采伐作业对长白山阔叶红松林林下植被影响的研究(英文)

森林采伐是森林经营中的一个关键环节。采伐对树木天然更新的影响直接关系到森林的结构、组成及森林的可持续经营。本文以长白山阔叶红松林为研究对象,对择伐5 年后不同作业迹地(集材道、楞场和采伐后林窗)的幼苗更新及灌木、草本的多样性进行研究,并以没有进行采伐干扰的林地作为对照。共设计样地23 块,在各个小样地内调查幼树的种类、树木、高度;灌木和草本植物的种类、树木、高度和盖度,应用SPSS 软件进行一元方差分析。结果表明,择伐作业影响幼苗的更新,灌木和草本的多样性在作业迹地增加,尤其在林隙处的多样性最高。适当采伐能促进幼苗的更新,特别是阔叶树种的更新。集材道对幼苗密度的影响比对幼苗高度的影响大,幼苗密度在集材道上密度最高;采伐作业对针叶树种有严重的影响,在三种作业迹地上针叶树种的密度都比对照低。为了维持森林的结构和组成,在森林收获和楞场的建立时,必须采取科学措施保护针叶树种。此外,采伐林窗和楞场的面积都应该减小。为了确保采伐迹地建群树种的更新,经营中应该在保护生物多样性的前提下,适当控制过于旺盛的灌木和草本层的生长。本研究结果将为采伐后作业迹地的恢复和森林的可持续经营提供科学的依据。图1 表4 参14。     

Impact of mining on tree diversity of the silica mining forest area at Shankargarh,Allahabad, India

The Shankargarh forest area is rich in silica, a major mineral used in glass industry. Extensive open cast silica mining has severely damaged the forest as well as productivity of the region. An understanding of the impact of mining on the environment particularly on vegetation characteristics is a prerequisite for further management of these mining sites, especially in the selection of species for reclamation works. The present paper deals with the study of the tree composition of silica mining area of Shgankargarh forest, at both disturbed and undisturbed sites. Tree vegetation study was conducted at undisturbed and disturbed sites of Shankargarh forests using standard quadrate method. Density, abundance and frequency values of tree species were calculated. Species were categorized into different classes according to their frequency. The importance value index (IVI) for each species was determined. Species diversity, Concentration of dominance, Species richness and Evenness index were calculated for the undisturbed and disturbed sites. The distribution pattern of the species was studied by using Whitford’s index. Similarity index between tree composition of disturbed and undisturbed sites was determined by using Jaccard’s and Sorenson’s index of similarity. Tree species showed a drastic reduction in their numbers in disturbed sites compared to that of the undisturbed sites. The phytosociological indices also illustrated the impact of mining on the tree composition of the area. The present study led to the conclusion that resultant tree vegetation analysis can be used as important tool for predicting the suitability of particular species for revegetating the mined areas.     

燕山山地油松人工林林隙大小对更新的影响

林隙(gap)又称林窗,指森林群落中老龄树木死亡或偶然因素(如干旱、台风、火灾等)造成成熟阶段的优势种死亡,从而形成林冠空隙的现象(Watt,1947)。美国生态学家Runkle(1982)将林隙分为2种,一种是林冠空隙(canopy gap),指上方无树冠遮荫的部分,即太阳处于正中位置时无树冠遮荫的面积之和;另一种是扩展林隙(expanded gap),指形成该林隙周围树木基部所包围的面积(Runkle,1981)。本研究所指的林隙为应用最为广泛的扩展林隙。近年来的森林动态学研究表明:林隙在森林结构和多样性维持中起着重要的作用,已     

Tree recruitment in gaps of various size,clearcuts and undisturbed mixed forest of interior British Columbia,Canada

Tree seedling recruitment was monitored after various types of logging in mixed conifer and deciduous forests of northern British Columbia, Canada. Predicting tree seedling recruitment after disturbance is fundamental to understanding forest dynamics and succession and is vital for forest management purposes. Seedling recruitment success in multi-species northern latitude forests varied as a function of mature tree canopy cover, gap size and position in a gap. Recruitment was abundant within canopy gaps across a wide range of gap sizes (20–5000 m²), but recruit numbers dropped off rapidly under the closed forest canopy and in the open conditions of clearcuts. Inside canopy gaps, recruitment was similar by gap position in small gaps (<300 m²) but, in these northern latitude forests, exhibited a trend of increasing density from the sunny north to shady south end of larger gaps. This was true for all tree species regardless of their shade tolerance ranking. There was no evidence of gap partitioning by any of the tree species during the regeneration phase suggesting that adaptation to the subtleties of gap size during early recruitment are not well developed in these tree species. Favorable locations for emergence and early establishment of germinants were less favorable for growth and survival of established seedlings, i.e. the regeneration niches in these forests were discordant. Tree abundance and species diversity appears to be controlled more by differentiation among growth and survival niches than by the regeneration niches. From the perspective of forest management, abundant natural regeneration of all the dominant tree species of these mixed-species forests can be obtained after partial cutting.     

Effects of treefall gaps created by windthrow on bat assemblages in a temperate forest

Determining the way in which spatial distribution and diversity of forest-dwelling mammals varies with natural disturbance is essential to understanding the spatial dynamics of mammal assemblages in forests. Bats are the only forest-dwelling mammals capable of true flight. At a local scale, bat flight ability, which may be related to ecomorphological traits, is an important factor influencing spatial distribution. We tested two postulates: (1) the spatial distribution of bats is affected by sizes of forest gaps created by natural disturbances and (2) species-specific responses can be predicted from bat ecomorphological traits (aspect ratio (AR) and wingtip shape index (WT)) that influence bat flight ability. We found that sizes of forest gaps affected the occurrence of each bat species and species richness of bats at local scales; species-specific responses were related to the ecomorphological traits of individual species. Bat species with high AR and low WT were not affected by variation in canopy gap size. In contrast, bat species with low AR and high WT responded negatively to gap size, and those with intermediate AR and WT responded positively to canopy gap size at sites with small-sized gaps but responded negatively to large-sized gaps. Overall bat species richness responded negatively to gap size. Thus, ecomorphological traits may be important determinants of bat spatial distributions and species diversity at local scales in disturbed habitats. In this study, forest edges might have been undersampled due to the location of bat detectors. However, this potential undersampling should not have affected the interpretation of occurrence patterns of bat species responding to gap size. Our results imply that bat conservation efforts in forest lands should take into consideration specific responses related to ecomorphological traits of species inhabiting an area. The results also suggest that quantifying the effects of natural disturbances on bat assemblages may provide a knowledge base for forest management to minimize the impacts of unavoidable anthropogenic disturbances on bat species diversity. Rare or infrequent natural disturbances can provide models for forest management aimed at maintaining bat species diversity.     

林窗效应研究综述

森林群落常发生一些小规模的内源干扰,从而形成林窗。林窗的形成对推动森林群落的演替更新和生态系统发展至关重要。林窗面积的大小与树木的倒伏方式和林冠冠幅及大小有关。林窗面积及林窗内位置的不同,导致其小气候和土壤理化性质等环境因子发生改变,进而影响到林窗内树种更新和物种组成、林窗植被的物种多样性及其微生物和土壤动物的种类、数量等方面。未来林窗研究重点应该放在次生林和人工林的林窗效应,林窗对森林生态系统碳储量影响机制,林窗凋落物分解因子间的相互关系、作用机理和养分循环,不同树种的林窗与最适更新面积的关系,林窗的边缘效应,林窗的土壤动物和微生物动态及过程。     

The influence of gap creation on the regeneration of Pinus tabuliformis planted forest and its role in the near-natural cultivation strategy for planted forest management

It remains unclear whether or not creating gaps in planted forests can increase the plant species composition, structure, and biodiversity, and also whether it can be helpful for restoring planted forests (to a more natural state). Based on a comparison of species composition and structure among forest patches, small gaps (4-25 m²), medium gaps (25-150 m²) and large gaps (150-450 m²), we found that (1) creating gaps enhanced vascular plant diversity. Both the species richness and Shannon diversity indices of small, medium and large gaps were significantly higher than in the understory. The pattern of increasing diversity of vascular plants with gap creation could be partly attributed to the emergence of novel shade intolerant species in gaps. (2) Creating gaps favored the colonization and regeneration of native species. Gap size influenced not only the emergence and density of individuals of different species, but also the emergence of different life form types. Small gaps promoted the regeneration of some shrub species, such as Ostryopsis davidiana, Rosa hugonis, and Forsythia suspense, leading to these species becoming canopy dominants early on in succession. The medium and large gaps favored the growth of tree species, such as Populus davidiana and Betula platyphylla (early successional stage), and Quercus liaotungensis and Pinus tabulaeformis (later successional phase). (3) The canonical correspondence analysis showed that plant species composition and distribution were mainly influenced by gap size and slope aspect, and that the recorded plant species could be divided into three life forms (trees, shrubs and herbs) on the biplot diagram. (4) Finally, creating gaps provided opportunities not only for native pioneer species in the early successional stage, but also for climatic climax species to grow to canopy dominants in later successional phases, suggesting that a more natural forest will develop with plant succession. Gap size plays an important role in plant regeneration, and it could be used to produce desired successional communities in near natural management for planted forests.     

Effects of gaps on regeneration of woody plants: a meta-analysis

Forest gaps, openings in the canopy caused by death of one or more trees, have a profound effect on forest regeneration and drive the forest growth cycle. It is therefore necessary to understand the effects of forest gaps on regeneration for modern forest management. In order to provide a quantitative assessment of the effects of forest gaps on regen-eration of woody plants, we conducted this review of gap effects on woody plant regeneration on the basis of 527 observations from 42 indi-vidual papers, and reported the results of these data in a meta-analysis. Overall, densities of regenerated woody plants were significantly greater （359%） in forest gaps than on the closed-canopy forest floor. The regen-eration density in gaps of plantation forests was significantly greater （P＆lt;0.05） than that of natural forest because the regeneration in gaps of plan-tation forests was improved by both gap effects and experimental meas-ures. Similarly, in comparison to natural gaps, regeneration was better enhanced in artificial gaps. Regeneration density exhibited a significantly positive correlation with gap size, but a negative correlation with gap age because the gap size decreased with increasing gap age. Shade tolerance of woody plants affected regeneration density in gaps and understory. Average regeneration density of shade-tolerant species exhibited a sig-nificantly positive response to gaps but densities remained lower in total than those of intermediate and shade-intolerant species. Gap effects on regeneration decreased in response to increasing temperature and pre-cipitation because of the limiting effects of lower temperature and moisture on woody plant regeneration. In summary, forest gaps enhance woody plant regeneration, and the effects of gaps varied by forest type, gap characteristics, environmental factors and plant traits. The results of this meta-analysis are useful for better understanding the effects and roles of gaps on forest regeneration and forest management.     

林窗定义及林窗特征测定方法研究进展

林窗形成导致环境资源再分配,进而影响物种入侵、种子萌发、幼苗更新和幼树存活;林窗改变了森林空间结构和植物群落组成,增加了群落异质性,这为保持森林群落相对稳定奠定了基础,也为多种生物共存发展提供了适宜的条件。目前,关于林窗对植被更新、植物多样性影响研究较多,但至今没有一个公认的客观量化林窗定义的方法,导致研究者采用的林窗识别标准不同,无法对现有林窗研究进行有效比较,且林窗特征测定方法也尚未完善。文中对林窗的概念形成、定义发展及林窗特征（形状、大小、形成、年龄及边缘木）测定方法进行梳理,探讨目前林窗研究存在的问题,以期为未来林窗干扰相关研究提供历史脉络与研究方向。     

Gap phase regeneration recruitment of mixed conifer-broadleaf forests in Wolong Nature Reserve

Wolong Nature Reserve is the largest reserve for protecting the endangered wild giant panda. Due to historical factors, even in many sections of the core protection area, the forests have been seriously destroyed and natural forests are poorly restored. However, the relative importance of the determinants for recruitment of communities under disturbance is rarely explored. In our study the endogenous and exterior factors in a forest gap that affect the conifer-broad-leaved mixed forest regeneration were investigated near Wuyipeng, one of the observation stations at Wolong, to explore which determinant had the greatest effect on gap regeneration and to discover the recruitment of seedling establishment in forest gaps. With a linear sampling method, environmental factors, gap characteristics and recruitment of new individuals were measured and examined in every forest gap along three sampling lines. Data of environmental factors in the gaps were collected for a Pearson correlation analysis in order to explore the disturbance and preprocessed characteristics of the gaps, using principal component analysis in SPSS. Correlation analysis was applied to further explore the relationship between changes in the gaps and the response of the regenerating seedlings. The results show that a range of natural and human disturbances affected the pattern and characteristics of the forest gaps in this area. The richness in the composition of the seedlings was higher than that of gap makers, but the order of dominance of the composition was different between seedlings and gap makers. The success of dominant species in establishing themselves was affected by different environmental factors. For instance, the establishment of Betula spp. was correlated significantly with topographic factors, while that of Abies faxoniana was affected by soil characteristics and that of Rhododendron spp. correlated significantly with topographic factors and characteristics of gap makers. Moreover, all the biodiversity indices of regenerating seedlings were significantly correlated with environmental principle components mostly reflecting soil properties. From this we can infer that soil characteristics are the most important factors affecting the regeneration of dominant species and seedling diversity under gap disturbances. __________ Translated from Journal of Plant Ecology (Chinese Version), 2007, 31 (3): 425–430 [译自: 植物生态学报]     

Canopy gap characteristics and its influence on the regeneration of broad-leaved Korean pine forests in Changbai Mountain

lntroductionBroad-leavedKoreanpineforestisacldriaxforestli1tCm-peratezoneofNortheaster11Cl1ina.ltscharacteristichasIligllstabilityandproductivity.Therewerenlanyresearcl1worksfocusedontl1estructure,dynandcsanditsmain-tainingmechanisn1.Researchontheforestregeneratio11wasamali1approaclltoundersta11dtlleforestdynanlicsI;j.ThereweremanyresearcI1eso11tl1erege11erationofh1iskindofforestsincel95ol5~ll].Butfewoftl1emweredoneundertheconsiderationofcanopygapdistUrbance.Sincel99O's,moreandmoreresearch…     

Factors affecting soil-based natural regeneration of <Emphasis Type="Italic">Abies sachalinensis</Emphasis> following timber harvesting in a sub-boreal forest in Japan

Natural regeneration of Abies sachalinensis on soil was studied in a natural sub-boreal forest managed by single tree selection cutting located within the Hokkaido Tokyo University Forest. First, seedlings of A. sachalinensis on skid trails, soil mounds, soil pits, and undisturbed soil were counted, and their areas were measured. Most seedlings were found on skid trails. Seedling densities of A. sachalinensis on sites with soil disturbance, with the exception of soil pits, were significantly greater than that on undisturbed soil. Second, densities of Sasa senanensis, heights of S. senanensis, and seedling densities of A. sachalinensis on skid trails and undisturbed soil in both a closed-canopy stand and in a canopy gap were compared. Seedling density of A. sachalinensis on skid trails in the closed-canopy stand was significantly greater than that in the canopy gap. Average density and average height of S. senanensis, which is known to interfere with regeneration of many woody species, were significantly greater on skid trails and undisturbed soil in the canopy gap than those in the closed-canopy stand. It could be concluded from this study that both crown closure and soil disturbance are essential for natural regeneration of A. sachalinensis on soil in sub-boreal forests.     

Influences of life history, environmental gradients, and disturbance on riparian tree regeneration in Western Oregon

In two related field studies: (1) a multiscale riparian forest inventory and (2) a comparative study of natural forest gap and nongap environments, we explored regeneration patterns of native riparian trees in relation to large- and small-scale ecological drivers in four western Oregon watersheds spanning a climatic gradient from dry to wet. Twenty-three tree species were classified by life history traits into five functional groups that differed in shade and drought tolerance; distribution and abundance of tree regeneration were analyzed by group. For most groups, seedling abundance varied substantially across the large scale climate gradient. In particular, drought tolerant species decreased sharply in abundance from the drier to wetter watersheds. Overall seedling frequency and diversity also decreased from the driest to wettest watersheds, while nurse log use increased. Regeneration of most, but not all, species was greater in gaps.Principal conclusions are that life history information was useful for categorizing species and quite predictive of regeneration behavior overall, yet for many species contextual factors such as climate, forest structure, and adjoining species were also very important. Species appeared to regenerate opportunistically wherever local conditions fell within their environmental tolerances and competitive abilities. Management or restoration of riparian forests, therefore, requires knowledge of site conditions, the life history of the riparian trees present, and in particular an understanding of the species’ environmental tolerances, disturbance responses, and competitive abilities relative to one another.     

林窗对南酸枣林灌木层植物多样性的影响

于湖南省宁乡县青羊湖国有林场南酸枣林内的41个林窗和41个非林窗中设置样方,对其灌木层植物群落进行调查,采用物种多样性指数研究林窗干扰对南酸枣林灌木层植物多样性的影响。结果表明:林窗干扰改善了微环境,林窗内与非林窗中林下灌木层树种重要值及多样性存在差异。林窗内的灌木层树种组成数量、累计出现的平均密度、频度、丰富度指数R、Shannon-Wienner指数H、Pielout均匀度指数E、综合多样性用均优多指数Z均高于非林窗中的,而其生态优势度指数λ稍低于非林窗中的。     

A review of the roles of forest canopy gaps

Treefall gap, canopy opening caused by the death of one or more trees, is the dominant form of disturbance in many forest systems worldwide. Gaps play an important role in forest ecology helping to pre- serve bio- and pedo-diversity, influencing nutrient cycles, and maintain- ing the complex structure of the late-successional forests. Over the last 30 years, numerous reviews have been written describing gap dynamics. Here we synthesize current understanding on gap dynamics relating to tree regeneration with particular emphasis on gap characteristics consid- ered critical to develop ecologically sustainable forest management sys- tems and to conserve native biodiversity. Specifically, we addressed the question： how do gaps influence forest structure？ From the literature re- viewed, the size of gaps induces important changes in factors such as light intensity, soil humidity and soil biological properties that influence tree species regeneration and differ in gaps of different sizes. Shade- tolerant species can colonize small gaps; shade-intolerant species need large gaps for successful regeneration. Additionally, gap dynamics differ between temperate, boreal, and tropical forests, showing the importance of climate differences in driving forest regeneration. This review summa- rizes information of use to forest managers who design cutting regimes that mimic natural disturbances and who must consider forest structure, forest climate, and the role of natural disturbance in their designs.     

冰雪灾害对九连山常绿阔叶林幼苗更新的影响

以九连山亚热带常绿阔叶林为研究对象,对雪灾林隙与非林隙条件下幼苗的组成、多样性等进行研究,分析冰雪灾害对九连山常绿阔叶林幼苗更新的影响.研究结果表明:(1)幼苗的个体密度在林隙内外差别不大;(2)乔木幼苗在林隙中的重要值大于非林隙,灌木幼苗在非林隙中的重要值大于林隙;(3)主要乔木树种的幼苗对林隙的反应存在很大的差异;...     

Bottomland hardwood forest recovery following tornado disturbance and salvage logging

Catastrophic wind events, including tornado, hurricane, and linear winds, are significant disturbances in temperate forested wetlands. Information is lacking on how post-disturbance salvage logging may impact short and long-term objectives in conservation areas where natural stands are typically managed passively. Woody regeneration and herbaceous cover were assessed for three years in a bottomland hardwood forest across a gradient of damage from an F4 tornado, with and without subsequent salvage logging. Soil disturbance intensity and recovery associated with salvage logging within wind-disturbed sites were also assessed. Woody stem density and proportion of potential overstory species (species with the potential to occupy a position in the canopy) increased as a function of wind disturbance intensity. Stem density, proportion of overstory trees, or species diversity did not differ between wind + salvage and wind-disturbed-only plots. Significant dissimilarity occurred among soil disturbance classes within salvaged sites. By the third growing season, vegetation in soil disturbance classes in wind + salvage areas was converging toward undisturbed conditions and bottomland hardwood forest recovery was underway in all vegetation disturbance types and soil disturbance classes. Post-tornado salvage logging, applied judiciously, may contribute to microsite and vegetation diversity.     

Explaining conifer dominance in Afrotemperate forests: Shade tolerance favours Podocarpus latifolius over angiosperm species

We contrasted the regeneration success of angiosperm canopy species with that of Podocarpus latifolius to test whether this conifer was sufficiently shade tolerant to be favoured on infrequently disturbed and well-shaded sites. Seedling and sapling population structures were measured in several habitats representing a light gradient in warm temperate forest in the Drakenberg mountains of South Africa. The angiosperm-dominated forest was well-shaded (∼5.5% PAR) and lacked ground vegetation. Seedlings of angiosperm canopy species were abundant but the absence of saplings indicated regeneration failure. In contrast, P. latifolius was represented by all size classes beneath the intact canopy indicating continuous regeneration in angiosperm-dominated forest. Angiosperm regeneration was similarly poor in forest gaps, which were dominated by grasses, ferns and vines. P. latifolius entered the advanced regeneration in gaps by establishing in shade before gap formation. The floor of the Podocarpus-dominated forest was less shaded (∼7.5% PAR) than angiosperm-dominated forest and dominated by grass, which suppressed most conifer regeneration. Gaps in Podocarpus forest were more than double the size of gaps in angiosperm forest and dominated by vines and understorey shrubs. There was no clear winner of the angiosperm–conifer contest in Podocarpus forest gaps. Consequently, gaps maintain limited angiosperm diversity in Podocarpus forest. Angiosperm canopy species regenerated continuously in the high light (∼11.5% PAR) thicket environment. Few seedlings of P. latifolius were recorded in the thicket environment. Our findings are consistent with the hypothesis that shaded and infrequently disturbed forest sites favour shade-tolerant conifers over relatively light-demanding angiosperm species. The outcome of the competitive interaction between angiosperm and conifer at the regeneration phase depends on the relative shade-tolerance of associated species.     

Assessment of natural regeneration status and diversity of tree species in the biodiversity conservation areas of Northeastern Bangladesh

A study was conducted at two-biodiversity conservation areas of Northeastern Bangladesh (a part of Sylhet Forest Division) to assess the species composition, diversity and density of natural regeneration of tree species both indigenous and exotic species. Data were collected by stratified random quadrate method during January 2010 to July 2010. Totally 200 circular plots of 2 m×2 m in size had 5 different habitat types of plants namely; forest, roadside, homestead (surrounding forest dwellers house), fallow land and others (canals, streams and tea gardens side), which included a total of 55 regenerating species belonging to 28 families. Meliaceae is the dominant family and shows the highest family importance value (26.3), having six species, followed by Moraceae (24.24). Among the five habitat types, forest (43 species) possess the highest number of species, followed by roadside (23 species). Total 15 exotic species among 9 families and 40 indigenous species with 24 families were recorded. For exotic species, Tectona grandis possess the highest relative density (11.7%) and relative frequency (10.5%); Senna siamea had highest relative abundance (7.83%). In case of indigenous species, Chickrassia tabularis possess the highest relative density (4.23%) and relative frequency (4%); Dipterocarpus turbinatus had the highest relative abundance (3.92%). Tectona grandis (29.66) and Chickrassia tabularis (10.8) had the highest IVI for exotic and indigenous species respectively. Different diversity indices such as Shanon-Winner diversity index, species diversity index, species richness index, species evenness index, Simpson index and species dominance index, etc. were applied to quantify definite diversity. The regeneration of species associated with low levels of disturbance was in the exotic species. Study suggests that proper protection from human disturbances and scientific management of natural regeneration of two-study forests may lead a rich biodiversity site in the country.