Structural development of pinus sylvestris stands with varying initial density: A simulation model

The growth and structural development of Scots pine (Pinus sylvestris L.) trees growing at different spacing was simulated using a model based on the dry matter production per needle biomass unit and its allocation to needles, branches and stem. Special emphasis was given to the effect of stand density on the growth of the crown system and its implications on branchiness and timber quality. The simulations showed that the needle biomass culminates considerably earlier than the branch biomass with a time lag inversely related to the stand density. The lengths of living and dead crown were also inversely related to stand density. The resulting differences in branchiness were especially obvious in the early development of the tree stands. In the long run these differences tend to disappear, indicating equal external branchiness independently of the initial spacing for mature stands of Scots pine. The internal branchiness, however, was particularly sensitive to the initial spacing.     

Relation between crown parameters and volume increment of Picea abies stands damaged by environmental pollution

Although a good correlation has been found between the mean degree of damage and the increment trend of larger Norway spruce collectives, individual spruce trees often betray no relationship between crown damage and the development of the volume increment. Detailed investigations show that stem and crown dimensions, growing space as well as the competition of neighboring stems can substantially outweigh the influence of crown damage on the volume increment. Furthermore, current methods to determine crown parameters and needle loss are in many cases quite error‐prone. Use of more accurate techniques for measuring spruce trees produced a stronger relation between the volume increment per square meter crown surface area and the degree of crown damage (needle loss).     

辽宁杨树纸浆林发展可行性及技术对策

论述了杨树纸浆材市场需求及展望，杨树纸浆试验林在辽西地区的生长及产量指标，杨树纸浆林主要品种的制浆造纸性能指标，辽宁省杨树纸浆林栽培适宜区域及主要栽培模式，杨树纸浆林栽培技术要点和病虫害防治。     

Branch mortality and potential litterfall from Douglas-fir trees in stands of varying density

Forest floor characteristics influence nutrient cycling and energy flow properties of forest ecosystems, and determine quality of habitat for many forest plants and animals. Differential crown recession and crown development among stands of differing density suggest that an opportunity may exist to control the input of fine woody litter into the system by manipulating stand density. The objective was to measure the rate of branch mortality among stands of differing density and to estimate the range in total per hectare necromass inputs. Although litter traps are reliable for estimating per hectare rates of litterfall, branch mortality dating on sectioned stems uniquely allows assessment of several other litterfall components: (1) individual tree contributions to total litterfall; (2) the amount of branch material released by mortality, regardless of whether the branches are shed to the forest floor; (3) the distribution of basal diameters characterizing the litterfall from a given tree and stand. Twenty-four trees were felled and sectioned on permanent plots that were part of a silvicultural study of stand density regimes in Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco.). Whorl branches were dissected out of bole sections to determine the dates of mortality, and a branch biomass equation was applied to estimate potential rate of litterfall. Periodic annual rates were expressed in four ways: (1) number of branches per tree; (2) mass of branches per tree; (3) mass of branches per unit of crown projection area; (4) mass of branches per hectare. For the growth periods investigated, larger trees and trees growing on denser plots tended to release a greater necromass through branch mortality. Average branch basal diameter generally decreased with increasing stand density. Annual branch mortality ranged from 33 to 430 g m⁻² crown projection area for individual trees, and from 236 to 1035 kg ha⁻¹ for individual plots. These rates approached the low end of the range of previously published fine litterfall rates for Douglas-fir. Rates on these plots were relatively low owing to the temporary delay in crown recession imposed by artificial thinning. A conceptual model of branch litter dynamics is presented to depict consistencies with crown development among stands managed under different density regimes.     

不同林分密度对桉树幼林木材材性的影响

对两种桉树无性系(巨细桉DH201-2和尾巨桉DH32-29)在5种不同林分密度下(1250、1657、2500、5000和6667株.hm-2)的木材材性(包括木材基本密度和纤维形态特征)进行了测量和研究分析(纤维形态特征分两种不同材型分析:心材与边材)。结果表明:无性系DH201-2的林分密度3(2 500株.hm-2)木材基本密度最大,为452 kg.m-3,DH32-29以林分密度2(1657株.hm-2)的木材密度最大,为436 kg.m-3;相同无性系相同林分密度下,边材纤维长度比心材纤维长度长,边材长宽比一般比心材长宽比要大,纤维宽度则无显著差异;无性系DH201-2的林分密度3(2500株.hm-2)和林分密度5(6667株.hm-2)是符合林分蓄积量、木材基本密度和纤维长度最大值的组合。     

Water use by Eucalyptus tereticornis stands of differing density in southern India

We studied water use by Eucalyptus tereticornis Sm. in two plantations, differing in tree density (1800 stems ha(-1) at Site I and 1090 stems ha(-1) at Site II), in different years. At both sites, stomatal conductance, predawn and midday water potentials and microclimate were measured and used to estimate hourly transpiration by the Penman-Monteith equation. Growth in girth was also measured. Stomatal conductance was closely correlated with atmospheric vapor pressure deficit (D); however, stomata did not close completely even at high D ( approximately 5.0 kPa). Midday leaf water potentials did not fall below -2.0 MPa during any part of the year at either site. Predawn leaf water potentials were greater than -0.25 MPa during the postmonsoon period, but declined to -0.7 MPa at Site I during the premonsoon period. Transpiration estimates ranged from 0.6 to 1.2 mm h(-1) at Site I and from 0.2 to 0.6 mm h(-1) at Site II. The extrapolated transpiration values for the rain-free days of the year were 1563 mm and 853 mm for Sites I and II, respectively. Growth in girth was negligible during the premonsoon period. Photosynthesis was not affected by the minor water stress that developed during the premonsoon period.     

Canopy density in stands of Picea abies and Pinus sylvestris after different thinning methods

Altogether 82 plots (261 estimations) of Picea abies (L.) Karst, and 193 plots (360 estimations) of Pinus sylvestris (L.) stands were estimated by a vertical tube. The “crown free projection”, CFP, of stands thinned in three methods with different thinning grades was measured: unthinned, heavily and very heavily thinned, heavily thinned delayed first thinning, extra heavily thinned and thinned from the top. Basal area (m²ha^?1) density (stems ha^?1) and diameter sum (m ha^?1) were plotted against CFP. Basal area was the best practical measure of stand in this study. Generally Scots pine stands have higher CFP and the curves are steeper than in Norway spruce stands. Depending on the grade of thinning, heavily and very heavily thinned spruce stands, delayed first thinning included, have CFP values of 10–15% and stands thinned from the top, 20–40%, compared with 30–80% and 30–60% respectively in pine stands. Extra heavily thinned stands have the highest CFP, 20–80% in spruce and 50–90% in pine stands. The CFP levels after thinning are too high in pine stands for avoidance of sucker and sprout production of aspen and birch. In dense Norway spruce stands thinned from the top or heavily and very heavily thinned, the CFP values are low enough (≤30%) to diminish the production of suckers.     

Comparison between old-growth stands and secondary stands regenerating after clear-felling in warm-temperate forests of Yakushima, southern Japan

In order to infer successional changes in structure, species composition and diversity of warm-temperate forest, we compared secondary stands regenerating after clear-felling (41–64-years old) with old-growth stands at altitudes between 300 and 800 m on Yakushima Island, southern Japan. Stem density and maximum stem diameter differed between secondary and old-growth stands, but basal area and aboveground biomass did not. At lower altitudes, the dominant species in old-growth stands with a strong sprouting capacity (Castanopsis cuspidata) also dominated secondary stands, and species composition of secondary and old-growth stands was similar. At higher altitudes, by contrast, the dominant species in old-growth stands (Distylium racemosum) had little sprouting capacity and was poorly represented in diverse secondary stands, which were dominated by Castanopsis or other less abundant species. Secondary stands had greater species diversity (Shannon–Wiener index) than old-growth stands, particularly at higher altitudes. This was due to greater species richness resulting from higher stem density per area, but not to greater evenness. We grouped the component species that share ecologically similar traits into four guilds (fagaceous, primary evergreen, secondary evergreen and deciduous species). Secondary stands were characterized by greater numbers of deciduous and secondary evergreen species. We concluded that different sprouting capacities of dominant species and different regeneration traits among guilds are responsible for the change in species composition and diversity during succession.     

Comparison of tree size structure and growth for partially harvested and even-aged hemlock-spruce stands in southeast Alaska

The effects of partial cutting on tree size structure and stand growth were evaluated in 52 plots in 13 stands in southeast Alaska that were partially harvested 53–96 years ago and compared with 50-year-old even-aged stands that developed after clearcutting. The net basal-area growth was greater in the partially cut plots than in the uncut plots, and basal-area growth generally increased with increasing cutting intensity. However, the basal-area growth of all of the partially harvested stands was significantly less than the growth of 50-year-old even-aged stands, and net basal area growth over the 50 year period since partial harvesting was about 33–43% of the growth of the even-aged stands. Partial cutting maintained stand structures similar to uncut old-growth stands, and the cutting had no significant effect on tree species composition. The tree size distribution of the partially harvested stands was far more complex and well distributed in comparison with the 50-year-old even-aged stands, and included the presence of several trees with diameters of more than 100 cm. These trees included both large-diameter spruce and hemlock trees and were a distinctive structural feature that was noticeably lacking in the even-aged stands.     

Relationship between tree morphology and growth stress in mature European beech stands

Aims

In European Beech (Fagus sylvatica L.) large growth stresses lead to severe log end splitting that devaluate beech timber. Our study aimed at detecting relationships between growth stress and some morphology parameters in trees.

Methods

Growth stress indicators were recorded for 440 mature trees in nine stands from five European countries, together with morphology parameters.

Results

Most trees displayed an uneven distribution of growth stress around the trunk. Moreover, growth stress intensity varied largely between individual trees. Geometry of the trunk was a poor predictor of growth stress intensity. Crown asymmetry resulted in a larger stress dissymmetry within trees. Trunk inclination was not correlated to max tension stress, contrary to what is usually found in younger trees. In the case of small inclination, growth stress was close to expected from biomechanics of restoring verticality. Trees exhibiting a larger inclination probably evolved a different mechanical solution: a rather large crown, lower tree slenderness and a sufficient asymmetry in growth stress as to prevent a higher inclination due to growth.

Conclusion

A large slenderness is the best accurate predictor of a large growth stress, although variations in the ratio height/diameter at breast height explained only 10 % of the variability of growth stress. A large crown surface was the best predictor of a low level of growth stress. A large spacing between trees seems a good solution to lower the risk of growth stress in mature beech.     

ESTIMATING DENSITY AND POPULATION SIZE OF TIBETAN SNOWCOCK IN TIBET

PREFACETibetansnowcock(Tetragallustibetanus)isakindofplateaubird,whichlivesinhighestlandamongseveralpheas-antsintheworld.ThesnowcockismainlydistributinginTibetanditshabitatstretchesfromeastinQinlingMountainstowestinPamirPlateau.Tibetisavastexpanseofplateauterritorywithe1evationofmorethan4,OOom.TherearemanysnowmountainsinTibet,especial1yinwesternparts.Theriversand1akesspreadovera1lplaces.BecauseofhightopographyfeaturesandtheHimalayasmountainsys-temacrossfromeasttowest,thechimateisveryco…     

Using density management diagrams to assess crown fire potential in Pinus pinaster Ait. stands

Context

Density management diagrams (DMDs) are useful for designing, displaying and evaluating alternative density management regimes for a given stand-level management objective. The inclusion of variables related to crown fire potential within DMDs has not previously been considered.

Aims

The aim of this study was to include isolines of variables related to crown fire initiation and spread in DMDs to enable identification of stand structures associated with different types of wildfire.

Methods

Biometric and fuel data from maritime pine (Pinus pinaster Ait.) stands in NW Spain were used to construct DMDs. Different surface and crown fire behaviour models were used together to estimate crown fire potential.

Results

The crown fire potential varied greatly throughout development of the maritime pine stands. Low stands were more prone to crowning. The type of crown fire was mainly determined by stand density.

Conclusion

The DMDs developed can be used to identify relationships between stand structure and crown fire potential, thus enabling the design of thinning schedules aimed at reducing the likelihood of crowning.     

论建筑与结构的关系

本文阐述了在建筑设计中,选择结构型式不仅是结构工程师的任务,也是建筑师的工作,一个 好的建筑设计应该是建筑师和结构工程师密切合作的成果,这种合作应当达到配合默契的程度。     

Variation in individual biomass decreases faster than mean biomass with increasing density of bamboo stands

The total biomass of a stand is an indicator of stand productivity and is closely related to the density of plants. According to the self-thinning law, mean individual biomass follows a negative power law with plant density. If the variance of individual biomass is constant, we can expect increased stand productivity with increasing plant density. However, Taylor's power law(TPL) that relates the variance and the mean of many biological measures(e.g. bilateral areal differences of a leaf, plant biomass atdifferent times, developmental rates at different temperatures, population densities on different spatial or temporal scales), affects the estimate of stand productivity when it is defined as the total biomass of large plants in a stand.Because the variance of individual biomass decreases faster than mean individual biomass, differences in individual biomass decline with increasing density, leading to more homogeneous timbers of greater economic value. We tested whether TPL in plant biomass holds for different species and whether the variance of individual biomass changes faster than the mean with increasing stand density.The height, ground diameter and fresh weight of 50 bamboo species were measured in 50 stands ranging from 1 m by 1 m to 30 m by 30 m to ensure more than 150 bamboos in every stand. We separately examined TPL in height,ground diameter, and weight, and found that TPL holds for all three biological measures, with the relationship strongest for weight. Using analysis of covariance to compare the regression slopes of logarithmic mean and variance against the logarithm of density, we found that the variance in individual biomass declined faster than the mean with increasing density. This suggests that dense planting reduced mean individual biomass but homogenized individual biomass. Thus, there exists a trade-off between effective stand productivity and stand density for optimal forest management. Sparse planting leads to large variation in individual biomass, whereas dense planting reduces mean individual biomass. Consequently, stand density for a plantation should be set based on this trade-off with reference to market demands.     

Trade-offs between seedling growth,thinning and stand stability in Sitka spruce stands: a modelling analysis

The forest industry is increasingly adopting alternative silvicultural systems, involving regeneration beneath an existing forest canopy, rather than clear-felling and replanting. To apply these silvicultural systems in windy regions such as Britain and Ireland, it is essential that the interactions between thinning intensity, stand stability and seedling growth are properly understood. Here, we present a modelling analysis of the three key relationships between: (i) stand density and the proportion of incident radiation transmitted through a forest canopy as a stand is thinned; (ii) transmitted radiation and seedling growth, and (iii) stand density and stand stability. These relationships were examined using separate models of radiative transfer (MAESTRO), seedling growth, and stand stability/wind risk (ForestGALES). Output from the three models was synthesised to calculate whether a given stand thinned to a pre-defined stability limit would allow sufficient light to penetrate the canopy for seedling growth. A minimum transmittance of 20% was identified as a requirement for seedling growth, which corresponds to removing 45% of stand basal area. A thinning of this intensity left some stands susceptible to unacceptable wind damage, especially in old or previously thinned stands on soils where rooting is impeded. The results emphasised the fact that rooting conditions, thinning history and age of intervention are major constraints on the silvicultural options. In general, older stands are not suitable for conversion to continuous cover forestry (CCF) systems, and the transformation process should begin at pole stage, when heavy thinning does not leave the stand unstable. The analysis approach used here illustrates the potential for combining models to address complex forest management issues.     

Stand density and fertilization effects on aboveground allocation patterns and stem form of Pinus sylvestris in young stands

The increasing demand for forest biomass for energy generation could be partially met by growing denser stands and use of fertilizer. Before this is done at large scale, more knowledge of the effects of stand density and fertilization on aboveground allocation patterns and stem form is needed. Therefore, effects of pre-commercial thinning (PCT) to 3000 stems ha^?1, an unthinned dense control (C), and PCT combined with two levels of fertilization (100 kg ha^?1 of nitrogen applied either during the establishment of the field experiment (F1) or annually (F2)) were examined in 23- to 26-year-old Scots pine (Pinus sylvestris L.) stands six years after the establishment of the field experiment. In total, 114 sample trees were harvested using destructive biomass sampling. The growth allocation and stem form of trees with diameter at breast height (DBH; 1.3 m height) >5.0 cm were not affected by either the PCT or fertilization. Small trees (DBH < 5 cm) in denser, unthinned control plots had more slender stems (lower DBH/height ratios) and allocated less growth to branches and foliage than trees in PCT plots. Fertilization had little effect on the stem form and growth allocation of the smallest trees. Therefore, effects of stem density and fertilization on stem form and growth allocation to foliage were only found for small suppressed trees, and the treatments had very little influence on dominant and codominant trees.     

The effect of density after precommercial thinning on volume and structure in Pinus Sylvestris and Picea Abies stands

The interaction between stand density and dominant height and the development of volume, mean diameter, mortality and distribution of volume during a period from precommercial thinning to first commercial thinning was studied on permanent plots in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) stands in Sweden. The reciprocal equation of the yield density effect was used to level the volume yield. Higher density after precommercial thinning resulted in higher yield and smaller mean diameter. The mortality up to first thinning was low, but is also dependent on density after precommercial thinning. The positive skewness of the volume distribution was higher in denser stands.