We tested the hypothesis that shallow-rooted crops and deep-rooted trees will share the available water in a complementary manner, when grown together, in a field trail in the Turkana district of northern Kenya during 1994 to 1996. Such studies have been few in dryland agroforestry. The effects of two different Acacia saligna (Labill.) H. Wendl. tree planting densities (2500 and 833 trees per ha), tree pruning (no pruning vs. pruning) and annual intercrops (no intercrop vs. intercrop) on total biomass production and their interactions were tested. In 1996 Sorghum bicolor (L.) Moench was used during the first vegetation period and Vigna unguiculata (L.) Walp. during the second. We used naturally generated runoff water for irrigation to supplement low rainfall amounts typical for the area. High biomass production (> 13 t ha^–1 over a two year period) was observed irrespective of intercropping of pruned trees or sole tree stands. Although the pruning treatment reduced total tree biomass yields by a quarter, the introduction of annual intercrops after the pruning of trees outweighed this loss. The yields of the intercrops in the pruned tree treatments were similar to their yields when grown as monocrops. The calculation of land equivalent ratios showed overyielding for intercropped, pruned systems. The high values for LER (1.36 at low and 1.47 at high density of trees) indicate that there is complementarity in resource use between the different species.This revised version was published online in November 2005 with corrections to the Cover Date.     

Lacquer-based agroforestry system in western Yunnan, China

The lacquer tree (Toxicodendron vernicifiuum) based agroforestry system is a very important farming system with development potential in western Yunnan, southwest China. It is, however, less understood in scientific fields. The Lemo people (a branch of the Bai minority nationality) traditionally grow lacquer trees interplanted with upland food crops in swidden fields. During a 10–15 year fallow period, farmers can harvest various products from lacquer trees, including resin for selling or trading, leafy shoots for vegetable, pericarps for making wax, roots and leaves for pesticide, dry resin for medicine, and seeds for vegetable oil extraction. The Lemo people believe the lacquer tree is the most important crop in their community. The lacquer agroforestry system provides the Lemo people with food, cash income and environmental benefits. Further studies on the lacquer agroforestry system will be indispensable to improve this system so as to disseminate it to other communities.This revised version was published online in November 2005 with corrections to the Cover Date.     

Using 3D architectural models to assess light availability and root bulkiness in coconut agroforestry systems

Using 3D architectural models to assess light availability and root bulkiness in agroforestry systems. In many parts of the humid tropics, coconut trees are frequently intercropped with food crops, or tree crops such as cocoa. The performance of such systems depends on planting patterns, but also on growing conditions for crops below the coconut canopy throughout the development of the coconut trees. We used a modelling approach providing indicators for assessing above-ground competition for light and below-ground competition for space, in order to optimize intercropping in coconut smallholdings. Light transmission and the number of coconut roots in the interrow were assessed in coconut smallholdings from 6 to 60 years old. The modelling of light transmission through coconut stands was based on three-dimensional virtual coconut trees and a numerical light model that computed the shade cast by coconut trees on underlying crops. Root colonization in the interrow was assessed with virtual 3D coconut root systems. Our results showed that intercropping with shade-tolerant species was not limited by light transmission from the 35th year after coconut tree planting. However, at that stage of coconut tree development, the density of primary roots in the interrow limited intercrop development, especially for root and tuber crops. Alteration of the planting pattern over time increased light transmission but did not significantly affect root density. This modelling approach, which involved little parameterization that was easily done, appeared to be an efficient tool for recommending coconut tree planting patterns and densities, as well as indicating intercrop potential depending on their location in the most sunlit areas with minimum root competition.

Light-related competitive effects of overstory trees on the understory conifer saplings in a subalpine forest

To clarify the mechanism by which overstory trees shade understory saplings, we investigated the relationships among light conditions of the saplings (measured as indirect site factor; ISF and direct site factor; DSF), the calculated competition effects of overstory trees on the saplings (W), and relative height growth rate of the saplings (RHGR). We calculated several W values in order to find a W value which can express the light conditions as appropriately as possible, and the results indicated that W explained only 21.9%–24.7% of the total variance of light conditions in the cases where W gave the best fit. In this study, W was calculated based on the basal areas of overstory trees. However, it is known that canopy structure also affects the light regimes in the forest understory, and this might yield the possible errors even within W representing the shading effects most adequately. Therefore, although W significantly represents the shading effect from overstory trees, a great proportion of the variance remained without being explained by W. RHGR was negatively correlated with W, and the W value which had the most adequate explanation of the shading effect also showed the best negative correlation with RHGR. This provides the evidence that the competitive effect of overstory trees on sapling growth is mediated by the shading effect, indicating that competition for light clearly exists within this forest. Such competition for light may closely relate to the well-known phenomenon of gap regeneration in subalpine forests in central Japan.     

The banana (Musa spp.) — Coffee robusta: traditional agroforestry system of Uganda

Banana (Musa spp.) — with Coffee robusta a traditional agroforestry system in Uganda is a purely mixed cropping system managed on a sustained production basis. The basic primary production is banana as a main food crop and Coffee robusta as the main cash crop. The variety of trees deliberately retained, managed and planted, animals and crops in the system are managed by the farmers to maximize production by supplying timber, fruits, fuelwood, fodder, food, medicinal and other miscellaneous uses.